Albunea occultus, BOYKO, 2002

BOYKO, CHRISTOPHER B., 2002, A Worldwide Revision Of The Recent And Fossil Sand Crabs Of The Albuneidae Stimpson And Blepharipodidae, New Family (Crustacea: Decapoda: Anomura: Hippoidea), Bulletin of the American Museum of Natural History 2002 (272), pp. 1-396 : 313-327

Albunea occultus , new species Figures 100 , 101

Albunea symnista [sic]: Gordon, 1938: 187 (part)*. – Serène and Umali, 1965: 90–95 (part)*. – Miyake, 1991: 157, pl. 53, fig. 2.— Asakura, 1995: 376, fig. 21–285 (not Albunea symmysta ( Linnaeus, 1758)) .

Albunea symmista [sic]: Miyake, 1965: 651 * (not Albunea symmysta ( Linnaeus, 1758)) .

MATERIAL EXAMINED: Japan: Tosa­Shimizu , Tosa Bay, Shikoku Island, 4–7 m, May 1959, coll. K. Kurohara: 2 ♀, 8.5–15.3 mm cl, paratypes ( ZLKU 7040–7041 ) ; Tosa Bay , Shikoku Island, July 6, 1959, coll. S. Nakayama: 1 Ƌ, 16.7 mm cl, 1 ♀, 18.1 mm cl, paratypes ( ZLKU 7062–7063 ) .

Taiwan: Taiwan Strait, south of Formosa Banks to the Penghu (Pescadores) Islands , 30–50 m, May 5, 1972, coll. commercial fishermen: 1 ♀, 23.0 m cl, paratype ( CASIZ 109246 ) ; Kaohsiung, 15 m, May 10, 1991, coll. unknown: 1 ♀, 24.0 mm cl, paratype ( NTOU) ; Kaohsiung, 20 m, April 3, 1988, coll. unknown: 1 ♀, 25.3 mm cl, paratype ( NTOU) ; ‘‘ Taiwan ,’’ May 17, 1992, coll. unknown: 1 ♀, 22.3 mm cl, paratype ( NTOU) ; ‘‘ southern Taiwan ,’’ 5 m, Jan. 24, 1992, coll. unknown: 1 Ƌ, 19.7 mm cl, paratype ( NTOU) .

Philippines: Mindoro , coll. P. de Mesa: 1 Ƌ, 10.1 mm cl, paratype ( MCZ 10261) ; Morong , Bataan, Feb. 28, 1999, coll. J. D. Williams: 1 juvenile, 4.0 mm cl, paratype ( AMNH 18094 ) ; Nasasa Bay, Zambales Province, 5–18 fms (= 9.1–32.9 m), Jan. 30&ndash ♀ b. 8, 1960, coll. F. G. Dayrit and J. E. Norton: 1 Ƌ, 6.7 mm cl, 1 ♀, 7.3 mm cl, paratypes ( USNM 267779 ) ; south lagoon, Sibutu, Sulu Archipelago , 04°31̍N, 119°22̍E, 13 fms (= 23.8 m), Feb. 25–26, 1964, coll. B. R. Wilson on ‘‘ Pele’ ’: 1 Ƌ, 8.6 mm cl, paratype ( WAM 10411) .

Australia: Western Australia: Coral Bay , 1989, coll. C. Simpson: 1 ♀, 20.2 mm cl, paratype ( WAM 20261) ; Sta. 3, northeast corner of North Pasco Island, 20°54̍S, 115°19̍E, Sept. 17, 1966, coll. WAM­USNM Barrow Island Expedition 1966: 1 Ƌ, 16.4 mm cl, paratype ( WAM 23392) ; Sta. 05D06BT, northwest shelf, 19°29.7̍S, 118°52.1̍E, 38–39 m, Oct. 25, 1983, coll. CSIRO: 1 Ƌ, 12.9 mm cl, 3 ♀, 6.2–12.4 mm cl, paratypes ( QM W22322) ; Sta. 05B07BT, northwest shelf, 19°30.9̍S, 118°49.2̍E, 38– 39 m, Oct. 25, 1983, coll. CSIRO: 1 ♀, 17.7 mm cl, paratype ( QM W22323) ; Sta. 05D02BT, northwest shelf, 19°29.4̍S, 118°52.4̍E, 37–38 m, Oct. 24, 1983, coll. CSIRO: 3 Ƌ, 6.1–14.0 mm cl, 3 ♀, 7.5–9.2 mm cl, paratypes ( QM W22326) ; Sta. 04B09BT, northwest shelf, 19°28.4̍S, 118°55.2̍E, 39 m, Aug. 31, 1983, coll. CSI­ RO: 1 ♀, 7.7 mm cl, paratype ( QM W22329) ; Sta. 05B12BT, northwest shelf, 19°03.6̍S, 119°03.4̍E, 82 m, Oct. 23, 1983, coll. CSIRO: 2 Ƌ, 6.3–10.3 mm cl, paratypes ( QM W22338) ; Sta. 04B14S, northwest shelf, 19°43.9̍S, 117°54.5̍E, 52–53 m, Sept. 2, 1983, coll. CSIRO: 2 ♀, 5.6–6.2 mm cl, paratypes ( QM W22340) ; ‘‘northwest Australia ,’’ coll. unknown: 1 ♀, 10.4 mm cl, paratype ( BMNH 1932.11.30.64) ; Northern Territory: Western shore, Darwin Harbour , Nov. 3, 1963, coll. unknown: 1 Ƌ, 18.6 mm cl, paratype ( WAM 23393) ; Waigait, Darwin , Nov. 11, 1969, coll. O. J. Cameron: 2 Ƌ, 15.1–17.5 mm cl, 1 ♀, 18.1 mm cl, paratype (AM P19435) ; Mandora, Darwin , Nov. 22, 1972, coll. N. Coleman: 1 oviger, 23.7 mm cl, paratype (AM P19436) ; Mandora, Darwin , Nov. 21, 1972, coll. N. Coleman: 2 Ƌ, 14.0– 17.5 mm cl, paratypes (AM P23809) ; Dudley Point, Darwin , Sept. 17, 1970, coll. O. J. and J. Cameron: 1 ♀, 21.0 mm cl, paratype (AM P19993) ; sand bar no. 1, Darwin , Nov. 14, 1970, coll. O. J. and J. Cameron: 1 Ƌ, 19.7 mm cl, holotype (AM P20486) ; Queensland: Michaelmas Cay, Great Barrier Reef, north of Cairns , Sept. 14, 1963, coll. W. Goode: 1 ♀, 11.1 mm cl, paratype ( WAM 24510) ; Bowen , Feb. 27, 1934, coll. unknown: 1 Ƌ, 16.5 mm cl, paratype ( QM W25188) ; Mornington Island, Gulf of Carpentaria , 16°29̍S, 139°34̍E, coll. unknown: 1 Ƌ, 9.1 mm cl, paratype ( MOV J44733 ) ; off Cape Moreton , April 1964, coll. B. Harris: 1 ♀, 26.7 mm cl, paratype ( QM W2380 ) ; New South Wales: Off Richmond River mouth, off Ballina , 28°52̍S, 153°34̍E, 16 m, Oct. 6, 1962, coll. unknown: 1 Ƌ, 17.3 mm cl, paratype (AM P15354) ; 3 mi off mouth of Clarence River , 15 fms (= 27.4 m), late 1968, coll. J. J. Toyer: 1 ♀, 22.2 mm cl, allotype (AM P16778) ; within 2 km north of Clarence River bar, 8–9 fms (= 14.6–16.5 m), June 10, 1976, coll. A. E. Ford: 1 ♀, 25.0 mm cl, paratype (AM P24477) .

