Lagocheirus plantaris plantaris Erichson, 1847

Santos-Silva, Antonio, Nascimento, Francisco E. De L. & Kozlov, Anton Olegovich, 2019, The megadiverse fauna of Neotropical Cerambycidae (Coleoptera): Notes, descriptions, new records, new species, and revalidations, Zootaxa 4603 (3), pp. 441-472 : 446

Lagocheirus plantaris plantaris Erichson, 1847

( Figures 18–21 )

Lagochirus plantaris Erichson, 1847: 144 ; White, 1855: 365; Gemminger, 1873: 3148; Aurivillius, 1923: 392 (cat.); Soukup, 1942: 309 (distr.); Blackwelder, 1946: 612 (checklist).

Lagocheirus plantaris plantaris ; Dillon, 1957: 157; Monné & Giesbert, 1994: 247 (checklist); Monné, 1995: 45 (cat.); Martínez, 2000: 92 (distr.); Monné, 2005: 52 (cat.); Monné & Hovore, 2006: 183 (checklist); Wappes et al., 2006: 35 (distr.); Maes et al., 2010b: 471 (part; figs on page 472); Monné et al., 2012: 26 (distr.); Monné & Chaboo, 2015: 76 (distr.).

Lagocheirus plantaris ; Gilmour, 1965: 564 (cat.); Chemsak et al., 1992: 138 (checklist); Barbosa et al., 2009: 293 (distr.); Lagos & Barrios, 2014: 17 (distr.); Lanuza-Garay & Barrios, 2015: 66 (host); Lingafelter et al., 2017: 153.

Trypanidius fasciculatus White, 1855: 377 .

Lagochirus fasciculatus ; Gemminger, 1873: 3138 (cat.); Aurivillius, 1923: 392 (cat.); 1926: 6 (distr.).

Lagocheirus fasciculatus ; Bates, 1863a: 101.

Lagoschyrus fasciculatus ; Zischka, 1948: 7 (distr.).

Lagocheirus plantaris indistinctus Dillon, 1957: 158 ; Gilmour, 1965: 564 (cat.); Chemsak et al., 1992: 138 (checklist); Monné & Giesbert, 1994: 247 (checklist); Monné, 1995: 46 (cat.); Toledo, 1998: 31; Monné, 2005: 52 (cat.); Monné & Hovore, 2006: 183 (checklist); Swift et al., 2010: 38 (distr.); Lanuza-Garay, 2015: 10 (morphol.); Monné, 2018b: 64 (cat.). Syn. nov.

According to Dillon (1957) on Lagocheirus plantaris indistinctus : “Male. In many instances only slightly distinct from the typical form in having the basal angles of the scutellum pale or ochraceous pubescent. However, in well-marked examples the elytral pubescence is uniformly dull cinereous or fuscous, with the carina marked with ochraceous. The most important distinguishing feature is to be found in the tubercles of the pronotum, which is scarcely inflated. As consequence, the lateral tubercles are usually conical, and the tubercles of the disk are more pronounced.” However, comparing photograph of the holotype with several male specimens from South America, we see that the differences pointed out by Dillon (1957) do not exist, especially that reported as “most important”, because all males of Lagocheirus plantaris plantaris have the pronotum inflated, but it can be from slightly inflated to noticeably inflated. It is exactly how pronotum is inflated which cause the tubercles to be higher or lower. As for the shape of the lateral tubercles of the prothorax, the difference pointed out also does not exist. Furthermore, the shape of the lateral tubercle in several examined males are variable, from slightly rounded at apex to distinctly rounded and, in some of them, identical to that in the holotype of L. p. indistinctus. As for the difference in the pubescence of the scutellum, although nearly all specimens from South America do not have the basal sides with contrasting pubescence, it is present in some specimens; furthermore, it is not present in many specimens from Central America, independently of the pronotal shape, making clear that this is a variable feature in the species. Finally, the elytral pubescence is distinctly variable in the specimens examined, independently of the place from where the specimen is. Toledo (1998) also reported that the scutellum in Lagocheirus plantaris indistinctus has its sides with light pubescence. Unfortunately, he did not point out if he saw variation in this feature.

