Trichocoelina vitticollis ( Holmgren, 1883 ) Vilkamaa & Menzel, 2019

Vilkamaa, Pekka & Menzel, Frank, 2019, Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species, Zootaxa 4665 (1), pp. 1-67 : 47-50

Trichocoelina vitticollis ( Holmgren, 1883) comb. n.

Synonyms: = glacialis ( Lundbeck, 1898) [as Sciara ; preocc., not Sciara glacialis Rübsaamen, 1898 ]; = permutata ( Lundbeck, 1900) [as Sciara ; new name for Sciara glacialis Lundbeck, 1898 ].

Literature. Sciara glacialis Lundbeck—Lundbeck (1898) : 254, pl. 6, fig. 13. Sciara humicola Lundbeck—Lund-beck (1898) : 252, pl. 6, fig. 11 [misidentification]. Sciara permutata Lundbeck—Lundbeck (1900) : 313. Bradysia permutata (Lundbeck) — McAlpine (1964): 128. Bradysia (Hemineurina) permutata (Lundbeck) — Frey (1948): 66, 84; pl. 18, fig. 106. Lycoriella (Hemineurina) permutata (Lundbeck) — Tuomikoski (1959a): 36; Tuomikoski, (1960): 75, 76; Stone & Laffoon (1965): 232; Tuomikoski (1967): 48; Gerbachevskaja-Pavluchenko (1986): 31 [not sensu Mohrig et al. (1983a) 15; misidentification, = T. subpermutata ( Mohrig & Mamaev, 1990) ]. Sciara vitticollis Holmgren—Holmgren (1883) : 182; Jacobson (1898): 190; Gerbachevskaja-Pavluchenko (1986): 71. Lycoria (Neosciara) vitticollis (Holmgren) — Lengersdorf 1928 –30: 59. Lycoriella (Hemineurina) vitticollis (Holmgren) — Menzel & Mohrig (2000): 411, figs 380, 381; Coulson & Refseth (2004): 103; Coulson (2008): 162; Coulson (2013): 154; Mohrig et al. (2013): 271; Vilkamaa (2015): 551; Wirta et al. (2016): appendix, unpaginated p. 21 (table S1) and unpaginated p. 39 (cladogram).

