Glyphiulus subcostulifer, Golovatch & Geoffroy & Mauriès & Spiegel, 2007

Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & Spiegel, Didier Van Den, 2007, Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group, Zoosystema 29 (3), pp. 417-456 : 439-441

publication ID	https://doi.org/ 10.5281/zenodo.5393630
persistent identifier	https://treatment.plazi.org/id/B36487D8-FFD7-D520-D2D4-FB1DFDEEFE52
treatment provided by	Marcus
scientific name	Glyphiulus subcostulifer
status	sp. nov.

Treatment

Glyphiulus subcostulifer View in CoL n. sp.

( Figs 17-19 View FIG View FIG View FIG )

TYPE MATERIAL. — Laos. Vang Vieng, Tham None, 102.4338°E, 18.9513°N, cave, 1.I.2000, leg. A. Bedos & L. Deharveng (LAO-111), holotype ♂ ( MNHN GA 050), paratype ♂ ( SEM).

ETYMOLOGY. — To emphasize the very close resemblance to G. costulifer n. sp.

DIAGNOSIS. — Differs from congeners by the unique carinotaxy formulae, coupled with certain details of leg and gonopod structure (see also key below).

DESCRIPTION

Holotype with 50p+2a+ T, about 29 mm long and

1.7 mm wide; paratype 1.3 mm wide.

Coloration uniformly light brownish. Only three or four slightly infuscate and convex ocelli in each eye patch ( Fig. 17A View FIG ).

Antennae ( Fig. 19A View FIG ), postcollar constriction, dorsal crests (including ozoporiferous ones) and carinotaxy pattern on postcollar terga ( Fig. 17 View FIG C- E), segment’s texture and cross-section ( Figs 17C, D View FIG ; 18A, B View FIG ), paraprocts and hypoproct ( Fig. 17F View FIG ), ventral flaps of male segment 7 ( Fig. 18B View FIG ), male legs 2 and 3 ( Fig. 19D, E View FIG ) generally as in G. costulifer n. sp., but gnathochilarium (mentum divided, n = 3) ( Fig. 19B View FIG ) slightly more polytrichous; collum ( Fig. 17B View FIG ) with five, not three, anteriorly and six, not two, posteriorly abbreviated crests, median crest (ma) only scarcely shorter than both flanking ones (I-II+3c+4a+5c+VI+7c+8a+pc+ma +pc+8a+7c+VI+5c+4a+3c+II-I); all tergal crests ( Figs 17 View FIG A-E; 18A, B) on subsequent body segments subdivided, but anterior tooth on dorsal crests more rounded, and both lateral crests below ozoporiferous cone normal (2/2+I/i+3/3+I/i+2/2); epiproct ( Fig. 17E, F View FIG ) upturned and very broadly rounded in caudal part; legs slightly shorter than body diameter ( Fig. 18A View FIG ), always with an evident but short accessory spine at base of claw ( Fig. 19 View FIG D- F); male leg 1 with 4-segmented, considerably shortened telopodites and large paramedian coxal processes ( Fig. 19C View FIG ); each anterior gonopod coxosternal plate subsecuriform ( Fig. 19G View FIG ); posterior gonopods with distally slightly plumose flagella ( Fig. 19H View FIG ).