Limited Data: ‘‘Océan Indien:’’ 1 ♀, 20.6 mm cl (MNHN­Hi 89).

DIAGNOSIS: Carapace wider than long, covered with lightly setose grooves. Anterior margin with 8–11 spines on either side of ocular sinus. Setal field with narrow lateral elements and concave anterior margin. CG1 with separate posterior lateral elements; CG4 with one or two short, concave, anteriorly displaced medial elements between longer supralateral elements of CG4; CG5 of two triangular elements; CG6 and CG7 separate; CG8 broken; CG11 present; posterior submarginal groove entire. Rostrum present, not reaching posterior margin of ocular plate. Ocular plate triangular. Distal peduncular segments dorsoventrally flattened and triangular, tapering at tip, approximated along mesial margins, lateral margins convex, mesial margins straight. Cornea at tip. Dactylus of pereopod II with heel produced, tapered, and subacute. Dactylus of pereopod III with heel tapering, projecting, acute. Dactylus of pereopod IV sinuous from base to tip, with low rounded heel and shallow indent. Telson of male broadly triangular, laterally produced, length subequal to width, with broadly rounded produced tip; thickly calcified medially, inflated dorsally; distal two­thirds with lateral decalcified region, median row of thin setae. Telson of female flattened, ovate, and evenly calcified with slightly produced tip.

DESCRIPTION: Carapace (fig. 100A) slightly wider than long. Anterior margin slightly concave on either side of ocular sinus, becoming convex laterally, with 8–11 large spines (n = 6) along length. Rostrum as small acute tooth, not reaching proximal margin of ocular plate. Ocular sinus smoothly concave, with one or two large spines and two or three small spinules. Frontal region smooth; setal field narrow anteriorly and posteriorly; posterior lateral elements reduced to narrow bands of setae. CG1 parallel to anterior margin of carapace, sinuous, strongly crenulate, divided into medial fragment and curved, posteriorly displaced lateral ele­ ments. Mesogastric region smooth; CG2 present as one to three small medial elements; CG3 broken into two short medial and two to four longer lateral elements between posterior lateral elements of CG1, medial elements anteriorly displaced; CG4 with one or two short, concave, anteriorly displaced medial elements between longer supralateral elements of CG4. Hepatic region smooth, with oblique setose groove at median of lateral margin. Epibranchial region generally triangular, smooth; posterolateral margin without rows of setae. Metagastric region smooth; CG5 present as two triangular elements. CG6 strongly crenulate, strongly anteriorly concave medially and sloping out to anteriorly convex lateral thirds. CG7 oblique, not reaching lateral margins of median segment of CG6. Cardiac region smooth; CG8 present as one long or two short lateral elements. CG9 present as two short lateral grooves with gap at midline. CG10 present as two short lateral elements. CG11 present as one long or two short medial elements. Post­ CG11 element absent. Branchial region with numerous short, transverse rows of setae laterally. Posterior margin deeply and evenly convex, with submarginal groove reaching three­fourths up margin of posterior concavity to entire. Branchiostegite with short anterior submarginal spine; anterior region with scattered, short, transverse lines ventral to linea anomurica; with many short rows of setae and sparsely covered with long plumose setae ventrally; posterior region membranous, with numerous irregular fragments and sparsely covered with long plumose setae.

Ocular plate (fig. 100B) triangular with deep median indentation; median peduncular segments present as small ovate calcified areas lateral to ocular plate. Distal peduncular segments elongate, subtriangular, with slightly convex lateral and straight mesial margins, cornea covering lateral portion of distal tip; lateral margins with notch one­fourth distal from base; mesial margins approximated at base, with long plumose setae; tuft of plumose setae at proximolateral ventral angles and ventromedial oblique row of plumose setae extending from tuft to three­fourths length of segment.

Antennule (fig. 100C) with segment III narrow proximally, expanding distally to 1.5 times proximal width; with plumose setae on dorsal and ventral margins; dorsal exopodal flagellum with 69–88 articles (n = 6), long plumose setae on dorsal and ventral margins; ventral endopodal flagellum with two or three articles (n = 6), plumose setae on dorsal and ventral margins. Segment II medially inflated from dorsal view, with plumose setae on dorsal and ventral margins and scattered on ventrolateral quarter of surface. Segment I longer than wide, unarmed; dorsal third of lateral surface rugose, with long plumose setae; long plumose setae on dorsal and ventral margins.

Antenna (fig. 100D) with segment V approximately two times longer than wide, with long plumose setae on dorsal and ventral margins and scattered on lateral surface; flagellum with seven or eight articles (n = 6; but QM 2380 with 10 articles on one side and eight on other), long plumose setae on dorsal, ventral, and distal margins. Segment IV expanded distally, with long plumose setae on dorsal and ventral margins, and row of short setae on dorsolateral surface. Segment III with short plumose setae on dorsal margin, long plumose setae on ventral margin. Segment II short, widening distally, rugose, with plumose setae on margins and scattered on lateral surface; antennal acicle long, thin, and exceeding distal margin of segment IV by one­seventh length of segment IV, with long plumose setae on dorsal margin. Segment I rounded proximally, flattened ventrolaterally, with long plumose setae on margins, lobe, and in short row on surface rugae behind spine; lateral surface with acute spine dorsodistally, with low semicircular dorsolateral lobe ventrodistal to spine; segment with ventromesial antennal gland pore.

Mandible (fig. 100E) incisor process with three teeth; cutting edge smooth. Palp threesegmented, with plumose setae on margins and long, thick, simple setae arising from bend in second segment and on distal margin of terminal segment.

