Clivina molucca, Balkenohl, 2021

Balkenohl, Michael, 2021, Belgian Journal of Entomology, Belgian Journal of Entomology 115, pp. 1-83 : 40-42

publication ID

https://doi.org/ 10.5281/zenodo.5645953

DOI

https://doi.org/10.5281/zenodo.5645869

persistent identifier

https://treatment.plazi.org/id/961387D9-C821-FFC3-FDED-F973FC8CF73C

treatment provided by

Plazi

scientific name

Clivina molucca
status

sp. nov.

Clivina molucca View in CoL sp. nov.

urn:lsid:zoobank.org:act:F32E1EB6-11CD-408E-AD77-1713A536D0FE

( Figs 28 View Fig , 48 View Fig , 68 View Fig , 82 View Fig , 95 View Fig , 112 View Fig )

ETYMOLOGY. The name is a noun in the nominative case and refers to the Molucca Islands.

TYPE MATERIAL. Holotype: • ♂, with labels and data: white, black printed “ MALUKU, Seram 35 km E Pasahari 24–30.x.1998 J. Horák leg.” ( CADW) .

Paratypes: • 1 ♂, 2 ♀♀, same data as holotype ( CADW, CMBB) ; • 2 ♂♂, “ MALUKU, Seram UnitO. 35 km. Eof Pasahari , 24–30.x.1998. O. Mehl, leg. ( CMBM) ; • 3 ♂♂, 3 ♀♀, MALUKU, SERAM Solea 12 km SE Wahai , 17.i.–6.ii.1997, J. Horák leg. / Reich. castanea (Westw.) det. Dr. A. Dostal 1997 ( CADW) ; • 1 ♀, same data but 16.x–4.xi.1998, Ole Mehl, leg. ( CMBM) ; • 1 ♀, MALUKU, Ambon Laithatu, Soya 500 m, 8.xi.1998. O. Mehl leg. ( CMBM) ; • 2 ♀♀, same data but 16–21.x.1998 ( CADW) ; • 2 ♂♂, 1 ♀, same data but 31.x.–4.xi.1998 ( CADW) .

REMARK. Many of the specimens including the holotype exhibit some micro scratches on the head.

DIAGNOSIS. Amedium sized species with smooth lateral margin of the pronotum and elytron, with isodiametric reticulation on interval eight of the elytron, and triangle-like acute angled lobe of the mentum with slightly concave anterior margin, and with the median tooth much less projecting as the lobe. Asimilar combination of characters shows C. agona Putzeys. C. agona differs mainly by the size and the distinct lateral group of punctures in the basal half of the pronotum.

DESCRIPTION. Measurements. Holotype: Body length 8.11 mm, width 2.4 mm; ratio length/width of pronotum 0.93; ratio length/width of elytra 1.88. Other material: Body length 7.85–8.43 mm (: = 8.07 mm *), width 2.39-2.57 mm (: = 2.44 mm *), ratiolength/width of pronotum 0.9–0.94 (: = 0.92*), ratiolength/widthofelytra 1.81–1.89 (: = 1.84*); (*n = 10).

Colour: Glossy. Piceous. Intermediate and hind legs dark fuscous, antennae and tarsomeres fuscous.

Head: Athird narrower than pronotum. Clypeus slightly excised anteriorly; wing prominent, as protruding as clypeus, separated from clypeus by distinct notch; supraantennal plate convex; clypeus laterally with carina, wing amargined, supraantennal plate reflexed margined. Supraantennal plate extended up to mid-eye level, separated from wing by distinct notch. Clypeus slightly transverse, convex and glossy in posterior three quarters, in anterior quarter with isodiametric reticulation, separated from frons by distinct furrow. Frons moderately convex, with indistinct impression at middle, with blunt lateral carina at posterior eye-level. Clypeus and frons smooth, with minute scattered punctures, separated from wing and supraantennal plate by deep moderately wide furrow continuing posteriorly as wide supraorbital furrow with two supraorbital setigerous punctures at posterior eye level. Neck constriction developed as small step, with row of small punctures. Eye projecting, somewhat elongated in dorsal view. Gena covering eye by about a quarter in ventral view. Antenna just reaching posterior setaof pronotum, slightly flattened, antennomeres four to ten of moderate length (ratio L/ W 1.8 –1.88). Labrum indistinctly convex anteriorly, obtusely pointed at middle, with isodiametric reticulation, seven setose. Mentum ( Fig. 48 View Fig ) with distinct isodiametrically reticulated surface, with lobe concavely hollowed out, shape of lobe like a conspicuously askew trapezium, with indistinctly convex lateral margin, with slightly sinuate anterior margin and acute-angled lateral anterior angle, carinate laterally and anteriorly, with medial angle obtuse and indistinct, with short carina at middle, median tooth wide, distinctly less protruding anteriorly as lobe, straight anteriorly with small pointed knob at middle.

