Mycotretus Lacordaire, 1842

Pecci-Maddalena, Italo Salvatore de Castro, Lopes-Andrade, Cristiano & Skelley, Paul, 2023, Catalogue of Mycotretus Lacordaire, 1842 (Coleoptera: Erotylidae: Tritomini): an annotated, illustrated and historical approach, European Journal of Taxonomy 876 (1), pp. 1-182 : 11-16

Genus Mycotretus Lacordaire, 1842

Mycotretus Lacordaire, 1842: 132 . Type species: Erotylus lesueuri Chevrolat, 1835 . Designated by Boyle (1956), fixed by Skelley & Goodrich (1994); ICZN (1996).

Mycotretus Chevrolat in Dejean 1836: 428.

Mycotretus Lacordaire – Crotch 1873b: 144. — Gemminger & Harold 1876: 3691. — Gorham 1888: 46. — Deelder 1942: 54. — Mader 1951: 210. — Guerin 1953: 77. — Boyle 1956: 137.

Mycotretus Chevrolat, Dejean – Lacordaire 1842: 132. — Crotch 1873a: 354; 1876: 437. — Chapuis 1876: 32. — Kuhnt 1909: 70 [“ Mycotretus Dejean ”]. — Casey 1916: 158. — Alvarenga 1965: 87; 1994: 20. — Skelley & Goodrich 1994: 130. — ICZN 1996: 55 [suppression of the names Mycotretus Chevrolat and Mycotretus Dejean ].

Included species

Mycotretus adalioides Crotch, 1876 ( Fig. 4A )

Mycotretus aegrotus Gorham, 1888 ( Fig. 14E )

Mycotretus aestuans Lacordaire, 1842 ( Fig. 26C )

Mycotretus alvarengai Pecci-Maddalena & Lopes-Andrade, 2018 ( Fig. 39G )

Mycotretus ambulator Lacordaire, 1842 ( Fig. 26D )

Mycotretus anchoralis Guérin, 1956 ( Fig. 21D )

Mycotretus arcuatus Lacordaire, 1842

Mycotretus argus Lacordaire, 1842 ( Fig. 26F )

Mycotretus atricaudatus Gorham, 1888 ( Fig. 15A )

Mycotretus badius Gorham, 1888 ( Fig. 15B )

Mycotretus balteatus Crotch, 1876 ( Fig. 4B )

Mycotretus bicolor Taschenberg, 1870 ( Fig. 40A )

Mycotretus monrosi Guérin,1949 ( Fig. 21F ) syn. nov.

Mycotretus bicoloratus Kuhnt, 1911 ( Fig. 23E )

Mycotretus bipunctatus Gorham, 1888 ( Fig. 15C )

Mycotretus bistrigatus Lacordaire, 1842 ( Fig. 27A )

Mycotretus bistrioculatus Alvarenga, 1983 ( Fig. 2A )

Mycotretus brasilianus Crotch, 1876 ( Fig. 4E )

Mycotretus brevis Gorham, 1888 ( Fig. 15D )

Mycotretus centralis Arrow, 1909 ( Fig. 3B )

Mycotretus cercyonoides Gorham, 1888 ( Fig. 15E )

Mycotretus chilensis Crotch, 1876 ( Fig. 4F )

Mycotretus bicinctus Guérin, 1949a ( Fig. 21E )

Mycotretus cinctiger Crotch, 1876 ( Fig. 5B )

Mycotretus luizi Alvarenga, 1983 ( Fig. 2D ) syn. nov.

Mycotretus clitelliger Lacordaire, 1842 ( Fig. 27B )

Mycotretus coccidulinus Gorham, 1888 ( Fig. 15F )

Mycotretus coelestinus Lacordaire, 1842 ( Fig. 27D )

Mycotretus consanguineus Gorham, 1888 ( Fig. 16A )

Mycotretus cordiger Crotch, 1876 ( Fig. 5D )

Mycotretus zischkai Delkeskamp, 1957 ( Fig. 12F ) syn. nov.

Mycotretus cribratus Gorham, 1888 ( Fig. 16B )

Mycotretus cruciger Crotch, 1876 ( Fig. 5E )

Mycotretus crudus Gorham, 1888 ( Fig. 16C )

Mycotretus cruentus Gorham, 1888 ( Fig. 16D )

Mycotretus cunctans Mader, 1942 ( Fig. 38B )

Mycotretus cyanopterus Lacordaire, 1842 ( Fig. 27F )

Mycotretus decoratus ( Duponchel, 1825) ( Fig. 13B )

Mycotretus decorus Crotch, 1876 ( Fig. 5F )

Mycotretus deyrollei Crotch, 1876 ( Fig. 6A )

Mycotretus discipennis discipennis Kuhnt, 1910 ( Fig. 24A ) syn. nov.

Mycotretus discipennis conductus Kuhnt, 1910 ( Fig. 24B ) syn. nov.

Mycotretus sexlineatus Kuhnt, 1910 ( Fig. 25D ) syn. nov.

Mycotretus dichrous Kirsch, 1876 ( Fig. 22F )

Mycotretus dimidiatus Taschenberg, 1870 ( Fig. 40C )

Mycotretus distigma Lacordaire, 1842 ( Fig. 28B )

Mycotretus distinguendus Arrow, 1909 ( Fig. 3C )

Mycotretus dorsofasciatus Lacordaire, 1842 ( Fig. 28C )

Mycotretus nugator Lacordaire, 1842 ( Fig. 33G ) syn. nov.

Mycotretus dorsonotatus Lacordaire, 1842 ( Fig. 28D )

Mycotretus alternans Gorham, 1888 ( Fig. 14F ) syn. nov.