Maxillule (fig. 100F) distal endite proximally narrow, widening to inflated distal end, with thick simple setae on distal margin and thin simple setae on dorsal margin. Proximal endite with thick simple setae on distal mar­ gin. Endopodal external lobe truncate distally and curled under; internal lobe reduced with six thick setae at distolateral margin.

Maxilla (fig. 100G) exopod evenly round­ ed, with plumose setae along distal margin. Scaphognathite bluntly angled on posterior lobe, with plumose setae.

Maxilliped I (fig. 100H) epipod with plumose setae on margins and mesial surface. Endite tapered distally and subequal to first segment of exopod. Exopod with two segments; proximal segment narrow, margins parallel, with plumose setae; distal segment spatulate, longer than wide, broadest medially, margins, proximolateral and mesiomedial surface with long plumose setae. Endopod flattened and elongate, reaching twothirds to distal end of proximal exopodal segment; plumose setae on margins and median of lateral surface.

Maxilliped II (fig. 100I) dactylus evenly rounded, length slightly greater than width, with thick simple setae distally and on distolateral surface. Propodus two times wider than long, slightly produced at dorsodistal angle, with plumose setae on dorsal margin and patch of long simple setae on dorsodistal and ventrodistal angles. Carpus not produced dorsodistally, approximately two times longer than wide; long simple setae on dorsal and ventrodistal margins. Merus approximately three times longer than wide, margins parallel; with simple and plumose setae on margins and scattered on surface. Basis­ischium incompletely fused, with plumose setae on margins. Exopod one­third longer than merus, flagellum with one elongate article, approximately as long as carpus.

Maxilliped III (fig. 100J) dactylus oblong with rounded tip; long plumose setae on margins and lateral surface. Propodus dorsodistally inflated, with longitudinal median row of plumose setae on lateral surface; margins with plumose setae. Carpus produced onto propodus almost one­third length of propodus; lateral surface with two rows of plumose setae; long plumose setae on margins. Merus inflated, unarmed, with plumose setae on margins and scattered in short patches on lateral surface. Basis­ischium incompletely fused, with weak crista dentata of three or four teeth. Exopod two­segmented: proximal segment small; distal segment styliform, ta­ pering, approximately one­third length of merus; with plumose setae on margins; without flagellum.

Pereopod I (fig. 101A) dactylus curved and tapering; lateral and mesial surfaces smooth; dorsal margin with long plumose setae; ventral margin with short simple setae. Propodal lateral surface with numerous short, transverse rows of setose rugae; dorsal margin unarmed; ventral margin distally produced into acute spine; cutting edge lacking teeth, lined with long plumose setae; dorsal margin with long plumose setae, ventral margin with short simple setae. Carpus with dorsodistal angle produced into strong corneoustipped spine; dorsal margin with short trans­ verse grooves behind spine; dorsal and distal margins with long plumose setae; lateral surface with small distal rugose area, few transverse setose ridges on distal half of surface; mesial surface smooth, with subdorsal and medial transverse rows of long plumose setae, margins with long plumose setae. Merus unarmed; lateral surface with scattered transverse rows of long plumose setae, proximodorsal margin with long plumose setae; mesial surface with few scattered setae; fully calcified. Basis­ischium incompletely fused, unarmed. Coxa unarmed.

Pereopod II (fig. 101B) dactylus smooth; base to heel slightly concave, heel produced, broad and rounded at tip, heel to tip with narrow, acute indent, tip acute, tip to base broadly convex; lateral surface smooth, with several small tufts of short setae in generally straight line across medioproximal surface, several widely spaced submarginal tufts of short setae dorsodistally; mesial surface smooth, ventral margin with long plumose setae, dorsal margin with short simple setae, with patch of long plumose setae at base. Propodal dorsal surface smooth, with ventral margin inflated and rounded; oblique row of long plumose setae on distal margin of lateral surface; distal and ventral margins with long plumose setae; dorsolateral surface as narrow, oblique, flattened shelf, with short setae on dorsal margin and long plumose setae on ventral margin; mesial surface with elevated, curved, setose ridge from ventral junction with dactylus almost to ventral proximal junction with carpus. Carpus strongly produced and rounded dorsodistally, dorsal margin smooth; lateral surface smooth, produced area smooth, irregular, interrupted row of rugae and submarginal elevated ridge ventrally, rugae and ridge with long plumose setae; dorsal margin with short plumose setae, ventral margin with long plumose setae; mesial surface dorsal margin with numerous low, rounded spines, surface smooth, with row of long plumose setae distally and in oblique interrupted row. Merus with large median decalcified window covering nearly all of lateral surface, with few scattered tufts of long plumose setae on surface and on distodorsal margin; mesial surface nearly smooth, with two long rows of setae. Basis­ischium in­ completely fused and unarmed. Coxa unarmed.

Pereopod III (fig. 101C) dactylus with base to heel concave, heel narrow and acutely produced, heel to tip with concave area at base of heel and broadly concave indent, tip acute, tip to base smoothly convex; lateral surface smooth, with several small tufts of short setae in generally straight line across medioproximal surface, dorsodistal margin with tufts of short setae; ventral margin with long plumose setae, dorsal margin with short simple and plumose setae; mesial surface smooth, with plumose setae proximally at junction with propodus. Propodus not inflat­ ed dorsoventrally; lateral surface smooth, with long plumose setae in oblique row, simple setae on dorsal margin, plumose setae on ventral margin; dorsolateral surface narrow, oblique, flattened, with setose mat; mesial surface smooth, with few scattered setae. Carpus produced dorsodistally, exceeding proximal margin of propodus by one­third length of propodus; tip rounded, dorsolateral margin unarmed; lateral surface slightly rugose dorsodistally, with mat of short setae and two interrupted rows of setae ventrally; mesial surface smooth, with long plumose setae on distal margin and in transverse row on surface. Merus smooth, with large decalcified window covering nearly half of lateral surface medially; dorsal and ventral margins unarmed, with long plumose setae on distodorsal and mediolateral margins; mesial surface smooth. Basis­ischium incompletely fused and unarmed. Coxa unarmed. Female with large gonopore on anterior mesial margin of coxa, surrounded with short plumose setae; male with tiny pore located more mesially.

