Pholcus metta, Huber, 2019

Huber, Bernhard A., 2019, The pholcid spiders of Sri Lanka (Araneae: Pholcidae), Zootaxa 4550 (1), pp. 1-57 : 16

publication ID

https://doi.org/ 10.11646/zootaxa.4550.1.1

publication LSID

lsid:zoobank.org:pub:2F7D1EC4-D4ED-4FAE-B227-CF7B79EAE833

DOI

https://doi.org/10.5281/zenodo.4581627

persistent identifier

https://treatment.plazi.org/id/CA3B104C-FF9A-FF94-FF3D-FED1FCF7E6B1

treatment provided by

Plazi

scientific name

Pholcus metta
status

sp. nov.

Pholcus metta View in CoL sp. n.

Figures 56–58 View FIGURES 53–58 , 67–68 View FIGURES 65–72 , 76–78 View FIGURES 73–78 , 86 View FIGURES 85–88 , 90 View FIGURES 89–92

Diagnosis. Males are easily distinguished from other species in ceylonicus group by shape of genital bulb ( Figs 76–78 View FIGURES 73–78 ): distinctive main bulbal process (‘appendix’), i.e. large sclerotized process without retrolateral membrane and retrolateral conical projection (in contrast to P. ceylonicus ), narrower in dorsal view than in P. puranappui ; without prolateral process (in contrast to P. uva ); distinguished from P. uva also by shape of procursus ( Figs 67–68 View FIGURES 65–72 ; narrower and with subdistal dorsal process). Females are distinguished from P. ceylonicus and P. uva by larger posterior excavation of pre-epigynal plate ( Fig. 86 View FIGURES 85–88 ); from P. puranappui by narrower pre-epigynal plate (compare Figs 86–87 View FIGURES 85–88 ).

Etymology. The species name is derived from the Pāli word mettā , one of the four Buddhist virtues or brahmavihāras (active good will towards all); noun in apposition.

Type material. SRI LANKA: ♂ holotype, ZFMK (Ar 20048), North Central Province, Dimbulagala (7.860°N, 81.118°E), 140 m a.s.l., 11.iii.2017 (B.A. Huber) GoogleMaps .

Other material examined. SRI LANKA: 2♂ 5♀, ZFMK (Ar 20049), and 1♂ 1♀ in pure ethanol, ZFMK ( SL126 About ZFMK ), same data as holotype GoogleMaps .

Description. Male (holotype). MEASUREMENTS. Total body length 7.3, carapace width 1.8. Distance PME- PME 240 µm, diameter PME 170 µm, distance PME-ALE 60 µm, distance AME-AME 40 µm, diameter AME 85 µm. Sternum width/length: 1.23/0.97. Leg 1: 55.7 (13.3 + 0.8 + 13.5 + 24.8 + 3.3), tibia 2: 9.5, tibia 3: 6.9, tibia 4: 8.8; tibia 1 L/d: 71. Femora 1–4 width (at half length): 0.24, 0.24, 0.23, 0.22.

COLOR (in ethanol). Carapace ochre-yellow with large dark brown posterior mark, ocular area also brown, clypeus not darkened; sternum ochre-yellow with star-shaped light brown median mark; legs ochre to light brown, with indistinct darker rings subdistally on femora and tibiae, tips of femora and tibiae whitish; abdomen ochregray, with series of cuticular dorsal marks, brown area in front of gonopore and pair of smaller dark marks lateral of gonopore, indistinct wide median band between gonopore and spinnerets.

BODY. Habitus as in close relatives (cf. Figs 47, 50–52 View FIGURES 47–52 ); ocular area slightly raised, each triad on low hump; carapace without median furrow; clypeus and sternum unmodified.

CHELICERAE. As in P. ceylonicus (cf. Huber & Benjamin 2005: fig. 2d), with pair of distal apophyses close to laminae, each with two modified hairs, and pair of small proximal lateral apophyses.

PALPS. As in Figs 56–58 View FIGURES 53–58 ; coxa unmodified; trochanter with strong retrolateral apophysis with large modified hair at its tip and rounded subdistal process prolaterally; femur large, with distinct dorsal process and two small ventral processes; procursus simple compared to most congeners ( Figs 67–68 View FIGURES 65–72 ), with dorsal and prolateral processes distally; genital bulb as in Figs 76–78 View FIGURES 73–78 , partly whitish, partly sclerotized and brown, with three processes: conical membranous process (putative embolus), small sclerotized process close to proximal bulbal sclerite, and large mostly sclerotized main bulbal process (‘appendix’).

LEGS. Without spines and curved hairs, few vertical hairs; retrolateral trichobothrium on tibia 1 at 5.5%; prolateral trichobothrium absent on tibia 1; tarsus 1 with many pseudosegments but only distally ~10 distinct.

Male (variation). Tibia 1 in 2 other males: 14.5, 14.7. Carapace and sternum marks as in holotype, abdominal pattern variably distinct.

Female. In general similar to male; tibia 1 in 4 females: 11.1, 12.0, 12.3, 13.9. Epigynum as in Fig. 86 View FIGURES 85–88 , with large sclerotized area (pre-epigynal plate) in front of small epigynal plate; the latter in circular excavation of preepigynal plate and provided with finger-shaped median process (‘knob’); width of circular excavation: 0.17–0.18. Internal genitalia as in Fig. 90 View FIGURES 89–92 , with pair of oval pore plates.

Natural history. This species was very abundant among large rocks at the type locality. Webs and behavior at disturbance as in P. ceylonicus .

Distribution. Known from type locality only ( Fig. 223 View FIGURES 222–223 ).

ZFMK

Zoologisches Forschungsmuseum Alexander Koenig

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Pholcidae

Genus

Pholcus

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