Curiously, what neither Dillon (1957) nor Toledo (1998) reported is that the pronotum in females is always less inflated than in all males, making the pronotal tubercles distinctly more elevated, and the lateral tubercles of the prothorax more distinctly conical, as in the holotype of Lagocheirus plantaris plantaris (see photograph of the holotype in Bezark 2018a).

Based on the absence of reliable differences between Lagocheirus plantaris plantaris and L. p. indistinctus, the latter is formally considered a junior synonym of the former.

Maes et al. (2010b) recorded L. plantaris indistinctus for Nicaragua, based on a single specimen: “ Nicaragua: RAAN (Zelaya): Cerro Saslaya: Camp 3, UTM 713150 – 1521450, 950 m, IV-1999, col. J.M. Maes & B. Hernandez (1 ex. col. Museo Entomológico de León).” Although it is very probable that L. plantaris really occurs in Nicaragua, the specimen illustrated in Maes et al. (2010b) is a female of L. binumeratus Thomson, 1860. Accordingly, for now, L. plantaris plantaris is formally excluded from the fauna of Nicaragua. As the work also illustrated two specimens of the true L. p. plantaris , it needs to be listed under the references of both species as “part”. The current distribution of the species can be seen in the map ( Fig. 84 ).

Material examined (all belonging to the MZSP, Fig. 84 ). COLOMBIA, Huila: Gigante , 1 female, IV.1974 ; Caquetá, 1 male; Amazonas ( PNN Amacayacu, Matamata , 3º23’S 70º6’W, 150m, Malaise trap), 1 male, 27.III– 3.IV.2000, A. Parente col. ; Boyacá: Muzo , 1 female, 1930, H. Apolinar, Maria col. ; Cundinamarca ( Reserva Bosque Guajira, 4°47’34”N 75º46’60”W, 2,910M), 1 female, 8–13.III.2018. PERU, Satipo, 1 male, IV (no year indicated), A. Maller col.; 1 male, III (no year indicated), A. Maller col.; 1 male, IV.1945, A. Maller col.; Loreto: Estirón (Rio Ampiyacu) 1 male, 15–22.V.1966, Malkin col.; Ucayali: Pucallpá , 1 female, 15.VIII.1974. BRAZIL, Amazonas: Benjamin Constant (Rio Jatay, alto Amazonas), 4 males, 5 females, XI.1960, Dirings col.; 1 female, II.1960; 1 male, X.1961, Dirings col.; 1 male, 1 female, XI.1961, Dirings col.; 1 male, 1 female, III.1962, Dirings col.; 1 male, IX.1963, Dirings col.; 1male, XI.1963, Dirings col.; 1 male, XII.1963, Dirings col.; 1 male. S„o Paulo de Olivença ( Rio Solimıes ), 1 female, XI.1960, Dirings col. ; Manicoré ( Rio Madeira ), 1 male, VI. 1921, J.F. Zikán col. BOLIVIA, La Paz: Larecaja (Guanay, Uyapi ), 1 female, X–XI.1992 . PARAGUAY, Concepción: Horqueta , 1 male, Alberto Shultze col. PANAMA, Panama: Isla Barro Colorado , 1 female, 3.VIII.1961, J.M. Campbell col. ; 1 male, 8.XII.1963, L.J. Bottime col. ; 1 female, XI.1965, H. Britski col.; Capira ( Cerro Campana , 2900 m), 1 male, 31.VII.1970, J.M. Campbell col.

Figures

FIGURES 12–21. 12–16) Holonotus latithorax: 12) Ventral habitus, female; 13) Lateral habitus, female; 14) Dorsal habitus, lectotype female; 15) Labels, lectotype female; 16) Elytral apex, male. 17) Holonotus laevithorax, elytral apex, male. 18–21) Lagocheirus plantaris plantaris: 18) Dorsal habitus, male; 19) Head and prothorax, male; 20) Dorsal habitus, female; 21) Head and prothorax, female. Figures 14–15 by Jiří Pirkl.

FIGURES 80–84. 80) Obrium costaricum, female, dorsal habitus. 81) Pygmodeon obtusum, male, dorsal habitus. 82) Pantonyssus nigriceps, female, dorsal habitus. 83) Stizocera geniculata, female, dorsal habitus. 84) Map of the geographical distribution of Lagocheirus plantaris plantaris.