Mycotretus quadristriolatus Kuhnt, 1910 ( Fig. 25C ) syn. nov.

Mycotretus duodecimguttatus duodecimguttatus ( Duponchel, 1825) ( Fig. 13C )

Mycotretus duodecimguttatus consociatus ( Kuhnt, 1910) comb. nov.

Mycotretus durius Lacordaire, 1842 ( Fig. 28F )

Mycotretus dytiscoides Lacordaire, 1842 ( Fig. 29A )

Mycotretus egae Crotch, 1876 ( Fig. 6C )

Mycotretus basalis Crotch, 1876 ( Fig. 4C ) syn. nov.

Mycotretus elegans Gorham, 1888 ( Fig. 16E )

Mycotretus episcaphoides Crotch, 1876 ( Fig. 6D )

Mycotretus episcopalis Lacordaire, 1842 ( Fig. 29B )

Mycotretus epopterus Gorham, 1888 ( Fig. 16F )

Mycotretus erraticus Gorham, 1898 ( Fig. 17A )

Mycotretus expressus Kuhnt, 1910 ( Fig. 24C )

Mycotretus fallax ( Guérin-Méneville, 1841) ( Fig. 22C )

Mycotretus fasciolatus Lacordaire, 1842 ( Fig. 29C )

Mycotretus fascipennis Kuhnt, 1910 ( Fig. 24D )

Mycotretus fidelis Delkeskamp, 1939 ( Fig. 12C )

Mycotretus flavomarginatus Lacordaire, 1842 ( Fig. 29E )

Mycotretus major Mader, 1955 ( Fig. 38D ) syn. nov.

Mycotretus floriger Lacordaire, 1842 ( Fig. 29F )

Mycotretus fulviceps Crotch, 1876 ( Fig. 6E )

Mycotretus fulvilabris Crotch, 1876 ( Fig. 6F )

Mycotretus rastratus Crotch, 1876 ( Fig. 10B ) syn. nov.

Mycotretus geminus Gorham, 1888 ( Fig. 17B )

Mycotretus guatemalae Crotch, 1876 ( Fig. 7A )

Mycotretus haemapterus Gorham, 1888 ( Fig. 17C )

Mycotretus haematicus Gorham, 1888 ( Fig. 17D )

Mycotretus hepaticus Lacordaire, 1842 ( Fig. 31A )

Mycotretus hilaris Lacordaire, 1842 ( Fig. 31B )

Mycotretus hirudo Gorham, 1888 ( Fig. 17E )

Mycotretus humeralis ( Germar, 1824)

Mycotretus humilis Lacordaire, 1842

Mycotretus illustris Crotch, 1876 ( Fig. 7B )

Mycotretus incarnatus Gorham, 1888 ( Fig. 17F )

Mycotretus interstictus Gorham, 1888 ( Fig. 18A )

Mycotretus interstitialis Kuhnt, 1910 ( Fig. 25A ) syn. nov.

Mycotretus jocosus Lacordaire, 1842 ( Fig. 31D )

Mycotretus laccophilinus Gorham, 1888 ( Fig. 18B )

Mycotretus lacertosus Lacordaire, 1842 ( Fig. 31E )

Mycotretus laeviventris Crotch, 1876 ( Fig. 7C )

Mycotretus lepidus Lacordaire, 1842 ( Fig. 31F )

Mycotretus graniformis Lacordaire, 1842 ( Fig. 30E ) syn. nov.

Mycotretus chontalesi Crotch, 1873 ( Fig. 5A ) syn. nov.

Mycotretus leprosus Lacordaire, 1842 ( Fig. 32A )

Mycotretus lesueuri ( Chevrolat, 1835) ( Fig. 3F )

Mycotretus savignyi Lacordaire, 1842 ( Fig. 36C ) syn. nov.

Mycotretus limbatus ( Lacordaire, 1842) ( Fig. 32B )

Mycotretus lissomoides Crotch, 1876 ( Fig. 7D )

M ycotretus luteipes Lacordaire, 1842 ( Fig. 32C )

Mycotretus luteolus Gorham, 1888 ( Fig. 18C )

Mycotretus maculatus ( Olivier, 1792) ( Fig. 39E )

Mycotretus figuratus Lacordaire, 1842 ( Fig. 29D )

Mycotretus mutabilis Crotch, 1876 ( Fig. 7E ) syn. nov.

Mycotretus magus Lacordaire, 1842 ( Fig. 32D )

Mycotretus marginicollis Lacordaire, 1842 ( Fig. 32E )

Mycotretus melanopterus Lacordaire, 1842 ( Fig. 32F )

Mycotretus xanthosomus Lacordaire, 1842 ( Fig. 37F )

Mycotretus melanotus Gorham, 1888 ( Fig. 18D )

Mycotretus miniatus Lacordaire, 1842 ( Fig. 33B )

Mycotretus minutus ( Duponchel, 1825) ( Fig. 13E )

Mycotretus quadrinus Lacordaire, 1842 ( Fig. 35D )

Mycotretus misellus Lacordaire, 1842 ( Fig. 33C )

Mycotretus mycetophagoides mycetophagoides Crotch, 1876 ( Fig. 7F )

Mycotretus mycetophagoides erythrocerus Crotch, 1876 ( Fig. 8A )

Mycotretus mycetophiloides mycetophiloides Crotch, 1876 ( Fig. 8B )

Mycotretus mycetophiloides careus Crotch, 1876 ( Fig. 8C )

Mycotretus nigricollis Gorham, 1888 ( Fig. 18E )

Mycotretus nigripes Gorham, 1888 ( Fig. 18F )

Mycotretus nigrivittis Lacordaire, 1842 ( Fig. 33D )

Mycotretus nigromanicatus Boyle, 1954 ( Fig. 3D )

Mycotretus nitescens Crotch, 1876 ( Fig. 8E )

Mycotretus normalis Gorham, 1888 ( Fig. 19A )

Mycotretus noterinus Gorham, 1888 ( Fig. 19B )

Mycotretus octoculatus Alvarenga, 1983 ( Fig. 2E )

Mycotretus opalescens Crotch, 1876 ( Fig. 8F )

Mycotretus pelliciens Kirsch, 1876 ( Fig. 23A ) syn. nov.