Pereopod IV (fig. 101D) dactylus with base to tip convex proximally, concave distally, tip acute, tip to base concave distally to convex proximally; lateral surface smooth, ventral margin with long plumose setae, dorsal margin with short simple setae; mesial surface with dorsal decalcified region, demarcated ventrally by longitudinal elevated ridge with row of short setae; with setose punctations ventral to decalcified window. Propodus expanded dorsally and ventrally; ventral expansion reaching ventral margin of dactylus, ventral margin with long plumose setae; dorsal expansion with row of long plumose setae dorsally, oblique area with mat of short simple setae; lateral surface smooth, mesial surface smooth, with distoventral area of few patches of long plumose setae. Carpus slightly produced and subacute dorsodistally; ventral four­fifths of lateral surface and mesial surface smooth, dorsodistal one­fifth of lateral surface with mat of short setae; dorsal margin with short simple and long plumose setae; ventral margin with short simple setae; mesial surface decalcified medially, distal margin with long plumose setae. Merus with scattered, short, transverse rows of short simple setae on lateral surface, dorsal and medioventral margins with long plumose setae; proximoventral half of mesial surface with large decalcified window. Basis­ischium incompletely fused and unarmed. Coxa unarmed.

Abdomen (fig. 101E) somite I wider than long, widest posteriorly; dorsal surface with anterior margin straight; posterior margin curved with elevated submarginal row of short setae; small transverse decalcified windows laterad of segment median. Somite II dorsal surface with irregular submarginal transverse ridge anteriorly; with small transverse decalcified windows laterad of segment median just anterior to submarginal ridge; pleura expanded and directed laterally; anterolateral margins angled, anterior and lateral margins with long plumose setae, posterolateral angle rounded, posterior margin with short setae; posteromesial angle with mat of short simple setae. Somite III similar to somite II, but narrower, shorter, anterior submarginal windows present; pleura thinner and shorter than on somite II, directed posterolaterally, with setae as in somite II; anterolateral angle subacute; dorsal surface obliquely flattened anterolaterally, with submarginal row of short setae. Somite IV similar to somite III, but thinner and shorter, anterior submarginal windows present; dorsal surface with few short setae anterolaterally; pleura thinner and shorter than on somite III, directed posterolaterally; dorsal surface obliquely flattened anterolaterally; margins with long plumose setae. Somite V wider than somite IV, anterior submarginal windows present; lateral margins with plumose setae; pleura absent. Somite VI slightly broader than somite V, anterior submarginal windows present; dorsal surface with short transverse rows of setae laterad of midline; pleura absent.

Females with uniramous, paired pleopods on somites II–V; males without pleopods.

Telson of male (fig. 101F) broadly triangular, laterally produced, length subequal to width, with broadly rounded produced tip; thickly calcified medially, inflated dorsally; distal two­thirds with lateral decalcified region; median longitudinal groove extending one­half of length, row of long simple setae of either side of median groove beginning at median and continuing almost to distal margin of calcified area; proximolateral angles with few long simple setae; margins with long simple setae. Telson of female (fig. 101G) flattened, ovate, and evenly calcified with slightly produced rounded tip; median groove similar to male, setal row from midpoint of median groove to near distal margin of telson with simple setae approximately one­half size of those of male; proximolateral angle with few short setae, margins with long simple setae.

DISTRIBUTION: From southern Japan southward to the eastern and western coasts of Australia, in up to 82 m depth.

MAXIMUM SIZE: Males: 19.7 mm cl; females: 26.7 mm cl.

TYPE SPECIMENS: AM P20486 (holotype), AM P16778 (allotype), AM P15354 (paratype), AM P19435 (3 paratypes), AM P19436 (paratype), AM P19993 (paratype), AM P23809 (2 paratypes), AM P24477 (paratype), AMNH 18094 (paratype), BMNH 1932.11.30.64 (paratype), CASIZ 109246 (paratype), MCZ 10261 (paratype), MOV J44733 (paratype), NTOU (paratype), NTOU (paratype), NTOU (paratype), NTOU (paratype), QM W2380 (paratype), QM W22322 (4 paratypes), QM W22323 (paratype), QM W22326 (6 paratypes), QM W22338 (2 paratypes), QM W22329 (paratype), QM W22340 (2 paratypes), QM W25188 (paratype), USNM 267779 (2 paratypes), WAM 10411 (paratype), WAM 20261 (paratype), WAM 23392 (paratype), WAM 23393 (paratype), WAM 24510 (paratype), ZLKU 7040– 7041 (2 paratypes), ZLKU 7062–7063 (2 paratypes).

TYPE LOCALITY: Sand bar no. 1, Darwin, Northern Territory, Australia.

ETYMOLOGY: The specific name of this taxon is derived from the Latin word meaning ‘‘hidden,’’ and refers to the confusion of this species with A. symmysta both in the literature and in museum collections.

REMARKS: The specimen illustrated by Asakura (1995) offers a rare example of a record that can be confirmed based on a photograph alone. Asakura’s (1995) specimen clearly has an entire posterior submarginal groove on the carapace. Of the three species of Albunea known from Japan ( A. symmysta , A. groeningi , A. occultus ), only A. occultus possesses this diagnostic character.

It is difficult to know what species Serène and Umali (1965) were dealing with from the Philippines, but at least some of their material was probably A. occultus , based on their comments regarding the shape of the dactyli of pereopod IV. Their material also likely contained A. symmysta , but all of it needs to be examined for confirmation.

This species is most closely related to A. symmysta .

Albunea elegans

A. Milne Edwards and Bouvier, 1898 Figures 102 , 103

Albunea symnista [sic]: Brulle´, 1836–1844: 17. – Ozorio, 1888: 186 (not Albunea symmysta ( Linnaeus, 1758)) .

Albunea elegans A. Milne Edwards and Bouvier, 1898: 236–238 .—A. Milne Edwards and Bouvier, 1900: 273–275, pl. 28, figs. 22–25. – Gordon, 1938: 187 (list). – Bouvier, 1940: 181. – Balss, 1957: 1598.

Albunea carabus: Balss, 1916a: 37 *. – Monod, 1933: 473 (part) (not Albunea carabus ( Linnaeus, 1758)) .

Albunea paretoi: Monod, 1956: 37–40 , figs. 2–9. – Buchanan, 1957: 56. – Sourie, 1957: 13, 31, 78. – Forest, 1958: 147 (list). – Gauld, 1960: 66 (list). – Rossignol, 1962: 124. – Fransen, 1991: 56, 77 (unjustified emendation) (not Albunea paretii Guérin Méneville, 1853 ).

Albunea paretii: Rossignol, 1957: 97 , fig. 12. – Holthuis and Manning, 1970: 251–252*. – Kaestner, 1980: 336 (part). – Türkay, 1982: 98, 101, 110. – d’Udekem d’Acoz, 1999: 171 (part) (not Albunea paretii Guérin Méneville, 1853 ).

? Albunea sp. A Lebour, 1959: 129 , fig. 14.

