Batrisini, Reitter, 1882

Yin, Zi-Wei, 2022, The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth’s “ Third Pole ” (Coleoptera, Staphylinidae, Pselaphinae), Zootaxa 5111 (1), pp. 1-211 : 8-9

Key to Tibetan genera of Batrisini

1 Whole dorsal surface of body bears coarse punctation; each elytron with four basal foveae; aedeagus simple, plate-like ( Figs 52–61 )............................................................................... Sathytes Westwood

- Dorsal surface of body finely punctate to glabrous, or only parts of body coarsely punctate; each elytron with one to three basal foveae; aedeagus usually composed of median and dorsal lobes................................................. 2

2 Each elytron with two basal foveae; pronotum with transverse antebasal sulcus connecting lateral antebasal foveae; aedeagus with articulated dorsal lobe............................................................................. 3

- Each elytron with one or three basal foveae; pronotum lacking transverse antebasal sulcus; aedeagus with partially fixed dorsal lobe................................................................................................ 6

3 Tergite 1 (IV) with pair of oblique inner marginal carinae ( Fig. 76 ).................................. Trisinus Raffray

- Tergite 1 (IV) lacking inner marginal carinae............................................................... 4

4 Aedeagus with small, constricted basal capsule ( Fig. 29 )..................................... Batriscenaulax Jeannel

- Aedeagus with large or extended basal capsule.............................................................. 5

5 Scape lacking inner glandular structure at apicolateral margin; dorsal lobe of aedeagus erect ( Figs 2–28 )................................................................................................... Arthromelodes Jeannel

- Scape with inner glandular structure at apicolateral margin; dorsal lobe of aedeagus recumbent or strongly bent at base ( Figs 30–37 )................................................................................ Batrisiella Raffray

6 Head and pronotum strongly transverse; each elytron with one basal fovea ( Yin & Li 2015a: figs 1, 3)..................................................................................................... Hingstoniella Jeannel

- Head at most moderately transverse; pronotum usually with length and width subequal; each elytron with three basal foveae 7

7 Head lacking carina or sulcus; antennomeres moniliform; pronotum lacking discal or lateral longitudinal sulcus/carina..... 8

- Head carinate or sulcate; antennomeres non-moniliform; pronotum with discal or lateral longitudinal sulcus/carina........ 9

8 Body glabrous; head sub-trapezoidal, with trichome at posterolateral margin ( Yin et al. 2012b: fig. 1; Yin & Li 2013a: fig. 1)....................................................................................... Tangius Yin & Li

- Body pubescent; head sub-rectangular, lacking trichome at posterolateral margin ( Yin 2021: fig. 1)....................................................................................................... Myrmicophila Yin & Li

9 Pronotum with pairs of sinuate discal and lateral longitudinal sulci; metatibia of male with long pencil of setae at apex ( Figs 38 , 39 )................................................................................... Batrisodes Reitter

- Pronotum at most with one pair of longitudinal sulci, often with pair of discal longitudinal carinae; metatibia lacking or with short tuft of setae at apex.............................................................................. 10

10 Pronotum with pairs of anterolateral and antero-hypomeral foveae ( Fig. 40E ).................. Coryphomimus gen. nov.

- Pronotum lacking anterolateral or antero-hypomeral foveae................................................... 11

11 Elytra with inner two basal foveae close, disc coarsely punctate ( Figs 43–45 ).................... Coryphomodes Jeannel

- Elytra with inner two basal foveae clearly separated, disc finely punctate........................................ 12

12 Body length less than 1.7 mm; vertex strongly convex; lacking lateral frontal fovea; pronotum with short discal longitudinal carina present only at basal 1/2; lacking external male sexual characters ( Figs 46–48 )................. Gyrongita gen. nov.