Mycotretus opalizans Mader, 1942 ( Fig. 38E )

Mycotretus oppositipunctum Gorham, 1888 ( Fig. 19C )

Mycotretus ornatus ( Duponchel, 1825) ( Fig. 14A )

Mycotretus cognatus Lacordaire, 1842 ( Fig. 27E ) syn. nov.

Erotylus coronatus Duponchel, 1825 ( Fig. 13A ) syn. nov.

Mycotretus difficilis Lacordaire, 1842 ( Fig. 28A ) syn. nov.

Mycotretus dispar Taschenberg, 1870 ( Fig. 40E ) syn. nov.

Mycotretus dubius Lacordaire, 1842 ( Fig. 28E ) syn. nov.

Mycotretus godarti Lacordaire, 1842 ( Fig. 30D )

Mycotretus graphoderus graphoderus Lacordaire, 1842 ( Fig. 30F ) syn. nov.

Mycotretus graphoderus strigipennis Kuhnt, 1910 ( Fig. 24E ) syn. nov.

Mycotretus graphoderus thoracicus Kuhnt, 1910 ( Fig. 24F ) syn. nov.

Mycotretus intermedius Lacordaire, 1842 ( Fig. 31C ) syn. nov.

Erotylus maculosus Duponchel, 1825 ( Fig. 14B )

Mycotretus melanostictus Lacordaire, 1842 ( Fig. 33A )

Erotylus nigropunctatus Duponchel, 1825 ( Fig. 13F ) syn. nov.

Mycotretus ornatus partitialis Deelder, 1942 ( Fig. 12B ) syn. nov.

Mycotretus partitialis Mader, 1940 ( Fig. 39A ) syn. nov.

Mycotretus posticus Lacordaire, 1842 ( Fig. 14A ) syn. nov.

Erotylus puncticollis Duponchel, 1825 syn. nov.

Mycotretus terminalis Lacordaire, 1842 ( Fig. 37C )

Mycotretus pallidior ( Crotch, 1876) ( Fig. 9A )

Mycotretus nigrotinctus Crotch, 1876 ( Fig. 8D )

Mycotretus palmiphilus Lacordaire, 1842 ( Fig. 34A )

Mycotretus panamanus Gorham, 1888 ( Fig. 19D )

Mycotretus parallelus Crotch, 1876 ( Fig. 9B )

Mycotretus partitus Lacordaire, 1842 ( Fig. 34B )

Mycotretus pebasensis Crotch, 1876 ( Fig. 9C )

Mycotretus pecari Lacordaire, 1842 ( Fig. 34C )

Mycotretus quattuordecimguttatus conjunctus Crotch, 1876 ( Fig. 10A ) syn. nov.

Mycotretus peruae Crotch, 1876 ( Fig. 9D )

Mycotretus pictopiceus Gorham, 1888 ( Fig. 19E )

Mycotretus planus Gorham, 1888 ( Fig. 19F )

Mycotretus polyophthalmus Lacordaire, 1842 ( Fig. 34D )

Mycotretus prioteloides Mader, 1942 ( Fig. 39B )

Mycotretus psittacus Lacordaire, 1842 ( Fig. 34E )

Mycotretus psylloboroides Crotch, 1876 ( Fig. 9E )

Mycotretus pulchellus Lacordaire, 1842 ( Fig. 34F )

Mycotretus puncticeps Kirsch, 1865 ( Fig. 23C )

Mycotretus pusillus Lacordaire, 1842 ( Fig. 35B )

Mycotretus pygmaeus Lacordaire, 1842 ( Fig. 35C )

Mycotretus quadrioculatus Alvarenga, 1983 ( Fig. 2F )

Mycotretus quadripunctatus Crotch, 1876 ( Fig. 9F )

Mycotretus quattuordecimguttatus Lacordaire, 1842 ( Fig. 35E )

Mycotretus bisseptemguttatus Crotch, 1876 ( Fig. 4D ) syn. nov.

Mycotretus reticulatus Crotch, 1876 ( Fig. 10C )

Mycotretus lopesi Alvarenga, 1989 ( Fig. 2C ) syn. nov.

Mycotretus rhodosomus Lacordaire, 1842 ( Fig. 35F )

Brachysphaenus (Iphiclus) nigromaculatus Kuhnt, 1909 ( Fig. 25B ) syn. nov.

Mycotretus rubidus Gorham, 1888 ( Fig. 20A )

Mycotretus rufilabris ( Lacordaire, 1842) ( Fig. 36A )

Mycotretus rufipennis Gorham, 1888 ( Fig. 20B )

Mycotretus sallaei Crotch, 1876 ( Fig. 10D )

Mycotretus sandicatus Gorham, 1888 ( Fig. 20C )

Mycotretus sanguineus ( Duponchel, 1825) ( Fig. 14C )

Mycotretus sannio Lacordaire, 1842 ( Fig. 36B )

Mycotretus scalaris Lacordaire, 1842 ( Fig. 36D )

Mycotretus scitulus Lacordaire, 1842 ( Fig. 36E )

Mycotretus derasofasciatus Kuhnt, 1910 ( Fig. 23F ) syn. nov.