MATERIAL EXAMINED: Portugal: Cape

Verde Islands: La Praya, 10–30 m, 1883, coll. ‘‘Talisman’’: 2 Ƌ, 12.3–13.1 mm cl, 4 ♀, 13.4–18.4 mm cl, syntypes (MNHN­Hi 7); Sta. 7.V06, 16°10̍N, 22°57̍W, west coast, Boa Vista, Aug. 27–28, 1986, coll. CANCAP: 3 ♀, 14.5–19.6 mm cl ( RMNH 38604 ), 2 ♀, 17.9–19.4 mm cl ( AMNH 18078 ex RMNH 38604 ) ; Plage de Mindolo , Île de São Vicente, May 15, 1950, coll. J. Cadenat: 1 Ƌ, 12.7 mm cl (MNHN­Hi 12) ; Sta. 38, 7 m, coll.? ‘‘ Calypso’ ’: 1 ♀, 12.5 mm cl (MNHN­Hi 181) .

Senegal: Virage, Cape Vert, Dakar, April 12, 1982, coll. F. C. Roest: 5 Ƌ, 8.6–10.8 mm cl, 2 ♀, 11.7–12.9 mm cl ( RMNH 35926), 2 Ƌ, 9.8 mm cl, 1 ♀, 13.8 mm cl ( AMNH 18079 ex RMNH 35926).

Liberia: Sta. 289, Net 70B, 03°04̍45̎N, 16°62̍W, 0–132 m, Aug. 23–24, 1927, coll. R/V ‘‘Discovery’’: 1 zoea? V larva, 6.0 mm ( BMNH 1951.2.17.2355); Sta. 295, Net TYF, 05°30̍30̎N, 17°45̍W, 2500–2700 m (see below), Aug. 25, 1927, coll. ‘‘Discovery’’: 1 zoea? V larva, 7.0 mm ( BMNH 1951.2.17.2356).

Ghana: Manford, Gold Coast, coll. C. Hupfer: 1 Ƌ, 19.1 mm cl ( ZMH K–5136).

Ivory Coast: Grand Bassam, 1901, coll. V. Mayet: 2 ♀, 14.5–18.9 mm cl (MNHN­Hi 13).

Dahomey: Chenal de Cotonov, near Cotonou, July 19, 1963, coll. H. Hoestlandt: 3 ♀, 6.2–7.4 mm cl ( RMNH 20957).

Nigeria: West Mole , Lagos, June 16, 1963, coll. A. R. Longhurst: 2 unsexable, unmeasurable specimens ( RMNH 20959 ) ; Sta. 224, Lagos, May 9, 1965, coll. ‘‘ Pillsbury’ ’: 1 Ƌ, 11.5 mm cl ( RMNH 22664 ) .

Gabon: Sta. 277, 01°44̍S, 08°38̍E, 0–88 m, Aug. 7, 1927, coll. R/V ‘‘Discovery’’: 1 zoea? V larva, 5.9 mm ( BMNH 1951.2.17.2354).

Congo: Baie de Pointe­Noire, Jan. 1969, coll. unknown: 1 Ƌ, 11.9 mm cl (MNHN­Hi 14).

Zaire: Sta. 276, Net 100B, 05°54̍S, 11°19̍E, 0–110 m, Aug. 5, 1927, coll. R/V ‘‘Discovery’’: 2 zoea? V larvae, 6.5–6.7 mm ( BMNH 1951.2.17.2353).

DIAGNOSIS: Carapace wider than long, covered with lightly setose grooves. Anterior margin with 10–12 spines on either side of ocular sinus. Setal field with narrow lateral elements and straight anterior margin. CG1 with separate posterior lateral elements; CG4 with two to four short, anteriorly displaced medial elements between longer supralateral elements of CG4; CG5 of two oblique triangular elements; CG6 and CG7 fused; CG8 broken; CG11 present. Rostrum present, not reaching posterior margin of ocular plate. Ocular plate triangular. Distal peduncular segments dorsoventrally flattened and triangular in shape, tapering at tip, approximated along mesial margins, lateral margins convex, mesial margins straight. Cornea at tip. Dactylus of pereopod II with heel produced, tapered, and acute. Dactylus of pereopod III with heel broad, projecting, acute. Dactylus of pereopod IV sinuous from base to tip, with low rounded heel and shallow indent. Telson of male broadly triangular, tip broadly round­ ed, thickly calcified medially, inflated dorsally, distal two­thirds with lateral decalcified region, median row of thin setae. Telson of female flattened, ovate, and evenly calcified with slightly produced tip.

DESCRIPTION: Carapace (fig. 102A) slightly wider than long. Anterior margin slightly concave on either side of ocular sinus, becoming convex laterally with 10–12 large spines (n = 6) along length. Rostrum as small acute tooth, not reaching proximal margin of ocular plate. Ocular sinus smoothly concave, surface smooth or with one small spinule. Frontal region smooth; setal field narrow anteriorly, widening posteriorly; posterior lateral elements reduced to narrow bands of setae. CG1 parallel to anterior margin of carapace, sinuous, strongly crenulate, divided into medial fragment and curved, posteriorly displaced lateral elements. Mesogastric region smooth; CG2 present as two short medial elements; CG3 broken into six to eight short elements between posterior lateral elements of CG1; CG4 with two to four short, anteriorly displaced medial elements between longer supralateral elements of CG4. Hepatic region smooth, with oblique setose groove at median of lateral margin. Epibranchial region generally triangular, smooth; posterolateral margin with two or three tiny rows of setae. Metagastric region smooth; CG5 present as two oblique, triangular elements. CG6 strongly crenulate, strongly anteriorly concave medially and sloping out to anteriorly convex lateral thirds. CG7 oblique, reaching lateral margins of median segment of CG6. Cardiac region smooth; CG8 present as two short medial and two short lateral elements. CG9 present as two short lateral grooves with gap at midline. CG10 present as two short elements. CG11 present as one or two short medial elements. Post­CG11 element absent. Branchial region with numerous short, transverse rows of setae. Posterior margin deeply and evenly convex, with submarginal groove reaching twothirds up margin of posterior concavity. Branchiostegite with strong anterior submarginal spine; anterior region with scattered short, transverse lines ventral to linea anomurica; with many short rows of setae and sparsely covered with long plumose setae ventrally; posterior region membranous, with numerous irregular fragments and sparsely covered with long plumose setae.

Ocular plate (fig. 102B) triangular with shallow median indentation; median peduncular segments present as small ovate calcified areas lateral to ocular plate. Distal peduncular segments elongate, subtriangular, 0.27–0.32 length of carapace, with convex lateral and straight mesial margins, cornea covering distal tip; lateral margins with notch one­third distal from base; mesial margins approximated at base; mesial and lateral margins with long plumose setae; tuft of plumose setae at proximolateral ventral angle and ventromedial row of plumose setae extending from tuft to base of cornea.