New records. CANADA, Northwest Territories, Mackenzie Delta, Reindeer Depot , 29.VI.1948, J.R. Vockeroth, 1 male (in MZH) ; NE GREENLAND, Zackenberg , 74°28’N, 20°34’W, 1.VIII.1991, J. Böcher, 1 male (in ZMUC) ; NW GREENLAND, Thule , 8.VII.1940, C. Wibe, 1 male (in ZMUC) ; SE GREENLAND, Skoldängen , 14–27.VII.1992, 2 males (1 in MZH, 1 in ZMUC) ; SW GREENLAND, Nugssuak , 18.VII.1949, C. Wibe, 1 male (in ZMUC) ; Saputit , 6.VII.1949, C. Wibe, 1 male (in MZH) ; W GREENLAND, Arnangarnup kua, 11.VII.1984, J. Böcher, 1 male (im ZMUC) ; Skjoklungen, ‘ Bygder’ , 19–27.VII.1992, S. Andersen, 1 male (in ZMUC) ; Söndre Stormfjord , 21.VI.1952, C. Wibe, 1 male (in MZH) ; same locality but 67°02’N, 50°40’W, 2.VIII.1992, J. Böcher, 1 male (in MZH) ; NORWAY, ‘ Svalbard, Bjorndalen’ [= Svalbard, Spitsbergen, Nordenskiöld Land, valley Bjørndalen W of Adventfjorden], 78.2320°N, 15.3270°E, 13.VII.2012, G.E.E. Søli, 1 male (NHMO 261403 and BOLD Sample ID SV985, in NHMO) ; ‘ Svalbard, Bolterdalen’ [= Svalbard, Spitsbergen, Nordenskiöld Land, valley Bolterdalen on the southern side of Adventdalen] , 78.1640°N, 15.9900°E, 100 m, 10.VII.2012, G.E.E. Søli, 3 males (NHMO 260984 and BOLD Sample ID SV1061, NHMO 260985 and BOLD Sample ID SV1062, 2 in NHMO; BOLD Sample ID SV1064, in SDEI); ‘Svalbard, Colesbukta’ [= Svalbard, Spitsbergen, Nordenskiöld Land, bay Colesbukta on the southern side of Isfjorden] 78.1120°N, 15.0290°E, 11.VII.2012, T. Ekrem, E. Stur & G.E.E. Søli, 1 male (NHMO 261328 and BOLD Sample ID SV910, in NHMO) ; ‘ Svalbard, Hanaskogdalen’ [= Svalbard, Spitsbergen, Nordenskiöld Land, valley Hanaskogdalen on the eastern side of Adventfjorden] , 78.2830°N, 15.6050°E, 25 m, 12.VII.2012, T. Ekrem, E. Stur & G.E.E. Søli, 4 males (NHMO 261385 and BOLD Sample ID SV967, NHMO 261390 and BOLD Sample ID SV972, 2 in NHMO; BOLD Sample ID SV945, 1 in NTNU-VM; BOLD Sample ID SV974, 1 in SDEI); ‘Svalbard, Krossfjorden, 14. juli bukta’ [= Svalbard, Spitsbergen, bay Fjortende Julibukta (northern side) on the eastern side of Krossfjorden], 79.1284°N, 11.8582°E, 3 m, 18.VII.2013, T. Ekrem, K. Harsaker & G.E.E. Søli, 1 male (NHMO 261046 and BOLD Sample ID SV1137, in NHMO); ‘ Svalbard, Lakselva’ [= Svalbard, Bjørnøya, river Lakselva], 74.4994°N, 18.9776°E, 15 m, 28.VII.2009, T. Ekrem, 1 male (NTNU-VM 50612 and BOLD Sample ID BJ262, in NTNU-VM) ; same locality but 3.VIII.2009, T. Ekrem, 2 males (NTNU-VM 50606 and BOLD Sample ID BJ256, NTNU-VM 50608 and BOLD Sample ID BJ258, in NTNU-VM) ; ‘ Svalbard, Lillehoeoekfjorden , Nilspynten’ [Sval- bard, Spitsbergen, Albert I Land, Lillehøkfjorden, E part of Mitrahalvøya, Nilspynten], 79.2660°N, 11.5690°E, 10 m, 18.VII.2013, G.E.E. Søli, 6 males (NHMO 261244 and BOLD Sample ID SV825, NHMO 261245 and BOLD Sample ID SV826, NHMO 261253 and BOLD Sample ID SV834, NHMO 261254 and BOLD Sample ID SV835, 4 in NHMO; BOLD Sample ID SV828, in SDEI; BOLD Sample ID SV 831 in MZH); ‘ Svalbard, Longyearbyen’ [= Svalbard, Spitsbergen, Nordenskiöld Land, Longyearbyen in the Longyeardalen S of Adventfjorden], 78.2170°N, 15.6180°E, 16.VII.2012, G.E.E. Søli, T. Ekrem & E. Stur, 2 males (NHMO 261508 and BOLD Sample ID SV1263, NHMO 261175 and BOLD Sample ID SV1266, in NHMO); ‘Svalbard, Longyearbyen’ [= Svalbard, Spitsbergen, Nordenskiöld Land, Longyearbyen in the Longyeardalen S of Adventfjorden], 78.2088°N, 15.5889°E, 14.VII.2012, G.E.E. Søli, 2 males (NHMO 261508 and BOLD Sample ID SV-NHMO91, in NHMO; BOLD Sample ID SV-NHMO94, in SDEI); ‘ Svalbard, Longyearbyen’ [= Svalbard, Spitsbergen, Nordenskiöld Land, Longyearbyen in the Longyeardalen S of Adventfjorden], 78.2130°N, 15.6040°E, 14.VII.2012, E. Stur, 2 males (NHMO 261191 and BOLD Sample ID SV1282, in NHMO; BOLD Sample ID SV1283, in MZH); Svalbard, NW part of Spitsbergen , southern coast of Kongsfjord , W of Ny Ålesund, yellow pan trap, 2–15.VII.1974, Stephan , 2 males (in NHMO, SDEI); SWEDEN, Lapland , Abisko, Naturvetenskapliga Station, meadow at the station, sweep-net, 28.VI.1988, M. von Tschirnhaus, 4 males (2 in NHMO, 2 in SDEI) .

BIN. BOLD:ABA5288.

Discussion. The species was described from one female from Novaya Zemlya by Holmgren (1883) and later described again based on some males and females from Greenland as Sciara glacialis ( Lundbeck 1898) and from Spitsbergen as Sciara permutata ( Lundbeck 1900) . The species was redescribed with a list of literature, synonyms and combinations by Menzel & Mohrig (2000). Trichocoelina vitticollis (Holmgren) can be distinguished from all other species of the genus by its very large, setose intergonocoxal lobe of hypopygium and in having the gonostylus with a group of slender, oblique medial megasetae. See also under Trichocoelina tecta sp. n.