- Body length usually no less than 2.0 mm; lateral frontal fovea present; vertex weakly to moderately convex; pronotum often with complete discal carina; male sexual characters usually present on antennae or legs, rarely on head ( Figs 62–74 )............................................................................................ Tribasodites Jeannel

Figures

FIGURE 2. Morphology of Arthromelodes alpitorus sp. nov. (A–H. Male. I. Female). A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter. D. Mesotibia. E. Tergite 1 (IV). F. Sternite 7 (IX). G, H. Aedeagus, lateral (G) and ventral (H). I. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C–E; 0.1 mm in F–I.

FIGURE 3. Morphology of Arthromelodes angulatus sp. nov. (A–G. Male. H. Female). A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter. D. Tergite 1 (IV). E. Sternite 7 (IX). F, G. Aedeagus, lateral (F) and ventral (G). H. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, D; 0.1 mm in E–H.

FIGURE 4. Morphology of Arthromelodes aniqiao sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter and mesofemur. D. Tergite 1 (IV). E. Sternite 7 (IX). F, G. Aedeagus, lateral (F) and ventral (G). Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in D, F, G; 0.1 mm in E.

FIGURE 5. Morphology of Arthromelodes bicolor sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Mesotibia. D. Tergite 1 (IV). E. Sternite 7 (IX). F, G. Aedeagus, lateral (F) and ventral (G). Scale bars: 0.5 mm in A; 0.3 mm in B, D; 0.2 mm in C, F, G; 0.1 mm in E.

FIGURE 6. Morphology of Arthromelodes championi (Jeannel) comb. nov. (A–G. Male. H. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 7–11. D. Mesotibia. E. Sternite 7 (IX). F, G. Aedeagus, lateral (F) and ventral (G). H. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, D, F, G; 0.1 mm in E, H.

FIGURE 7. Morphology of Arthromelodes complexus sp. nov. (A–I. Male. J. Female). A. Dorsal habitus. B. Head and pronotum. C. Protibia. D. Mesotrochanter. E. Mesotibia. F. Tergite 1 (IV). G. Sternite 7 (IX). H, I. Aedeagus, lateral (H) and ventral (I). J. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C–F; 0.1 mm in G–J.

FIGURE 8. Morphology of Arthromelodes cona sp. nov. (A–H. Male. I. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Mesotrochanter. E. Mesotibia. F. Sternite 7 (IX). G, H. Aedeagus, dorsal (G) and lateral (H). I. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, E; 0.2 mm in C, G, H; 0.1 mm in D, I, F.

FIGURE 9. Morphology of Arthromelodes crassicornis sp. nov. (A–H. Male. I, J. Female). A. Dorsal habitus. B. Head and pronotum. C, I. Antennomeres 9–11. D. Metaventrite and sternite 2 (IV). E. Mesotibia. F. Sternite 7 (IX). G, H. Aedeagus, dorsal (F) and lateral (G). J. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, D; 0.2 mm in C, E, G–J; 0.1 mm in F.

FIGURE 10. Morphology of Arthromelodes criniger sp. nov. (A–I. Male. J. Female). A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter. D. Apex of protibia. E. Metatrochanter. F. Tergite 1 (IV). G. Sternite 7 (IX). H, I. Aedeagus, lateral (H) and ventral (I). J. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, F; 0.1 mm in D, E, G, H–J.

FIGURE 11. Morphology of Arthromelodes cylindricus sp. nov. (A–G. Male. H. Female). A. Dorsal habitus. B. Head and pronotum. C. Metafemur. D. Tergite 1 (IV). E. Sternite 7 (IX). F, G. Aedeagus, dorsal (F) and lateral (G). H. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, D; 0.1 mm in C, E, H; 0.2 mm in F, G.

FIGURE 12. Morphology of Arthromelodes flosculus sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Tergite 1 (IV). D. Sternite 7 (IX). E, F. Aedeagus, lateral (E) and ventral (F). Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in E; 0.2 mm in E, F.