Antennule (fig. 102C) with segment III narrow proximally, expanding distally to twice proximal width; with plumose setae on dorsal and ventral margins and sparsely scattered on lateral surface; dorsal exopodal flagellum with 100–120 articles (n = 6), long plumose setae on dorsal and ventral margins; ventral endopodal flagellum short with two articles (n = 6) and plumose setae on dorsal and ventral margins. Segment II medially inflated in dorsal view, with plumose setae on dorsal and ventral margins and scattered on ventrolateral third of surface. Segment I wid­ er than long, with small dorsodistal spine; dorsal third of lateral surface rugose, with long plumose setae; long plumose setae on dorsal and ventral margins.

Antenna (fig. 102D) with segment V ap­ proximately two times longer than wide, with long plumose setae on dorsal and ventral margins and scattered on lateral surface; flagellum with seven articles (n = 6), long plumose setae on dorsal, ventral, and distal margins. Segment IV expanded distally, with long plumose setae on dorsal, ventral, and distal margins, and row of setae on dorsolateral surface. Segment III with long plumose setae on dorsal and ventral margin and in short row on surface. Segment II short, widening distally, rugose, with plumose setae on margins and scattered on lateral surface; antennal acicle long, thin, and exceeding distal margin of segment IV by one­third length of segment IV, with long plumose setae on dorsal margin. Segment I rounded proximally, flattened ventrolaterally, with long plumose setae on margins and scattered in line of surface rugae behind spine; lateral surface with acute spine dorsodistally, with low semicircular dorsolateral lobe ventrodistal to spine; segment with ventromesial antennal gland pore.

Mandible (fig. 102E) incisor process with two teeth; cutting edge with one tooth. Palp three­segmented, with plumose setae on margins and long, thick, simple setae arising from bend in second segment and on distal margin of terminal segment.

Maxillule (fig. 102F) distal endite proximally narrow, widening to inflated distal end, with thick simple setae on distal margin and thin simple setae on dorsal margin. Proximal endite with thick simple setae on distal margin. Endopodal external lobe truncate distally and curled under; internal lobe reduced with three thick setae at distolateral margin.

Maxilla (fig. 102G) exopod evenly round­ ed, with plumose setae along distal margin. Scaphognathite bluntly angled on posterior lobe, with plumose setae.

Maxilliped I (fig. 102H) epipod with plumose setae on margins, distolateral surface and mesial surface. Endite tapered distally and subequal to first segment of exopod. Exopod with two segments; proximal segment narrow, margins parallel with plumose setae; distal segment spatulate, longer than wide, broadest medially, margins and mesioventral surface with long plumose setae. Endopod flattened and elongate, reaching two­thirds to distal end of proximal exopodal segment; plumose setae on margins and median of lateral surface.

Maxilliped II (fig. 102I) dactylus evenly rounded, length slightly greater than width, with thick simple setae distally and on distolateral surface. Propodus 1.5 times wider than long, slightly produced at dorsodistal angle, with plumose setae on dorsal margin and patch of long simple setae on lateral surface and ventrolateral angle. Carpus not produced dorsodistally, approximately two times longer than wide; long simple setae on dorsal margin. Merus approximately three times longer than wide, margins parallel; with simple and plumose setae on margins and scattered on surface. Basis­ischium incompletely fused with plumose setae on margins. Exopod one­half longer than merus, flagellum with one elongate article, approximately as long as carpus is wide.

Maxilliped III (fig. 102J) dactylus with rounded tip; long plumose setae on margins and lateral surface. Propodus dorsodistally inflated, with longitudinal median row of plumose setae on lateral surface; margins with plumose setae. Carpus produced onto propodus almost one­fourth length of propodus; lateral surface with two rows of plumose setae on surface, plumose setae on margins. Merus inflated, unarmed, with plumose setae on margins and scattered on lateral surface. Basis­ischium incompletely fused, with weak crista dentata of two teeth. Exopod two­segmented: proximal segment small; distal segment styliform, tapering, approximately one­third length of merus; with plumose setae on margins; without flagellum.

Pereopod I (fig. 103A) dactylus curved and tapering; lateral and mesial surfaces smooth; dorsal margin with long plumose and short simple setae; ventral margin with short simple setae. Propodal lateral surface with numerous short, transverse rows of setose rugae; dorsal margin unarmed; ventral margin distally produced into acute spine; cutting edge lacking teeth, lined with long plumose setae; dorsal margin with long plumose setae, ventral margin with short simple setae. Carpus with dorsodistal angle produced into strong corneous­tipped spine; dorsal margin with few short, transverse grooves behind spine; dorsal and distal margins with long plumose setae; lateral surface with small distal rugose area, few transverse setose ridges on distal half of surface; mesial surface smooth, with medial transverse row of setae, margins with long plumose setae. Merus unarmed; lateral surface with scattered transverse rows of long plumose setae, margins with long plumose setae; mesial surface with few scattered setae; fully calcified. Basis­ischium incompletely fused, unarmed. Coxa of male with anterior spine; coxa of female unarmed.

Pereopod II (fig. 103B) dactylus smooth; base to heel concave, heel produced and acute, heel to tip with narrow, acute indent, tip acute, tip to base broadly convex; lateral surface smooth, with several small tufts of short setae in generally straight line across medioproximal surface, several widely spaced submarginal tufts of short setae dorsodistally; mesial surface smooth, ventral margin with long plumose setae, dorsal margin with short simple setae and patch of long plumose setae at base. Propodal dorsal surface smooth, with ventral margin inflated and rounded; oblique row of long plumose setae on distal margin of lateral surface; distal and ventral margins with long plumose setae; dorsolateral surface as narrow, oblique, flattened shelf, with short setae on dorsal margin and long plumose setae on ventral margin; mesial surface with elevated, curved setose ridge from ventral junction with dactylus almost to ventral proximal junction with carpus. Carpus strongly produced and rounded dorsodistally, dorsal margin smooth; lateral surface smooth, with small setose mat at tip of produced area and irregular, interrupted row of rugae and submarginal elevated ridge ventrally, rugae and ridge with long plumose setae; margins with short plumose setae; mesial surface smooth with row of long plumose setae distally and subdorsally. Merus with large median decalcified window covering nearly all of lateral surface, with few scattered long plumose setae on surface and margins; mesial surface nearly smooth, with two long rows of setae. Basis­ischium incompletely fused and unarmed. Male coxa with anterior margin spine; female coxa without spine.

