Ilyocoris andancensis, JATTIOT & TRINCAL & GARROUSTE & NEL, 2023

JATTIOT, ROMAIN, TRINCAL, VINCENT, GARROUSTE, ROMAIN & NEL, ANDRÉ, 2023, A new fossil species of the saucer bug genus Ilyocoris (Heteroptera, Naucoridae) from the Upper Miocene maar paleolake of la Montagne d’Andance (France), Palaeoentomology 6 (3), pp. 260-267 : 261

Ilyocoris andancensis sp. nov.

( Figs 1–4 ) urn:lsid:zoobank.org:act:3EB380F0-1632-4971-A5ED-7DFE8A2DC9FA

Material. Holotype MNHN-F.A71364 (Romain Jattiot coll.), paratypes MNHN-F.A71365, and MNHNF.A71366 (all compressed mummies), stored at the Muséum National d’Histoire Naturelle , Paris, France.

Diagnosis. Adult characters only. Color patterns identical on the three fossils. Connexivum brownish along posterior part and yellowish along anterior part; green-yellow anterior part of pronotum clearly narrower than black posterior part; outer margin of embolium black.

Type locality and horizon. Lower limit of Turolian to Tortonian-Messinian boundary, 7.30 ± 0.15 Ma ( Pastre et al., 2004), ‘ La Montagne d’Andance’, Saint-Bauzile village , Ardèche, France .

Description (based on all specimens). Head transverse; ocelli absent; eyes sessile; labium short and thick, arising near apex of head, conical, obviously segmented, without transverse sulcation; labium; apex of head without a laminated extension; vertex less than two times and half broader than transversal diameter of compound eye; head, pronotum, and scutellum shiny; ground color black and green-yellow; brown dots on head, some of which form two longitudinal, parallel, median lines, each of them widening posteriorly; anterior margin of pronotum straight behind interocular space; lateral pronotal margins limited inside by a strong sunken line; anterior part of pronotum green-yellow, and posterior part black; scutellum black; no meso- and metasternal midventral carinae (observed on holotype by partly removing the mummy); macropterous, hemelytra olive brown, opaque, very finely and densely punctate, clavus and margin paler, embolium with a less dense punctuation than the rest of hemelytra but with outer margin black; membrane without venation; fore legs raptorial, fore femur only moderately incrassate with inner antero-basal angle not produced, forming a continuous curve with trochanter, tarsus without a fringe of setae; hind legs flattened; tibiae with strong brownish spines and long swimming setae. Legs pale yellow; apex of abdomen without respiratory processes; connexivum brownish along posterior part and yellowish along anterior part.

Dimensions. Body 14.7 mm long, 6.4 mm wide (MNHN-F.A71364), 12.0 mm long, 6.0 mm wide (MNHN-F.A71365), 13.5 mm long, 6.0 mm wide (MNHN-F.A71366); head 1.3 mm long, 3.4 mm wide (MNHN-F.A71364); pronotum 2.0 mm long, 4.1 mm wide (MNHN-F.A71364); abdomen 9.7 mm long, 6.5 mm wide (MNHN-F.A71364); fore tibia 2.1 mm long, 0.9 mm wide; hind tibia 3.0 mm long, tarsus 4.1 mm long, hemelytra 8.6 mm long (MNHN-F.A71364).

Figures

FIGURE 1. Ilyocoris andancensis sp. nov., holotype MNHN-F.A71364. Photographs of general habitus. A, Dry. B, Under alcohol. Scale bars = 2 mm.

FIGURE 2. Ilyocoris andancensis sp. nov., holotype MNHN-F.A71364. Photographs of apex of abdomen. A, Under normal light, dry. B, Under UV light showing ovipositor, dry. Scale bar = 2 mm.

FIGURE 3. Ilyocoris andancensis sp. nov., paratype MNHN-F.A71365. Photographs. A, Habitus, dry. B, Head and right fore leg, dry. Scale bars = 2 mm.

FIGURE 4. Ilyocoris andancensis sp. nov., paratype MNHN-F.A71366. Photograph of habitus, dry. Scale bar = 2 mm.