FIGURE 13. Morphology of Arthromelodes gyamda sp. nov. (A–H. Male. I. Female). A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter and mesofemur. D. Tergites 1–4 (IV–VII). E. Sternite 7 (IX). F–H. Aedeagus, dorsal (F), lateral (G) and ventral (H). I. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.2 mm in D, F–H; 0.1 mm in E, I.

FIGURE 14. Morphology of Arthromelodes intricatus sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter. D. Metatrochanter. E. Tergite 1 (IV). F. Sternite 7 (IX). G, H. Aedeagus, lateral (G) and ventral (H). Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.2 mm in C–E; 0.1 mm in F–H.

FIGURE 15. Morphology of Arthromelodes lage sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter and mesofemur. D. Tergite 1 (IV). E. Sternite 7 (IX). F, G. Aedeagus, lateral (F) and ventral (G). Scale bars: 0.5 mm in A; 0.3 mm in B, D; 0.2 mm in C, F, G; 0.1 mm in E.

FIGURE 16. Morphology of Arthromelodes langjicuo sp. nov. (A–H. Male. I. Female). A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter. D. Apical part of mesotibia. E. Tergite 1 (IV). F. Sternite 7 (IX). G, H. Aedeagus, lateral (G) and ventral (H). I. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.1 mm in C, D, F–I; 0.2 mm in E.

FIGURE 17. Morphology of Arthromelodes latithorax sp. nov. (A–H. Male. I. Female). A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter and mesofemur. D. Mesotibia. E. Sternite 7 (IX). F–H. Aedeagus, dorsal (F), lateral (G) and ventral (H). I. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B–D; 0.1 mm in E, I; 0.2 mm in F–H.

FIGURE 18. Morphology of Arthromelodes lebus sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 7–11. D. Mesotibia. E. Sternite 7 (IX). F, G. Aedeagus, lateral (F) and ventral (G). Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, D; 0.05 mm in E; 0.1 mm in F, G.

FIGURE 19. Morphology of Arthromelodes lotus sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter. D. Apical part of mesotibia. E. Tergite 1 (IV). F. Sternite 7 (IX). G, H. Aedeagus, lateral (G) and ventral (H). Scale bars: 0.5 mm in A; 0.3 mm in B; 0.1 mm in C, D, G, H; 0.2 mm in E; 0.05 mm in F.

FIGURE 20. Morphology of Arthromelodes markam sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Mesotibia. D. Tergite 1 (IV). E. Sternite 7 (IX). F, H. Aedeagus, lateral (F) and ventral (G). Scale bars: 0.5 mm in A; 0.3 mm in B–D; 0.1 mm in E; 0.2 mm in F, G.

FIGURE 21. Morphology of Arthromelodes monba sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter and mesofemur. D. Mesotibia. E. Tergite 1 (IV). F. Sternite 7 (IX). G, H. Aedeagus, lateral (G) and ventral (H). Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C–E; 0.05 mm in F, 0.2 mm in G, H.

FIGURE 22. Morphology of Arthromelodes nepaeformis sp. nov. (A–H. Male. I. Female). A. Dorsal habitus. B. Head and pronotum. C. Metaventrite. D. Protibia. E. Apical part of mesotibia. F. Sternite 7 (IX). G–H. Aedeagus, ventral (G) and lateral (H). I. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C; 0.1 mm in D, E, G–I, 0.05 mm in F.

FIGURE 23. Morphology of Arthromelodes nomurai sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Mesotibia. D. Sternites 1–5 (III–VII). E. Sternite 7 (IX). F–H. Aedeagus, dorsal (F), lateral (G) and ventral (H). Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, D, F–H; 0.05 mm in E.

FIGURE 24. Morphology of Arthromelodes songxiaobini sp. nov. (A–G. Male. H. Female). A. Dorsal habitus. B. Head and pronotum. C. Mesotibia. D. Tergites 4 and 5 (VII and VIII). E. Sternite 7 (IX). F, G. Aedeagus, lateral (F) and ventral (G). H. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, D, F, G; 0.05 mm in E, 0.1 mm in H.