Pereopod III (fig. 103C) dactylus with base to heel concave, heel acutely produced, heel to tip with broadly concave indent and sinuous proximal half of margin, tip acute, tip to base smoothly convex; lateral surface smooth, with several small tufts of short setae in generally straight line across medioproximal surface, dorsodistal margin with tufts of short setae; ventral margin with long plumose setae, dorsal margin with short simple and plumose setae; mesial surface smooth with plumose setae proximally at junction with propodus. Propodus not inflat­ ed dorsoventrally; lateral surface smooth, with long plumose setae in oblique row, simple setae on dorsal margin; dorsolateral surface narrow, oblique, flattened, with setose mat; mesial surface smooth. Carpus produced dorsodistally, only slightly exceeding proximal margin of propodus; dorsolateral margin unarmed; lateral surface slightly rugose dorsodistally, with thin mat of short setae and two interrupted rows of setae ventrally; mesial surface smooth, with long plumose setae on distal margin and in oblique row on surface. Merus smooth, with large decalcified window covering nearly half of lateral surface medially; dorsal and ventral margins unarmed but with long plumose setae; distolateral margin with long plumose setae; mesial surface smooth. Basis­ischium incompletely fused and unarmed. Male coxa with spine on anterior margin; female lacking spine. Female with large gonopore on anterior mesial margin of coxa, surrounded with short plumose setae; male with small pore.

Pereopod IV (fig. 103D) dactylus with base to tip convex proximally, concave distally, indent smoothly joined with margin, tip acute, tip to base concave distally to convex proximally; lateral surface smooth, ventral margin with long plumose setae, dorsal margin with short simple setae; mesial surface with dorsal decalcified region, demarcated ventrally by longitudinal elevated ridge with row of short setae; with setose punctations ventral to decalcified window. Propodus expanded dorsally and ventrally; ventral expansion not reaching ventral margin of dactylus, margin with long plumose setae; dorsal expansion with row of long plumose setae dorsally, oblique area with mat of short simple setae; lateral and mesial surfaces smooth. Carpus slightly produced dorsodistally; ventral three­fourth of lateral surface and mesial surface smooth, dorsodistal quarter of lateral surface with mat of short setae; dorsal margin with short simple and long plumose setae; ventral margin with short simple setae; mesial surface decalcified medially. Merus with scattered short transverse rows of setae on lateral surface, dorsal and ventrodistal margins with long plumose setae; proximoventral half of mesial surface with large decalcified window. Basis­ischium incompletely fused and unarmed. Male coxa with spine on anterior margin; female coxa unarmed.

Abdomen (fig. 103E) with somite I length and width subequal, widest posteriorly; dorsal surface with anterior margin straight; posterior margin curved with elevated submarginal row of short setae; small transverse, decalcified windows laterad of segment medi­ an. Somite II dorsal surface with submarginal transverse ridge anteriorly; with small transverse, decalcified windows laterad of segment median just anterior to submarginal ridge; pleura expanded and directed anterolaterally; lateral margins rounded, anterior and lateral margins with long plumose setae, posterior margin with short setae; posteromesial angle with mat of short simple setae. Somite III similar to somite II, but narrower, shorter; pleura thinner and shorter than on somite II, directed anterolaterally, with setae as in somite II; anterolateral angle subacute; dorsal surface obliquely flattened anterolaterally. Somite IV similar to somite III, but thinner and shorter; dorsal surface with few short setae anterolaterally; pleura thinner and shorter than on somite III, directed posterolaterally; dorsal surface obliquely flattened anterolaterally; margins with long plumose setae. Somite V wider than somite IV; lateral margins with plumose setae; pleura absent. Somite VI slightly broader than somite V; dorsal surface with short transverse rows of setae laterad of midline; pleura absent.

Females with uniramous, paired pleopods on somites II–V; males without pleopods.

Telson of male (fig. 103F) broadly triangular, slightly longer than wide, with broadly rounded tip; thickly calcified medially, inflated dorsally; distal two­thirds with lateral decalcified region; median longitudinal groove extending one­half length, row of long simple setae of either side of median groove beginning at distal end and continu­ ing almost to distal margin of telson; proximolateral angles with patch of long simple setae; margins with long simple setae. Telson of female (fig. 103G) flattened, ovate, and evenly calcified with slightly produced tip; median groove similar to male, setal row from midpoint of median groove to near distal margin of telson with simple setae approximately one­fourth length of those on male; proximolateral angle with patch of setae, margins with long simple setae.

DISTRIBUTION: Cape Verde Islands and Senegal southward to Pointe­Noire, Congo, in up to 30 m depth.

MAXIMUM SIZE: Males: 19.1 mm cl; females: 19.6 mm cl.

TYPE SPECIMENS: MNHN­Hi 7 (6 syntypes).

TYPE LOCALITY: La Praya, Cape Verde Islands, Portugal, 10–30 m depth.

REMARKS: A. Milne Edwards and Bouvier (1898) erroneously cited this species as having a maxilliped III with one pleurobranch, two arthrobranchs, and lacking an epipod, and also pereopod V having two arthrobranchs. These authors subsequently (1900) correctly stated that there are no arthrobranchs on pereopod V, but they repeated the inaccuracies regarding maxilliped III. This error became propagated through time, as Balss (1957) repeated A. Milne Edwards and Bouvier’s (1900) incorrect statements and applied them to the genus as a whole in his monumental treatise on the Decapoda .

The west African larvae reported by Lebour (1959) as ‘‘ Albunea sp. A’’ strongly resemble the late stage (V or VI) zoeae of A. catherinae , n. sp., a closely related species ( Kurata, 1970). This suggests that these larvae are A. elegans , rather than the sympatric, but more distantly related, A. carabus . Several lots of larval specimens, probably stage V zoeae collected by R/V ‘‘Discovery’’ from east African localities, are referred to this species as they are extremely similar to the described larvae of A. paretii , another taxon closely related to A. elegans . The collection depth of 2500–2700 m for a putative larvae of this species from off Liberia ( BMNH 1951.2.17.2356) is a suspect depth for any albuneid larvae, as the deepest living adults are known in no more that 225 m of water.

Although this species is similar to A. par­ etii, and for many years was confused with that western Atlantic species, it is not as closely related to A. paretii , as is the Pacific A. lucasia . Albunea elegans is easily distinguished from A. catherinae , n. sp. by the shapes of the dactyli of pereopods III and IV. It can be separated from A. paretii by the longer length of the branchiostegite spine, broken CG8, and the different shape of the dactylus of pereopod III.

Albunea paretii Guérin Méneville, 1853 Figures 104 , 105

Albunea oxyophthalmus White, 1847: 57 (nomen nudum)*.