Mycotretus nubifer Casey, 1916 ( Fig. 3E ) syn. nov.

Mycotretus sedecimguttatus ( Guérin-Méneville, 1844) ( Fig. 22D )

Mycotretus seminiger Harold, 1876 ( Fig. 6B )

Mycotretus separandus Crotch, 1876 ( Fig. 10F )

Mycotretus sericeonitens sericeonitens Crotch, 1876 ( Fig. 11A )

Mycotretus sericeonitens monticola Crotch, 1876

Mycotretus sexoculatus sexoculatus Lacordaire, 1842 ( Fig. 36F )

Mycotretus sexoculatus chaparensis Delkeskamp, 1957 ( Fig. 12D )

Mycotretus sexpunctatus Gorham, 1888 ( Fig. 20D )

Mycotretus signatellus signatellus Crotch, 1876 ( Fig. 11B )

Mycotretus signatellus imperfectus Crotch, 1876 ( Fig. 11C ) syn. nov.

Mycotretus fragosoi Alvarenga, 1983 ( Fig. 2B ) syn. nov.

Mycotretus singularis Lacordaire, 1842 ( Fig. 37B )

Mycotretus sobrinus ( Guérin-Méneville, 1841) ( Fig. 22E )

Mycotretus silaceus Lacordaire, 1842 ( Fig. 37A )

Mycotretus spadiceus Gorham, 1888 ( Fig. 20E )

Mycotretus sticticollis Lacordaire, 1842

Mycotretus stillatus Kuhnt, 1910 ( Fig. 25E )

Mycotretus stramineus Gorham, 1888 ( Fig. 20F )

Mycotretus succinctus Crotch, 1876 ( Fig. 11D )

Mycotretus suturalis Kirsch, 1876 ( Fig. 23D )

Mycotretus ternotatus Gorham, 1888 ( Fig. 21A )

Mycotretus tesserarius Lacordaire, 1842

Mycotretus thoracicus Kuhnt, 1910 ( Fig. 24F )

Mycotretus tibialis Gorham, 1888 ( Fig. 21B )

Mycotretus tigratus Lacordaire, 1842 ( Fig. 37D )

Mycotretus nigrocinctus Lacordaire, 1842 ( Fig. 33E )

Mycotretus trabeatus Lacordaire, 1842 ( Fig. 37E ) syn. nov.

Mycotretus tigrinoides Mader, 1942 ( Fig. 39C )

Mycotretus tigrinus ( Olivier, 1792) ( Fig. 39F )

Erotylus conspersus Germar,1824

Mycotretus leopardus Crotch, 1876

Mycotretus multimaculatus Taschenberg, 1870 ( Fig. 40F )

Mycotretus tigrinus pardalis Crotch, 1876

Mycotretus tigripennis Mader, 1942 ( Fig. 39D )

Mycotretus tricolor Crotch, 1876 ( Fig. 11E )

Mycotretus trifasciatus Guérin, 1956 ( Fig. 22A )

Mycotretus tucuruiensis Alvarenga, 1983 ( Fig. 3A )

Mycotretus vilis Lacordaire, 1842

Mycotretus vittatus Gorham, 1888 ( Fig. 21C )

Mycotretus xanthomelas Crotch, 1876 ( Fig. 12A )

Mycotretus ziczac ziczac Kuhnt, 1910 ( Fig. 26B )

Mycotretus ziczac serenus Delkeskamp, 1957 ( Fig. 12E )

Excluded species

Iphiclus virgatus ( Kuhnt, 1910) comb. nov.

Mycolybas antesignatus ( Mader, 1942) comb. nov.

Figures

Fig. 4. Crotch primary types (dorsal, labels). A–A’. Lectotype of Mycotretus adalioides Crotch, 1876. B–B’. Lectotype of M. balteatus Crotch, 1876. C–C’. Lectotype of M. basalis Crotch, 1876. D–D’. Lectotype of M. bisseptemguttatus Crotch, 1876. E–E’. Lectotype of M. brasilianus Crotch, 1876. F–F’. Lectotype of M. chilensis Crotch, 1876. Scale bars = 1 mm.

Fig. 14. A–C, E–F. Primary types (dorsal, labels). A–A’. Lectotype of Erotylus ornatus Duponchel [currently Mycotretus ornatus (Duponchel)]. B–B’. Lectotype of E. maculosus Duponchel, 1825 [currently M. maculosus (Duponchel)]. C–C’. Lectotype of E. sanguineus Duponchel, 1825 [currently M. sanguineus (Duponchel)]. D–D’. Topotype of M. unicolor Fauvel, 1860 from RBINS (Belgium) [synonym of Mycomystes coccineus (Lacordaire, 1842)]. E–E’. Lectotype of M. aegrotus Gorham, 1888. F–F’. Lectotype of M. alternans Gorham, 1888. Scale bars = 1 mm.

Fig. 26. A–B, D–F. Primary types (dorsal, labels). A–A’. Lectotype of Mycotretus virgatus Kuhnt, 1910 [= Iphiclus virgatus (Kuhnt), comb. nov.]. B–B’. Holotype of M. ziczac ziczac Kuhnt, 1910. C–C’. Specimen of M. aestuans Lacordaire, 1842 from the Alvarenga collection (MNRJ, Brazil). D–D’. Lectotype of M. ambulator Lacordaire, 1842. E–E’. Lectotype of M. apicalis Lacordaire, 1842 [current combinationMycomystes apicalis (Lacordaire)]. F–F’. Lectotype of M. argus Lacordaire, 1842. Scale bars: A = see Material and methods; B–F = 1 mm.