FIGURE 25. Morphology of Arthromelodes speciosus sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Sternite 7 (IX). E, F. Aedeagus, lateral (F), lateral (E) and ventral (F). Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, E, F; 0.1 mm in D.

FIGURE 26. Morphology of Arthromelodes torus sp. nov. (A–H. Male. I. Female). A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter. D. Mesotibia. E. Tergite 1 (IV). F. Sternite 7 (IX). G, H. Aedeagus, lateral (G) and ventral (H). I. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C–E; 0.1 mm in F–I.

FIGURE 27. Morphology of Arthromelodes zhangmu sp. nov. (A–H. Male. I. Female). A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter. D. Mesotibia. E. Metatrochanter. F. Sternite 7 (IX). G, H. Aedeagus, dorsal (G) and lateral (H). I. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C–E; 0.05 mm in F, I, 0.1 mm in G, H.

FIGURE 28. Morphology of Arthromelodes zhentangensis sp. nov. (A–G. Male. H. Female). A. Dorsal habitus. B. Head and pronotum. C. Protibia. D. Tergite 1 (IV). E. Sternite 7 (IX). F, G. Aedeagus, lateral (F) and ventral (G). H. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.1 mm in C, F–H; 0.2 mm in D, 0.05 mm in E.

FIGURE 29. Morphology of Batriscenaulax campestris sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Apical part of protibia. D. Apical part of mesotibia. E. Tergite 1 (IV). F. Sternite 7 (IX). G, H. Aedeagus, ventral (G) and lateral (H). Scale bars: 0.5 mm in A; 0.3 mm in B; 0.1 mm in C, D, G, H; 0.2 mm in E, 0.05 mm in F.

FIGURE 30. Morphology of Batrisiella acclivis sp. nov. (A–H. Male. I. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomere 1. D. Apical part of profemur. E. Apical part of mesotibia. F. Sternite 7 (IX). G, H. Aedeagus, anterior (G) and lateral (H). I. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.1 mm in C–E, G–I; 0.05 mm in F.

FIGURE 31. Morphology of Batrisiella beibeng sp. nov. (A–J. Male. K. Female). A. Dorsal habitus. B. Head and pronotum. C. Profemur. D. Apical part of profemur, enlarged. E. Metatibia. F. Sternite 7 (IX). G–J. Aedeagus, dorsal (G), lateral (H), ventral (I) and anterior (J). K. Genitalia. Scale bars: 0.5 mm in A; 0.2 mm in B, C, E; 0.05 mm in F; 0.1 mm in G–K.

FIGURE 32. Morphology of Batrisiella bomea sp. nov. (A–F. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 8–11. D. Sternite 7 (IX). E, F. Aedeagus, anterior (E) and lateral (F). G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, E, F; 0.1 mm in D, G.

FIGURE 33. Morphology of Batrisiella concisa sp. nov. (A–H. Male. I. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomere 1. D. Tergite 1 (IV). E. Sternite 7 (IX). F–H. Aedeagus, dorsal (F), lateral (G) and ventral (H). I. Genitalia. Scale bars: 0.5 mm in A; 0.2 mm in B, D; 0.05 mm in C; 0.1 mm in E–I.

FIGURE 34. Morphology of Batrisiella curvitibialis sp. nov. (A–J. Male. K. Female). A. Dorsal habitus. B. Head and pronotum. C. Metaventrite. D. Mesotibia. E. Metafemur. F. Metatibia. G. Tergite 4 (VII). H. Sternite 7 (IX). I, J. Aedeagus, anterior (I) and lateral (J). K. Genitalia. Scale bars: 0.5 mm in A; 0.2 mm in C–G; 0.05 mm in H; 0.1 mm in I–K.

FIGURE 35. Morphology of Batrisiella ganma sp. nov. (A–F. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Tergite 1 (IV). D. Sternite 7 (IX). E, F. Aedeagus, lateral (E) and ventral (F). G. Genitalia. Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.05 mm in D; 0.1 mm in E–G.

