Drapetis, Meigen, 1822
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publication ID |
https://doi.org/10.11646/zootaxa.4624.3.11 |
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publication LSID |
lsid:zoobank.org:pub:2F12876C-2DA7-4452-9200-65F4BE1EC243 |
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persistent identifier |
https://treatment.plazi.org/id/7B5E8D08-AF0E-FFF9-FF08-49FB48C94494 |
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treatment provided by |
Plazi (2019-07-01 05:24:53, last updated 2024-11-29 16:24:55) |
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scientific name |
Drapetis |
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Key to males of NW European species of Drapetis View in CoL View at ENA
1 Radial-medial crossvein (r–m) situated at or before the middle of the basal medial cell (bm) ( Fig. 2A View FIGURE 2 ) … ( D. assimilis View in CoL -group)................................................................................................... 2
– Radial-medial crossvein (r–m) situated beyond the middle of the basal medial cell (bm) ( Figs 2B, C View FIGURE 2 ) … ( D exilis View in CoL -group)... 7
2 Proepisternum shiny black. Pedicel beneath with 1–2 elongate setae. Knob of halter dusky brown..................... 3
– Proepisternum grey pruinose. Pedicel with short setae only. Knob of halter pale yellow............................. 6
3 Proepisternum bare except for two setae at base. Hind femur with a row of long pale posteroventral setae in basal two-thirds ( Fig. 3A View FIGURE 3 )........................................................................................... 4
– Proepisternum also with a pale up-turned seta arising from the middle. Hind femur with pale posteroventral setae confined to basal third or absent.................................................................................… 5
4 Face and clypeus shiny black. Legs extensively blackish brown, paler brown on foreleg. Male terminalia ( Collin 1961, fig. 12a)................................................................................ … assimilis (Fallén) View in CoL
– Face and clypeus grey pruinose. Legs extensively yellowish to yellow brown, only more or less darkened dorsally on femora. Male terminalia ( Kovalev 1972, figs 1–3)...................................................… ingrica Kovalev
5 Hind femur with posteroventral row of pale setae in basal third. Male terminalia ( Collin 1961, fig. 13a)... … arcuata Loew
– Hind femur without posteroventral setae. Male terminalia ( Collin 1961, fig. 12b).................... … simulans Collin
6 Scape and pedicel black. Fore tarsi yellowish with blackish apical tarsomere...................... … convergens Collin
– Scape and pedicel orange yellow. All tarsi of uniform brownish colour. Male terminalia ( Grootaert et al. 2010, figs 5–8).......................................................................................... flavipes Macquart View in CoL
7 Mid tibia simple....................................................................................… 8
– Mid tibia ventrally expanded and compressed, with numerous denticles on its ventral edge ( Figs 3C View FIGURE 3 , 4A, C View FIGURE 4 ). Male terminalia ( Figs 1 View FIGURE 1 A–D).......................................................................... abrollensis sp. nov.
8 Mid femur in front with transverse cuticular ribs ( Figs 5C, E View FIGURE 5 , 6 View FIGURE 6 A–D) or fine furrows ( Figs 5B, D View FIGURE 5 ) and with usual ground setulae very short or absent.................................................................................… 9
– Mid femur simple, in front with practically smooth cuticle and a uniform cover of setulae ( Fig. 5A View FIGURE 5 ).................. 13
9 Mid femur in front with a variable pattern of transverse cuticular ribs ( Figs 5C, E View FIGURE 5 , 6 View FIGURE 6 A–D).......................... 10
– Mid femur in front with numerous linear transverse furrows ( Figs 5B, D View FIGURE 5 ). Male terminalia ( Collin 1961, fig. 14a)................................................................................................ … exilis Meigen View in CoL
10 Knob of halter dark. Scape and pedicel brownish black................................................... … 11
– Knob of halter white. Scape reddish brown, pedicel ochre yellow. Male terminalia ( Grootaert 2016, figs 6–8)............................................................................................ … bruscellensis Grootaert View in CoL