Fig. 39. A–D, G. Primary types (dorsal, labels). A–A’. Holotype of Mycotretus partitialis Mader, 1940. B–B’. Holotype of M. prioteloides Mader, 1942. C–C’. Holotype of M. tigrinoides Mader, 1942. D–D’. Holotype of M. tigripennis Mader, 1942. E–E’. Specimen of M. maculatus (Olivier, 1791) from the BMNH collection (UK). F–F’. Topotype of M. tigrinus (Olivier, 1791) from RBINS (Belgium). G–G’. Holotype of M. alvarengai Pecci-Maddalena & Lopes-Andrade, 2018. Scale bars: A–C, E–G = 1 mm; D = 2 mm.

Fig. 21. Primary types (dorsal, labels). A–A’. Holotype of Mycotretus ternotatus Gorham, 1888. B–B’. Holotype of M. tibialis Gorham, 1888. C–C’. Lectotype of M. vittatus Gorham, 1888. D–D’. Holotype of M. anchoralis Guérin, 1956. E–E’. Holotype of M. bicinctus Guérin, 1949. F–F’. Holotype of M. monrosi Guérin, 1949. Scale bars: A–C, F = 1 mm; D–E = 0.5 mm.

Fig. 15. Gorham primary types (dorsal, labels). A–A’. Holotype of Mycotretus atricaudatus Gorham, 1888. B–B’. Lectotype of M. badius Gorham, 1888. C–C’. Lectotype of M. bipunctatus Gorham, 1888. D–D’. Holotype of M. brevis Gorham, 1888. E–E’. Lectotype of M. cercyonoides Gorham, 1888. F–F’. Lectotype of M. coccidulinus Gorham, 1888. Scale bars = 1 mm.

Fig. 40. Taschenberg specimens. B–D, F. Primary types (dorsal, labels). A–A’. Specimen of Mycotretus bicolor Taschenberg, 1870 from Alvarenga collection (MNRJ, Brazil). B–B’. Lectotype of M. coccinelloides Taschenberg, 1870 [synonym of Mycomystes apicalis (Lacordaire, 1842)]. C–C’. Lectotype of M. dimidiatus Taschenberg, 1870. D–D’. Lectotype of M. discoidalis Taschenberg, 1870 [synonym of Mycomystes melanophthalmus (Duponchel, 1825)]. E–E’. Specimen of M. dispar Taschenberg, 1870 from the Alvarenga collection (MNRJ, Brazil).F–F’. Lectotype ofM. multimaculatus Taschenberg, 1870. Scale bars: A = 0.5 mm; E = 1 mm; B–D, F = see Material and methods.

Fig. 23. Primary types (dorsal, labels). A–A’. Lectotype of Mycotretus pelliciens Kirsch, 1876. B–B’. Lectotype of M. peruvianus (Kirsch, 1876) [present combination Mycomystes peruvianus (Kirsch)]. C–C’. Lectotype of M. puncticeps Kirsch, 1865. D–D’. Lectotype of M. suturalis Kirsch, 1876. E–E’. Lectotype of M. bicoloratus Kuhnt, 1911 [= M. bicolor Kirsch, 1876 (invalid name)]. F–F’. Lectotype of M. derasofasciatus Kuhnt, 1910. Scale bars: A–E = 1 mm; F = see Material and methods.

Fig. 27. Lacordaire specimens.A–B, D–F. Primary types (dorsal, labels).A–A’. Lectotype ofMycotretus bistrigatus Lacordaire, 1842. B–B’. Lectotype of M. clitelliger Lacordaire, 1842. C–C’. Specimen of M. coccineus (Lacordaire, 1842) from the Crotch collection (UMZC, United Kingdom) (current combination Mycomystes coccineus (Lacordaire)]. D–D’. Lectotype of M. coelestinus Lacordaire, 1842. E–E’. Lectotype of M. cognatus Lacordaire, 1842. F–F’. Lectotype of M. cyanopterus Lacordaire, 1842. Scale bars = 1 mm.

Fig. 2. Alvarenga holotypes (dorsal, labels). A–A’. Mycotretus bistrioculatus Alvarenga, 1983. B–B’. M. fragosoi Alvarenga, 1983. C–C’. M. lopesi Alvarenga, 1989. D–D’. M. luizi Alvarenga, 1983. E–E’. M. octoculatus Alvarenga, 1983. F–F’. M. quadrioculatus Alvarenga, 1983. Scale bars: A–B, D–F = 0.5 mm; C = 1 mm.

Fig. 3. Primary types (dorsal, labels). A–A’. Holotype of Mycotretus tucuriensis Alvarenga, 1983. B–B’. Lectotype of M. centralis Arrow, 1909. C–C’. Lectotype of M. distinguendus Arrow, 1909. D–D’. Holotype of M. nigromanicatus Boyle, 1954. E–E’. Lectotype of M. nubifer Casey, 1916. F–F’. Lectotype of M. lesueuri (Chevrolat, 1835). Scale bars = 1 mm.

Fig. 5. Crotch primary types (dorsal, labels) cont. A–A’. Lectotype of Mycotretus chontalesi Crotch, 1873. B–B’. Lectotype of M. cinctiger Crotch, 1876. C–C’. Lectotype of M. corallipennis Crotch, 1876 [synonym of Mycomystes apicalis (Lacordaire, 1842)]. D–D’. Lectotype of M. cordiger Crotch, 1876. E–E’. Lectotype of M. cruciger Crotch, 1876. F–F’. Lectotype of M. decorus Crotch, 1876. Scale bars = 1 mm.