FIGURE 36. Morphology of Batrisiella guttata sp. nov. (A–J. Male. K. Female). A. Dorsal habitus. B. Head and pronotum. C, D. Tergite 5 (VIII). E, F. Sternites 2–4 (IV–VI). G. Sternite 7 (IX). H–J. Aedeagus, dorsal (H), lateral (I) and ventral (J). K. Genitalia. Scale bars: 0.5 mm in A; 0.2 mm in C–F; 0.05 mm in F; 0.1 mm in G–K.

FIGURE 37. Morphology of Batrisiella zhujianqingi sp. nov. (A–G. Male. H. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomere 1. D. Tergite 1 (IV). E. Sternite 7 (IX). F, G. Aedeagus, dorsal (F) and lateral (G). H. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.05 mm in C, E; 0.2 mm in D, F, G; 0.1 mm in H.

FIGURE 38. Morphology of Batrisodes guoguotang sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Metatrochanter and metafemur. D. Metatibia. E. Sternite 7 (IX). F, G. Aedeagus, lateral (F) and ventral (G). Scale bars: 0.5 mm in A; 0.3 mm in B–D; 0.1 mm in E; 0.2 mm in F, G.

FIGURE 39. Morphology of Batrisodes hanmi sp. nov. (A–H. Male. I. Female). A. Dorsal habitus. B. Head and pronotum. C. Mesotrochanter. D. Metatrochanter. E. Metatibia. F. Sternite 7 (IX). G, H. Aedeagus, lateral (G) and ventral (H). I. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, E; 0.1 mm in C, D, F, I; 0.2 mm in G. H.

FIGURE 40. Morphology of Coryphomimus gen. nov. A–C. Head, in dorsal (A), lateral (B) and ventral (C). D–F. Prothorax, in dorsal (D), lateral (E) and ventral (F). G. Left elytron. H. Basal half of elytron, in lateral view.Abbreviations: alf—anterolateral fovea, ahf—antero-hypomeral fovea, bef—basolateral foveae, blf—basolateral fovea, dlc—discal longitudinal carina, ds—discal stria, gc—gular carina, gf—gular fovea, hr—hypomeral ridge, lpf—lateral procoxal fovea, laf—lateral antebasal fovea, llc—lateral longitudinal carina, mbc—mediobasal carina, mls—median longitudinal sulcus, ms—marginal stria, omc—ocularmandibular carina, shf—subhumeral fovea, vf—vertexal fovea. Scale bars: 0.2 mm.

FIGURE 43. Morphology of Coryphomodes budda sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Right elytron. D. Metatrochanter. E. Sternite 7 (IX). F, G. Aedeagus, lateral (F) and ventral (G). Scale bars: 0.3 mm in A; 0.2 mm in B, C; 0.05 mm in D, E; 0.1 mm in F, G.

FIGURE 44. Morphology of Coryphomodes cephalicus sp. nov. (A–H. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Right elytron. D. Metatrochanter. E, F. Aedeagus, lateral (E) and ventral (F). G. Genitalia. Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in D–G.

FIGURE 45. Morphology of Coryphomodes chenzhilini sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Right elytron. D. Mesotrochanter. E. Metatrochanter. F. Sternite 7 (IX). G, H. Aedeagus, lateral (G) and ventral (H). Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C–E; 0.1 mm in F–H.

FIGURE 46. Morphology of Gyrongita gen. nov. A–C. Head, in dorsal (A), lateral (B) and ventral (C). D–F. Prothorax, in dorsal (D), lateral (E) and ventral (F). G. Right elytron. H. Basal half of elytron, in lateral view. Abbreviations: bef—basolateral foveae, blf—basolateral fovea, dc—discal carina, ds—discal stria, gf—gular fovea, hr—hypomeral ridge, laf—lateral antebasal fovea, lls—lateral longitudinal sulcus, lpf—lateral procoxal fovea, mbc—mediobasal carina, mls—median longitudinal sulcus, ms—marginal stria, omc—ocular-mandibular carina, shf—subhumeral fovea, vf—vertexal fovea. Scale bars: 0.2 mm.