Albunea symnista [sic]: Gibbes, 1850a: 24 * (not Albunea symmysta ( Linnaeus, 1758)) .

Albunea Paretii Guérin Méneville, 1853: 47–48 , pl. 1, fig. 10*. – Bolivar, 1875: 21. – Seridji, 1988: 1298.

Albunaea [sic] oxyophthalma: Stimpson, 1858: 230 (list).

Albunaea [sic] Paretii: Stimpson, 1858: 230. – Stimpson, 1859: 78 (list).

Albunea oxyophthalma Miers, 1878: 329–330 , pl. 5, figs. 14, 15*. – Rathbun, 1897: 42 *. – Benedict, 1901: 139. – Verrill, 1901: 61–62, pl. 8, fig. 1*. – Verrill, 1908: 438, pl. 28, fig. 1*. – Gordon, 1938: 187 (part)*. – Monod, 1939: 560–561, fig. 5*. – Bouvier, 1940: 181. – Snodgrass, 1952: fig. 11a, d.

Albunea paretii: Miers, 1878: 330 . – Rodrigues da Costa, 1962: 6–7, pl. 1, figs. 5–7, pl. 3, figs. 1–3. – Williams, 1965: 137–138 (part)*. – Coêlho, 1966: 244. – Fausto Filho, 1967: 12– 13. – Coêlho and Ramos, 1972: 176 (part). – Gomes Corrêa and da Silva Brum, 1980: 60. – Kaestner, 1980: 336 (part). – Markham and McDermott, 1981: 1271. – Williams, 1984: 249–250 (part)*. – Chace et al., 1986: 338–339, fig. 112. – Calado, 1987: 106–118, pls. 9–12*. – Coêlho and Calado, 1987: 42–43, table 1. – Markham, 1988: 30. – Calado et al., 1990: 747 (part), fig. 2a, b*. – Hernández and Bolaños, 1995: 77. – Calado, 1995: 51–55, pl. 4, fig. f, pl. 5, fig. e, pl. 13, fig. a, pl. 14, figs. a, b, pl. 15, figs. a–e, pl. 16, figs. a–c*. – Calado, 1997a: 17. – Spivak, 1997: 74 (list). – Calado, 1998: 407. – Markham and Boyko, 1999: 4*. – d’Udekem d’Acoz, 1999: 171 (part). – Amaral et al. in Nucci et al., 2001: 479. – Morgado et al. in Nucci et al., 2001: 479.

Albunea pareti [sic]: Ortmann, 1896: 225 (list). – Ortmann, 1901: 1275. – Moreira, 1901: 30, 88 (list). – Boschi, 1981: 715, 740.

Figures

Fig. 100. Albunea occultus, n. sp.: A–J, ♀, 25.0 mm cl, AM P24477, paratype. A. Carapace, branchiostegite, and ocular peduncles, dorsal view. B. Ocular peduncles, dorsal view. C. Left antennule, lateral view. D. Left antenna, lateral view. E. Left mandible, mesial view. F. Left maxillule, lateral view. G. Left maxilla, lateral view. H. Right maxilliped I, lateral view. I. Left maxilliped II, lateral view. J. Left maxilliped III, lateral view. Scale = 3.3 mm (B, E, F), 4.4 mm (I), 6.7 mm (C, D, G, H, J), and 7.4 mm (A).

Fig. 101. Albunea occultus, n. sp.: A–E, ♀, 25.0 mm cl, AM P24477, paratype; F, Ƌ, 17.5 mm cl, AM P23809, paratype; G, ♀, 21.0 mm cl, AM P19993, paratype. A. Left pereopod I, lateral view. B. Left pereopod II, lateral view. C. Left pereopod III, lateral view. D. Left pereopod IV, lateral view. E. Abdominal somites I–VI, dorsal view. F. Telson of Ƌ, dorsal view. G. Telson of ♀, dorsal view. Scale = 3.3 mm (F, G), 6.7 mm (B–D), 7.4 mm (E), and 8.0 mm (A).

Fig. 102. Albunea elegans A. Milne Edwards and Bouvier, 1898: A–J, Ƌ, 10.4 mm cl, AMNH 18079. A. Carapace, branchiostegite, and ocular peduncles, dorsal view. B. Ocular peduncles, dorsal view. C. Left antennule, lateral view. D. Left antenna, lateral view. E. Left mandible, mesial view. F. Left maxillule, lateral view. G. Left maxilla, lateral view. H. Left maxilliped I, lateral view. I. Left maxilliped II, lateral view. J. Left maxilliped III, lateral view. Scale = 1.6 mm (B, E, F), 2.2 mm (I), and 3.3 mm (A, C, D, G, H, J).

Fig. 103. Albunea elegans A. Milne Edwards and Bouvier, 1898: A–F, Ƌ, 10.4 mm cl, AMNH 18079; G, ♀, 13.8 mm cl, AMNH 18079. A. Right pereopod I, lateral view. B. Left pereopod II, lateral view. C. Right pereopod III, lateral view. D. Left pereopod IV, lateral view. E. Abdominal somites I– VI, dorsal view. F. Telson of Ƌ, dorsal view. G. Telson of ♀, dorsal view. Scale = 2.2 mm (F, G), 3.3 mm (B, D, E), and 4.4 mm (A, C).

Fig. 104. Albunea paretii Guérin Méneville, 1853: A–D, H–J, ♀, 11.5 mm cl, AMNH 18089; E– G, ♀, 7.0 mm cl, USNM 260941. A. Carapace, branchiostegite, and ocular peduncles, dorsal view. B. Ocular peduncles, dorsal view. C. Left antennule, lateral view. D. Left antenna, lateral view. E. Left mandible, mesial view. F. Left maxillule, lateral view. G. Left maxilla, lateral view. H. Left maxilliped I, lateral view. I. Left maxilliped II, lateral view. J. Left maxilliped III, lateral view. Scale = 1.6 mm (B, E, F), 2.2 mm (G, I), 3.3 mm (D, H, J), and 4.4 mm (A, C).

Fig. 105. Albunea paretii Guérin Méneville, 1853: A–E, G, ♀, 11.5 mm cl, AMNH 18089; F, Ƌ, 11.5 mm cl, AMNH 18089. A. Left pereopod I, lateral view. B. Left pereopod II, lateral view. C. Left pereopod III, lateral view. D. Left pereopod IV, lateral view. E. Abdominal somites I–VI, dorsal view. F. Telson of Ƌ, dorsal view. G. Telson of ♀, dorsal view. Scale = 3.0 mm (F, G), 3.3 mm (A), and 4.4 mm (B–E).