Fig. 16. Gorham primary types (dorsal, labels) cont. A–A’. Lectotype of Mycotretus consanguineus Gorham, 1888. B–B’. Lectotype of M. cribratus Gorham, 1888. C–C’. Holotype of M. crudus Gorham, 1888. D–D’. Lectotype of M. cruentus Gorham, 1888. E–E’. Lectotype of M. elegans Gorham, 1888. F–F’. Holotype of M. epopterus Gorham, 1888. Scale bars = 1 mm.

Fig. 12. Primary types (dorsal, labels). A–A’. Lectotype of Mycotretus xanthomelas Crotch, 1876. B–B’. Holotype of M. ornatus partitialis Deelder, 1942. C–C’. Holotype of M. fidelis Deskeskamp, 1939 [= Brachysphaenus (Barytopus) parallelus Kuhnt, 1909]. D–D’. Holotype of M. sexoculatus chaparensis Delkeskamp, 1957.E–E’. Holotype ofM. ziczac serenus Delkeskamp, 1957. F–F’. Holotype of M. zischkai Delkeskamp, 1957. Scale bars: A, C, E = 1 mm; B, D, F = see Remarks.

Fig. 38. A–B, D–F. Mader primary types (dorsal, labels). A–A’. Holotype of Mycotretus antesignatus Mader, 1942 [= Mycolybas antesignatus (Mader) comv. nov.]. B–B’. Holotype of M. cunctans Mader, 1942. C–C’. M. quadripunctatus ab. luteobifasciatus Mader, 1942 (invalid name). D–D’. Holotype of M. major Mader, 1955. E–E’. Holotype of M. opalizans Mader, 1942. F–F’. M. opalizans ab. posticenigrum Mader, 1942 (invalid name). Scale bars: A, C–F = 1 mm; B = 2 mm.

Fig. 13. A–D. Duponchel primary types (dorsal, labels). A–A’. Lectotype of Erotylus coronatus Duponchel, 1825 [currently Mycotretus coronatus (Duponchel)]. B–B’. Lectotype of E. decoratus Duponchel, 1825 [currently M. decoratus (Duponchel)]. C–C’. Lectotype of E. duodecimguttatus Duponchel, 1825 [currentlyM. duodecimguttatus (Duponchel)].D–D’. Lectotype ofE. melanophthalmus Duponchel, 1825 [currently Mycomystes melanophthalmus (Duponchel)]. E–E’. Specimen of M. minutus (Duponchel, 1825) from the DZUP collection (Brazil).F–F’. Specimen of M. nigropunctatus (Duponchel, 1825) from the Brême collection (MRSN). Scale bars: A–D = 1 mm; E = 0.5 mm; F = see Material and methods.

Fig. 6. Crotch primary types (dorsal, labels) cont. A–A’. Lectotype of Mycotretus deyrollei Crotch, 1876. B–B’. Lectotype of M. dimidiatus Crotch, 1876 [= M. seminiger Harold, 1876]. C–C’. Lectotype of M. egae Crotch, 1876. D–D’. Lectotype of M. episcaphoides Crotch, 1876. E–E’. Lectotype of M. fulviceps Crotch, 1876. F–F’. Lectotype of M. fulvilabris Crotch, 1876. Scale bars = 1 mm.

Fig. 24. Kuhnt primary types (dorsal, labels). A–A’. Lectotype of Mycotretus discipennis Kuhnt, 1910. B–B’. Lectotype of M. discipennis conductus Kuhnt, 1910. C–C’. Lectotype of M. expressus Kuhnt, 1910. D–D’. Holotype of M. fascipennis Kuhnt, 1910. E–E’. Lectotype of M. graphoderus strigipennis Kuhnt, 1910. F–F’. Lectotype of M. graphoderus thoracicus Kuhnt, 1910. Scale bars: C–D = 1 mm; A–B, E–F = see Material and methods.

Fig. 25. Kuhnt primary types (dorsal, labels) cont. A–A’. Lectotype of Mycotretus interstitialis Kuhnt, 1910. B–B’. Holotype of Brachysphaenus (Iphiclus) nigromaculatus Kuhnt, 1909 [currently M. nigromaculatus (Kuhnt)]. C–C’. Holotype of M. quadristriolatus Kuhnt, 1910. D–D’. Lectotype of M. sexlineatus Kuhnt, 1910.E–E’. Holotype ofM. stillatus Kuhnt, 1910.F–F’. Holotype ofM. thoracicus Kuhnt, 1910. Scale bars: B, E–F = 1 mm; A, C–D = see Material and methods.

Fig. 22. A–C, D–F. Primary types (dorsal, labels). A–A’. Holotype of Mycotretus trifasciatus Guérin, 1956. B–B’. Lectotype of Erotylus cinctellus Guérin-Méneville, 1841 [= M. cinctellus; synonym of Mycomystes melanophthalmus (Duponchel, 1825)]. C–C’. Lectotype of M. fallax Guérin-Méneville, 1841. D–D’. Specimen of M. sedecimguttatus (Guérin-Méneville, 1844) from the UMZC collection. E–E’. Lectotype of Erotylus (Brachymerus) sobrinus Guérin-Méneville, 1841 [currently M. sobrinus (Guérin-Meneville)]. F–F’. Lectotype of M. dichrous Kirsch, 1876. Scale bars: A = 0.5 mm; B–F = 1 mm.

