Sussaba roberti, Klopfstein, Seraina, 2014

Klopfstein, Seraina, 2014, Revision of the Western Palaearctic Diplazontinae (Hymenoptera, Ichneumonidae), Zootaxa 3801 (1), pp. 1-143 : 92-93

Sussaba roberti sp. nov.

Etymology. This species is dedicated to my grandfather and talented scientist Prof. Robert Schenk, who introduced me to the scientific world.

Diagnosis. Fore wing length 3.5–4.1 mm. Antenna in females with 16, very rarely 15 flagellomeres, with multiporous plate sensilla present also ventrally on apical flagellomeres, although slightly sparser there than dorsally, without conspicuous velvety areas; in males with 18–19 flagellomeres, with uni-coloured tyloids on flagellomeres 5 to 9/10, first tyloid often distinctly smaller than second. Face mainly smooth and shining and strongly punctate over entire surface. Mesoscutum smooth and shining, in both sexes evenly punctate also over entire surface. Mesopleuron smooth and shining, distinctly punctate over most of its surface. Hind coxa with outer hind surface mostly smooth and shining, without any sculpture apart from around seta roots. Propodeum with a full set of carinae enclosing basal, lateral and petiolar areas, the areas with rugose sculpture. Metasoma compressed posterior to second segment, usually knife-like, first tergite 1.3–1.4 times longer than wide in females, 1.2–1.4 in males.

Description. Antenna with multiporous plate sensilla evenly distributed also on ventral surface of flagellomeres. Face centrally elevated, with very short and weak vertical depressions, a little protruding below but less so than S. erigator ; smooth and shining, with numerous distinct punctures. Clypeus separated from face by a distinct groove, elevated basally, remainder flat, very weakly bilobed, partly smooth, partly coriaceous, with some weak longitudinal striae. Head strongly constricted behind compound eyes. Mesoscutum smooth and shining between strong punctures, often punctures a bit less dense towards sides, without notauli; scutellum smooth and sparsely punctate, with lateral carinae clearly surpassing middle. Mesopleuron smooth and shining and distinctly punctate over entire surface except for a smooth area around mesopleural fovea; sternaulus weakly impressed, epicnemial carina complete ventrally. Metapleuron smooth and sparsely to densely punctate. Propodeum with a full set of strong carinae; areas between carinae rugose. Fore wing areolet open, vein 1cu-a opposite vein M; hind wing with one basal hamulus, vein CU+cu-a broken slightly below middle. Hind coxa smooth and punctate. Metasoma strongly compressed laterally from tergite 3, tergite 2 strongly tapering, almost triangular; hind margins of tergites 1–3 straight or convex, from tergite 4 concave, extending further back laterally than medially; tergite 1 smooth, with longitudinal carinae distinct on less than basal half, then replaced by some longitudinal rugae and sparse punctures; tergite 2 finely striate basally and punctate on about half the length, remainder entirely smooth, remaining tergites entirely smooth and shining. Second and third tergites with spiracles on laterotergites, distinctly below lateral folds. Ovipositor sheaths upcurved, slender, with tip closed apically; smooth and shining, with setae around tip and ventrally.

Colouration of females. Antenna dark brown, usually paler below. Head and mesosoma black, face sometimes with a yellow central face patch, clypeus black, yellow or orange on mouthparts, hind corner of pronotum, tegula, sometimes upper mesepimeron; subtegular ridge and scutellum black, without yellow shoulder marks. Legs orange, coxae and trochanters black; femora orange, sometimes marked with black below; hind tibia and tarsus orange to brown. Metasoma variously marked with orange, usually at least on hind margin of tergite 2 and entire tergites 3 and sometimes 4; this colouration can be dark.

Colouration of males. As in females but with yellow ventrally on entire antenna, entire face but sometimes with some black markings on face, often subtegular ridge. Fore and mid coxae with yellow apices, fore and mid trochanters often entirely yellow.

Similar species: Sussaba roberti is very close to S. erigator and has in the past been mistaken for the latter species. Females are easily separable by the longer antennae, the less projecting lower face, the evenly punctate mesoscutum and the more compressed metasoma. Males are more difficult to distinguish and the characters given in the key might not always be reliable, but male S. roberti usually have more tyloids, on flagellomeres 5 to 9 instead of 5 to 8 as in most erigator males, and the first tyloid is often distinctly smaller than the second.