Pronotum ( Fig. 68 View Fig ): Disk moderately convex in lateral and frontal view. Slightly wider than long. Reflexed lateral margin smooth, attenuate and indistinctly convex in anterior half, widest in posterior third; anterior angle obtuse, not projecting; posterior angle marked by small distinct and rounded tooth, tooth not protruding laterally over convexity of lateral margin. Lateral channel narrow, reticulated. Median line narrow, line-like; anterior transverse line minutely punctured, joining median line, just joining basal constriction. Surface smooth, with some transverse wrinkles and longitudinal wrinkles at middle, with distinct isodiametric reticulation in basal quarter and laterally with wide band of reticulation reaching nearly up to anterior angles.

Elytron: Disk flattened in anterior third in lateral view, convex on frontal view. Nearly twice as long as wide, slightly dilated in apical half with maximum width behind middle. Reflexed lateral margin smooth. Scutellar striole distinct; setigerous tubercle at base of first stria, with distinct tubercle at base of third interval. Striae moderately deep, punctuate-striate, one to three free at base, four to seven joining at humerus, one running up to apex, just not joining with seven, two ending free at apex, five and six joining apically, often three and four also, striae wide and deep apically. Intervals distinctly convex, all intervals distinctly convex at apex, eights with carina at humerus up to the end of the first quarter and in apical quarter. Third interval with four setigerous punctures, first situated at second stria near base, the other three approaching third stria. Surface of intervals glossy, interval eight with distinct isodiametric reticulation, interval seven with isodiametric reticulation in apical quarter, without reticulation at humerus.

Hind wings: Fully developed.

Lower surface: Proepisternum covered in lateral half with distinct isodiametric reticulation, with rough imbricate reticulation towards middle, with some transverse wrinkles ( Fig. 82 View Fig ). Terminal sternite of abdomen developed apically as flat moderately wide transverse fold sulcus.

Legs: Legs covered with longitudinal reticulation. Protibia with three spines of moderate length, not sulcate dorsally, movable spur slightly arcuate in basal three quarters, distinctly arcuate apically.

Male genitalia ( Fig. 95 View Fig ): Median lobe long, slender, slightly arcuate, apex moderately long, broad, regularly attenuated apically, pointed at apex, apex slightly turned ventrally, ventral rim moderately developed, base with apophysis. Endophallus apically with small group of minute bristles. Ventral paramere moderately wide, moderately arcuate, lateral apophysis medium sized, apex wide, with one long and two small setae at apex, with one additional moderately long seta preapically. Dorsal paramere strongly developed, at apex with one long seta and three moderately long setae, with two additional small setae in apical half.

Female coxostylus and laterotergite ( Fig. 112 View Fig ): Coxostylus slender, dorsally with ten long to medium sized and one distinctly elongated and robust nematiform setae in basal half, ventrally with three long setae arising from stepped knobs and with numerous smaller setae, with one minute SSO. Laterotergite with six moderately long setae basally and one isolated seta towards coxostylus.

VARIATION. On the frons of the head some of the paratypes exhibit a flat concave dint. On the pronotum, the lateral band of isodiametric reticulation is distinctly extended to the middle of the disk in two of the paratypes. In one male paratype, the second seta on the third interval is missing on both sides, in one female paratype this seta is missing on one side.

DISTRIBUTION. Known from the Molucca Islands.

CADW

CADW

CMBB

CMBB

CMBM

CMBM

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Carabidae

Genus

Clivina

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