FIGURE 47. Morphology of Gyrongita gen. nov. A. Meso- and metaventrite. B–D. Abdomen, in dorsal (B), lateral (C) and ventral (D). Abbreviations: amsp—antero mesoventral projection, blf—basolateral fovea, lmcf—lateral mesocoxal fovea, lmsf—lateral mesoventral fovea, lmtf—lateral metaventral fovea, mbf—mediobasal fovea, mc—marginal carina, mmsf—median mesoventral fovea, s2–5—sternites 2–5 (IV–VII), t1–4—tergites 1–4 (IV–VII). Scale bars: 0.2 mm in A; 0.3 mm in B–D.

FIGURE 48. Morphology of Gyrongita uniformis sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Sternite 7 (IX). E–F. Aedeagus, lateral (E) and ventral (F). Scale bars: 0.5 mm in A; 0.2 mm in B; 0.1 mm in C–F.

FIGURE 52. Morphology of Sathytes dawai sp. nov. (A–F. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in D–G.

FIGURE 53. Morphology of Sathytes duojii sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in D–F.

FIGURE 54. Morphology of Sathytes laticornis sp. nov. (A–F. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C; 0.1 mm in D–G.

FIGURE 55. Morphology of Sathytes pengzhongi sp. nov. (A–F. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C; 0.05 mm in D, 0.1 mm in E–G.

FIGURE 56. Morphology of Sathytes proclivis sp. nov. (A–F. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in D–G.

FIGURE 57. Morphology of Sathytes pseudograndis sp. nov. (A–F. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in D–G.

FIGURE 58. Morphology of Sathytes simplex Löbl, male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Sternite 7 (IX). E. Aedeagus, ventral. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, E; 0.1 mm in D.

FIGURE 59. Morphology of Sathytes sympatricus sp. nov. (A–F. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, C; 0.1 mm in D–G.

FIGURE 60. Morphology of Sathytes virupaksa sp. nov. (A–F. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C; 0.1 mm in D–G.

FIGURE 61. Morphology of Sathytes yigongensis sp. nov. (A–F. Male. G. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Antennomere 9. E. Sternite 7 (IX). F. Aedeagus, ventral. G. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C; 0.1 mm in D–G.

FIGURE 62. Morphology of Tribasodites bari sp. nov. (A–G, I. Male. H. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Metatrochanter. E. Sternite 7 (IX). F, G. Aedeagus, lateral (F) and ventral (G). H. Genitalia. I. Laboulbeniales on right femur. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, F, G; 0.1 mm in D; 0.05 mm in E, H, I.

FIGURE 63. Morphology of Tribasodites cellulanus sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Protrochanter. E. Mesotrochanter. F. Metatrochanter. G. Sternite 7 (IX). H, I. Aedeagus, lateral (H) and ventral (I). Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, H, I; 0.1 mm in D–G.

FIGURE 64. Morphology of Tribasodites constrictus sp. nov. (A–I. Male. J. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Mesotrochanter and base of mesofemur. E. Apex of mesotibia. F. Metatrochanter. G. Sternite 7 (IX). H, I. Aedeagus, lateral (H) and ventral (I). J. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, H, I; 0.1 mm in D–G, J.

FIGURE 65. Morphology of Tribasodites dilophus sp. nov. (A–H. Male. I. Female). A. Dorsal habitus. B. Head and pronotum. C. Pronotum, in dorsolateral view. D. Mesotrochanter. E. Metatrochanter. F. Sternite 7 (IX). G, H. Aedeagus, lateral (G) and ventral (H). I. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, G, H; 0.1 mm in D–F, I.