Type material. Holotype ♀ at NMBE, Switzerland, Grisons, Sur, NE Sur, 1770m, N46°31.451’ E9°38.055’, Malaise trap, leg. H.Baur & S.Klopfstein, 19.–27. VII. 2006. Labels: “Malaise-F. 3, FO-Nr. 834; CH/GR Sur, NE Sur; 1770m 768.450/154.500; leg. H.Baur; 19.– 27.7.2006 ”, “Extraction-Nb; 1-B9 ”, “Holotype; Sussaba ♀; roberti n.sp.; det. S.Klopfstein 2010 ” . Paratypes: CH, Luzern, Allmend and city, 10.VI.2006, leg. G. Bächli (1♂); at NMBE ; CH, Schaffhausen, Bargemer Randen , 600–800m, 26.VI.1975, leg. W. Sauter; at ETHZ ; UA, Lugansk reg., Provalie, leg. Osipov, 23.V.2002 (1♀), 19.–22.VII.2004 (1♂), 25.VIII.2004 (1♂) ; RU, Siberia, Irkutsk, Lisikha, leg. O. Berlov, 28.VIII.2003 (1♂), 17.VIII.2004 (1♀), 4.VIII.2005 (1♀), 2.VIII.2007 (1♀) ; RU, Siberia, 10 km E Irkutsk, leg. O. Berlov, 12.VIII.2004 (1♂), all at MR .

Additional material. Austria: Neusiedler See, Donnerskirchen, leg. Wachmann, 6.VIII.1964. 1♀, at KSch . France: Hautes-Alpes, Briançon, 1400m, leg. K.Zwakhals, 1.VIII.1971. 1♂, at KZ . Germany: Karlstadt i.M., leg. K.Schmidt, 4.VIII.1966. 1♀, at KSch ; Rheinland-Pfalz, Büchelberg, Sägewerk, leg. K.Schmidt, 6.VIII.1997. 1♀, at KSch ; Karlsruhe, Baden, Durlach, Bergwald, leg. Zmuszinski, 14.VII.1967. 18.VII.1967 1♀ each, at KSch. Italy: Bolzano, Brixen, 1500m, leg. K.Zwakhals, 23.VI.1976. 1♂, at KZ . Macedonia: Gorica, 5 km S. Ohrid, ca. 800m, leg. C.v.Achterberg, 13.IX.1979. 1♂, at KZ . Switzerland: Geneva, Russin , 400m, N46°11.16', E6°00.83', leg. J.de Beaumont, 26.VII.1959. 1♀, at MZL ; Vaud, Eclépens , Mormont , 500m, N46°39.06', E6°32.232', leg. J.de Beaumont, 6.VII.1958. 1♀, at MZL ; Vaud, Cudrefin , La Sauge , 433m, N46°58.44', E7°03.252', leg. J.de Beaumont, 17.VIII.1957. 1♀, at MZL . Turkey: Isparta, Dere Mahallesi, 1150m, leg. F. Gürbüz, 22.V.2006. 1♂, at SDU ; Isparta, Egridir, Kasnakmesesi, 1350m, leg. F. Gürbüz, 7.VI.2004. 3♂, 1♀, at SDU.

Distribution. Palaearctic.

Figures. Metasoma ♀ ( Fig. 23 B), habitus ( Fig. 34 C), male terminal sclerites ( Fig. 38 C).

Figures

FIGURE 23. Metasoma of Sussaba species viewed from above. A. Sussaba erigator. B. Sussaba roberti. C. Sussaba dorsalis. D. Sussaba pulchella. Scale bars represent 500 µm.

FIGURE 34. Habitus of Diplazontinae species. A. Sussaba montana. B. Sussaba pulchella. C. Sussaba roberti sp. nov. D. Syrphoctonus desvignesii. E. Syrphoctonus fissorius. F. Syrphoctonus tarsatorius.

FIGURE 38. Ninth sternite and ninth and tenth tergites of male Diplazontinae species. A. Sussaba erigator. B. Sussaba pulchella. C. Sussaba roberti. D. Syrphophilus asperatus. E. Tymmophorus erythrozonus. F. Tymmophorus obscuripes. G. Woldstedtius bauri. H. Woldstedtius citropectoralis. I. Woldstedtius holarcticus. J. Xestopelta gracillima.