Fig. 28. Lacordaire specimens cont. B, D–F. Primary types (dorsal, labels). A–A’. Specimen of Mycotretus difficilis Lacordaire, 1842 from the MZSP collection (Brazil).B–B’. Lectotype ofM. distigma Lacordaire, 1842. C–C’. Specimen of M. dorsofasciatus Lacordaire, 1842 from the Crotch collection (UMZC, UK). D–D’. Holotype of M. dorsonotatus Lacordaire, 1842. E–E’. Lectotype of M. dubius Lacordaire, 1842. F–F’. Lectotype of M. durius Lacordaire, 1842. Scale bars = 1 mm.

Fig. 33. Lacordaire specimens cont. A–B, E–F. Primary types (dorsal, labels). A–A’. Lectotype of Mycotretus melanostictus Lacordaire, 1842. B–B’. Lectotype of M. miniatus Lacordaire, 1842. C–C’. Specimen of M. misellus Lacordaire, 1842 from the Crotch collection (UMZC, UK). D–D’. Specimen of M. nigrivittis Lacordaire, 1842 from MZSP (Brazil). E–E’. Lectotype of M. nigrocinctus Lacordaire, 1842. F–F’. Lectotype of M. nigroterminatus Lacordaire, 1842 [synonym of Mycomystes apicalis (Lacordaire, 1842)]. G–G’. Specimen of M. nugator Lacordaire, 1842 from the Alvarenga collection (MNRJ, Brazil). Scale bars: A–F = 1 mm; G = 0.5 mm.

Fig. 29. Lacordaire specimens cont. A, C–F. Primary types (dorsal, labels). A–A’. Lectotype of Mycotretus dytiscoides Lacordaire, 1842. B–B’. Specimen of M. episcopalis Lacordaire, 1842 from the Alvarenga collection (MNRJ, Brazil). C–C’. Lectotype of M. fasciolatus Lacordaire, 1842. D–D’. Lectotype of M. figuratus Lacordaire, 1842. E–E’. Lectotype of M. flavomarginatus Lacordaire, 1842. F–F’. Holotype of M. floriger Lacordaire, 1842. Scale bars: A, C–F = 1 mm; B = 0.5 mm.

Fig. 17. Gorham primary types (dorsal, labels) cont. A–A’. Lectotype of Mycotretus erraticus Gorham, 1888. B–B’. Lectotype of M. geminus Gorham, 1888. C–C’. Lectotype of M. haemapterus Gorham, 1888. D–D’. Holotype of M. haematicus Gorham, 1888. E–E’. Holotype of M. hirudo Gorham, 1888. F–F’. Lectotype of M. incarnatus Gorham, 1888. Scale bars = 1 mm.

Fig. 10. Crotch primary types (dorsal, labels) cont.A–A’. Lectotype of Mycotretus quatuordecimguttatus conjunctus Crotch, 1876.B–B’. Lectotype ofM.rastratus Crotch, 1876. C–C’. Lectotype ofM. reticulatus Crotch, 1876. D–D’. Lectotype of M. sallaei Crotch, 1876. E–E’. Lectotype of M. sanguinosus Crotch, 1876 [synonym of Mycomystes coccineus (Lacordaire, 1842)]. F–F’. Lectotype of M. separandus Crotch, 1876. Scale bars = 1 mm.

Fig. 7. Crotch primary types (dorsal, labels) cont. A–A’. Lectotype of Mycotretus guatemalae Crotch, 1876. B–B’. Lectotype of M. illustris Crotch, 1876. C–C’. Lectotype of M. laeviventris Crotch, 1876. D–D’. Lectotype of M. lissomoides Crotch, 1876. E–E’. Lectotype of M. mutabilis Crotch, 1876. F–F’. Lectotype of M. mycetophagoides Crotch, 1876. Scale bars = 1 mm.

Fig. 31. Lacordaire specimens cont. A–D, F. Primary types (dorsal, labels). A–A’. Lectotype of Mycotretus hepaticusLacordaire, 1842. B–B’. Holotype of M. hilaris Lacordaire, 1842. C–C’. Lectotype of M. intermedius Lacordaire, 1842. D–D’. Lectotype of M. jocosus Lacordaire, 1842. E–E’. Specimen of M. lacertosus Lacordaire, 1842 from the Alvarenga collection (MNRJ, Brazil). F–F’. Lectotype of M. lepidus Lacordaire, 1842. Scale bars: A–D, F = 1 mm; E = 0.5 mm.

Fig. 18. Gorham primary types (dorsal, labels) cont. A–A’. Holotype of Mycotretus interstictus Gorham, 1888. B–B’. Holotype of M. laccophilinus Gorham, 1888. C–C’. Lectotype of M. luteolus Gorham, 1888. D–D’. Lectotype of M. melanotus Gorham, 1888. E–E’. Holotype of M. nigricollis Gorham, 1888. F–F’. Holotype of Mycotretus nigripes Gorham, 1888. Scale bars = 1 mm.

Fig. 30. Lacordaire primary types (dorsal, labels) cont. A–A’. Lectotype of Mycotretus fuscitarsis Lacordaire, 1842 [current combination Mycomystes fuscitarsis (Lacordaire)]. B–B’. Holotype of M. gemmula Lacordaire, 1842 [synonym of Mycomystes apicalis (Lacordaire)]. C–C’. Lectotype of M. gentilis Lacordaire, 1842 [synonym of Mycomystes apicalis (Lacordaire, 1842)]. D–D’. Lectotype of M. godarti Lacordaire, 1842. E–E’. Specimen of M. graniformis Lacordaire, 1842 from MRSN. F–F’. Lectotype of M. graphoderus Lacordaire, 1842. Scale bars = 1 mm.