11 Fore femur anteroventrally without a recurved apical seta. Mid femur with field of transverse ribs in anterior position ( Figs 5C View FIGURE 5 , 6A View FIGURE 6 ).............................................................................................… 12
– Fore femur anteroventrally with a recurved apical seta ( Fig. 3B View FIGURE 3 ). Mid femur with anterodorsal field of ca. 18 transverse ribs and valleys devoid of setulae ( Figs 6B, D View FIGURE 6 ). Male terminalia ( Grootaert 2016, figs. 12–14)................... … pusilla Loew View in CoL
12 Setulae beneath antennal pedicel reaching length of pedicel. Mid femur with anterior field of more than 20 transverse ribs crossed by a single row of short setulae ( Figs 6A, C View FIGURE 6 ). Male terminalia ( Grootaert 2016, figs 9–11)......... … parilis Collin View in CoL
– Setulae beneath antennal pedicel shorter than length of pedicel. Mid femur with anterior field of less than 15 short transverse ribs covered in short setulae ( Figs 5C, E View FIGURE 5 ). Male terminalia ( Collin 1961, fig. 14b).................... … infitialis Collin View in CoL
13 Longitudinal wing veins of regular thickness, membrane almost clear; vein R 4+5 fully developed until wing margin ( Fig. 2A View FIGURE 2 ) …................................................................................................ 14
– Longitudinal wing veins thickened, membrane conspicuously clouded; vein R 4+5 pale and faded shortly before wing margin ( Fig. 2C View FIGURE 2 ). Male terminalia ( Kovalev 1972, figs 8, 9)............................................ incompleta Collin View in CoL
14 Hind femur thinner than fore femur. Male terminalia with two strong, spine-like setae on left surstylus 1 and two similar setae on right surstylus ( Kovalev 1972, figs 15, 16, 24).........................................… stackelbergi Kovalev View in CoL
– Hind femur as thick as fore femur. Male terminalia without prominent, spine-like setae ( Kovalev 1972, figs 17–19, 24)........................................................................................ … completa Kovalev View in CoL
Collin, J. E. (1961) Empididae. In: British Flies. Vol. 6. Cambridge University Press, Cambridge, pp. i-viii + 1 - 782.
Grootaert, P., Shamshev, I. & Stark, A. (2010) Drapetis flavipes Macquart (Diptera, Hybotidae) new for the Belgian fauna, with a re-description of the species and a preliminary key to the West-European species of Drapetis. Bulletin de la Societe Royale Belge d'Entomologie, 146, 110 - 115.
Grootaert, P. (2016) Drapetis bruscellensis (Diptera, Hybotidae) a new species for science from the outskirts of Brussels, a not so cryptic species supported by COI barcoding. Belgian Journal of Entomology, 41, 1 - 14.
Kovalev, V. G. (1972) Diptera of the genera Drapetis Mg. and Crossopalpus Bigot (Diptera, Empididae) from the European part of the USSR. Entomologischeskoe Obozrenie, 51, 173 - 196. [in Russian]
FIGURE 1. Drapetis abrollensis sp. nov. Male terminalia. A. Right epandrial lamella with projections on apical border, right lateral view; B. Epandrium with cerci, dorsal view; C. Left surstylus, left lateral view; D. Epandrium and hypandrium, ventral view.Abreviations: ej = ejaculatory apodeme; hy = hypandrium; lc = left cercus; lel = left epandrial lamellae; ls 1 = left surstylus 1; ls 2 = left surstylus 2; p1, p2 = projections of the right epandrial lamella; rc = right cercus; rel = right epandrial lamella. Scale = 0.1 mm.
FIGURE 2. Drapetis males. Right wing. A. D. assimilis (Fallén); B. D. abrollensis sp. nov.; C. D. incompleta Collin. Scale = 0.1 mm.
FIGURE 3. Drapetis males. Legs. A. D. assimilis (Fallén), right hind femur; B. D. pusilla Loew, left fore femur; C. D. abrollensis sp. nov., right mid femur and tibia. Scale = 0.1 mm.
FIGURE 4. Drapetis abrollensis sp. nov. Right midleg, SEM images showing modified cuticle. A. Femur and tibia; B. Subapical cuticle of femur; C. Ventral edge of tibia; D. Detail of C showing accumulations of brochosomes. Scales: A = 0.1 mm; C = 1 μm.
FIGURE 5. Drapetis males. Left mid femur, SEM images showing modified cuticle. A. D. completa Kovalev; B. D. exilis Meigen; C. D. infitialis Collin; D. D. exilis, detail of anterior cuticle; E. D. infitialis, detail of anterior cuticle. Scales:A = 1 μm, C = 10 μm.
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