FIGURE 66. Morphology of Tribasodites elongatus sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C, D. Antennomeres 9–11, in dorsal (C) and ventral view (D). E. Protrochanter and profemur. F. Protibia. G. Mesotrochanter and mesofemur. H. Metatrochanter and metafemur. I. Tergite 5 (VIII). J. Sternite 6 (VIII). K. Sternite 7 (IX). L–N. Aedeagus, dorsal (L), lateral (M) and ventral (N). Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C–J, L–N; 0.1 mm in K.

FIGURE 67. Morphology of Tribasodites grandiceps sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Pronotum, in dorsolateral view. D. Apex of mesotibia. E. Metatrochanter. F. Metafemur. G. Sternite 7 (IX). H–J. Aedeagus, dorsal (H), lateral (I) and ventral (J). Scale bars: 0.5 mm in A; 0.3 mm in B, F; 0.2 mm in C, 0.1 mm D, E, H–J; 0.05 mm in G.

FIGURE 68. Morphology of Tribasodites gyirong sp. nov. (A–J. Male. K. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Antennomere 10, ventral. E. Mesotrochanter. F. Mesotibia. G. Metatrochanter. H. Sternite 7 (IX). I, J. Aedeagus, lateral (I) and ventral (J). K. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B, F; 0.2 mm in C, E, G, H–K; 0.1 mm in D.

FIGURE 69. Morphology of Tribasodites kiypu sp. nov. (A–G. Male. H. Female). A. Dorsal habitus. B. Head and pronotum. C. Pronotum, in dorsolateral view. D. Mesotrochanter. E. Metatrochanter. F, G. Aedeagus, ventral (F) and lateral (G). H. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C; 0.1 mm in D–H.

FIGURE 70. Morphology of Tribasodites mirabilis sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Pronotum, in dorsolateral view. D. Metatrochanter. E. Metafemur. F. Metatibia. G. Sternite 7 (IX). H–J. Aedeagus, dorsal (H), lateral (I) and ventral (J). Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, E, F; 0.1 mm D, G–J.

FIGURE 71. Morphology of Tribasodites prolixicornis sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Pronotum, in dorsolateral view. D. Apex of mesotibia. E. Metatrochanter. F. Metafemur. G. Sternite 7 (IX). H–J. Aedeagus, dorsal (H), lateral (I) and ventral (J). Scale bars: 0.5 mm in A; 0.3 mm in B, F; 0.2 mm in C; 0.1 mm D, E, H–J, 0.05 mm in G.

FIGURE 72. Morphology of Tribasodites pugiunculus sp. nov. (A–J. Male. K. Female). A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Antennomere 10, ventral. E. Mesotrochanter. F. Apical part of mesotibia. G. Metatrochanter. H. Sternite 7 (IX). I, J. Aedeagus, lateral (I) and ventral (J). K. Genitalia. Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, I, J; 0.1 mm in D–H, K.

FIGURE 73. Morphology of Tribasodites vertexalis sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Antennomeres 9–11. D. Mesotrochanter. E. Mesotibia. F. Metatrochanter. G–I. Aedeagus, ventral (G), lateral (H) and anterior (I). Scale bars: 0.5 mm in A; 0.3 mm in B; 0.2 mm in C, E; 0.1 mm in D, F–I.

FIGURE 74. Morphology of Tribasodites yatung sp. nov., male. A. Dorsal habitus. B. Head and pronotum. C. Pronotum, in dorsolateral view. D. Mesotrochanter. E. Apex of mesotibia. F. Metatrochanter. G. Sternite 7 (IX). H–J. Aedeagus, dorsal (H), lateral (I) and ventral (J). Scale bars: 0.5 mm in A; 0.3 mm in B, F; 0.2 mm in C; 0.1 mm D–F, H–J, 0.05 mm in G.

FIGURE 76. Morphology of Trisinus sp., female. A. Dorsal habitus. B. Head and pronotum. C. Apex of abdomen. D. Tergite 5 (VIII). E. Sternite 6 (VIII). F. Genitalia. Scale bars: 0.5 mm in A; 0.2 mm in B; 0.1 mm in C–F.