Fig. 32. Lacordaire specimens cont. A, C–F. Primary types (dorsal, labels). A–A’. Lectotype of Mycotretus leprosus Lacordaire, 1842. B–B’. Specimen of M. limbatus (Lacordaire, 1842) from the Crotch collection (UMZC, UK). C–C’. Lectotype of M. luteipes Lacordaire, 1842. D–D’. Lectotype of M. magus Lacordaire, 1842. E–E’. Holotype of M. marginicollis Lacordaire, 1842. F–F’. Lectotype of M. melanopterus Lacordaire, 1842. Scale bars = 1 mm.

Fig. 36. Lacordaire specimens cont. C–F. Primary types (dorsal, labels). A–A’. Specimen of Mycotretus rufilabris (Lacordaire, 1842) from the Alvarenga collection (MNRJ, Brazil). B–B’. Specimen of M. sannio Lacordaire, 1842 from the Crotch collection (UMZC, UK). C–C’. Lectotype of M. savignyi Lacordaire, 1842. D–D’. Lectotype of M. scalaris Lacordaire, 1842. E–E’. Lectotype of M. scitulus Lacordaire, 1842. F–F’. Lectotype of M. sexoculatus sexoculatus Lacordaire, 1842. Scale bars: A = 0.5 mm; B–F = 1 mm.

Fig. 37. Lacordaire primary types (dorsal, labels) cont. A–A’. Lectotype of Mycotretus silaceus (Lacordaire, 1842).B–B’. Lectotype of M. singularis Lacordaire, 1842. C–C’. Holotype ofM. terminalis Lacordaire, 1842. D–D’. Lectotype of M. tigratus Lacordaire, 1842. E–E’. Lectotype of M. trabeatus Lacordaire, 1842. F–F’. Lectotype of M. xanthosomus Lacordaire, 1842. Scale bars = 1 mm.

Fig. 35. Lacordaire primary types (dorsal, labels) cont. A–A’. Lectotype of Mycotretus pulicarius Lacordaire, 1842 [synonym of Mycomystes apicalis (Lacordaire, 1842)]. B–B’. Lectotype of M. pusillus Lacordaire, 1842. C–C’. Lectotype of M. pygmaeus Lacordaire, 1842. D–D’. Lectotype of M. quadrinus Lacordaire, 1842. E–E’. Lectotype of M. quatuordecimguttatus Lacordaire, 1842. F–F’. Lectotype of M. rhodosomus Lacordaire, 1842. Scale bars = 1 mm.

Fig. 8. Crotch primary types (dorsal, labels) cont. A–A’. Lectotype of Mycotretus mycetophagoides erythrocerus Crotch, 1876. B–B’. Lectotype of M. mycetophiloides Crotch, 1876;. C–C’. Lectotype of M. mycetophiloides careus Crotch, 1876. D–D’. Lectotype of M. nigrotinctus Crotch, 1876. E–E’. Lectotype of M. nitescens Crotch, 1876. F–F’. Lectotype of M. opalescens Crotch, 1876. Scale bars = 1 mm.

Fig. 19. Gorham primary types (dorsal, labels) cont. A–A’. Holotype of Mycotretus normalis Gorham, 1888. B–B’. Lectotype of M. noterinus Gorham, 1888. C–C’. Holotype of M. oppositipunctum Gorham, 1888. D–D’. Lectotype of M. panamanus Gorham, 1888. E–E’. Holotype of M. pictopiceus Gorham, 1888. F–F’. Holotype of M. planus Gorham, 1888. Scale bars = 1 mm.

Fig. 9. Crotch primary types (dorsal, labels) cont. A–A’. Lectotype of Ischyrus pallidior Crotch, 1876 [currently Mycotretus pallidior (Crotch)]. B–B’. Lectotype of Mycotretus parallelus Crotch, 1876. C–C’. Lectotype of M. pebasensis Crotch, 1876. D–D’. Lectotype of M. peruae Crotch, 1876. E–E’. Lectotype of M. psylloboroides Crotch, 1876. F–F’. Lectotype of M. quadripunctatus Crotch, 1876. Scale bars = 1 mm.

Fig. 34. Lacordaire primary types (dorsal, labels) cont. A–A’. Lectotype of Mycotretus palmiphilus Lacordaire, 1842. B–B’. Lectotype of M. partitus Lacordaire, 1842. C–C’. Lectotype of M. pecari Lacordaire, 1842. D–D’. Lectotype of M. polyophthalmus Lacordaire, 1842. E–E’. Lectotype of M. psittacus Lacordaire, 1842. F–F’. Holotype of M. pulchellus Lacordaire, 1842. Scale bars: A–B, E–F = 1 mm; C–D = see Material and methods.

Fig. 20. Gorham primary types (dorsal, labels) cont. A–A’. Lectotype of Mycotretus rubidus Gorham, 1888. B–B’. Holotype of M. rufipennis Gorham, 1888. C–C’. Lectotype of M. sandicatus Gorham, 1888. D–D’. Lectotype of M. sexpunctatus Gorham, 1888. E–E’. Lectotype of M. spadiceus Gorham, 1888. F–F’. Lectotype of M. stramineus Gorham, 1888. Scale bars = 1 mm.

Fig. 11. Crotch primary types (dorsal, labels) cont. A–A’. Lectotype of Mycotretus sericeonitens Crotch, 1876. B–B’. Lectotype of M. signatellus Crotch, 1876. C–C’. Lectotype of M. signatellus imperfectus Crotch, 1876. D–D’. Lectotype of M. succinctus Crotch, 1876. E–E’. Lectotype of M. tricolor Crotch, 1876. Scale bars = 1 mm.