Plusiocampa Silvestri, 1912

Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Kovac, L'ubomír, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante & Reboleira, Ana Sofia P. S., 2020, Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae), European Journal of Taxonomy 591, pp. 1-138 : 13-20

Plusiocampa Silvestri, 1912

Plusiocampa Silvestri, 1912: 141 .

Diagnosis ( Silvestri 1912; Condé 1955 a; Sendra & Weber 2018)

Apparently smooth epicuticle, usually reticulated at high magnification, rosetta pores absent. Head with a frontal process with or without tuberculate setae. In non-troglomorphic species, cupuliform organ with four or five spheroidal olfactory chemoreceptors with polygonal net, with pore surface made by at least one cup-shaped fold and a central structure ending in a terminal pore. In troglobiomorphic species, spheroidal olfactory chemoreceptor with more folds in spiral, radial or other complex shape and a more visible polygonal net with pore surface. Sensillum of third antennomere in ventral position. Meso- and metathorax with a few macrosetae but frequently with medial anterior, lateral anterior, lateral posterior and medial posterior (exceptionally medial intermediate and lateral intermediate) macrosetae. Femur with one to five dorsal macrosetae; tibia with one to three ventral macrosetae. Elbow-like claws with usually large lateral crests and setiform lateral processes, rarely with a few barbs on proximal part. Abdomen with lateral anterior and posterior macrosetae and never with medial anterior macrosetae. Sternal macrosetae: sternite I with 6+6 up to 60 macrosetae; sternites II–VII with 4+4 to up to 14+14 macrosetae; sternite VIII with 2+2 up to 4+4 macrosetae. Secondary sexual differences in shape of first urosternite appendages and number of glandular setae.

Habitat and distribution

This is the most diverse genus of Plusiocampinae , with a clear preference for the subterranean ecosystems ( Condé 1955 a; Sendra & Weber 2018). It has 72 species (including the seven new ones described here) and 10 subspecies, distributed around the Mediterranean Basin ( Figs 210–211 ). The exceptions are three Chinese species considered representatives of Plusiocampa by Silvestri (1931 a), Condé (1993 a) and Sendra et al. (2012 a): P. sinensis Silvestri, 1931 , P. lipsae Condé, 1993 and P. kashiensis Chou & Tong,

1980. Nevertheless, these species must be removed from Plusiocampa due to the macrosetal formula on urosternites II to VIII.

Key to the subgenera and species of Plusiocampa

The genus Plusiocampa has been divided into four subgenera based on two characters: the presence of the femoral dorsal macrosetae ( Condé 1955 a; Paclt 1957; Sendra & Weber 2018) and the number of tibial ventral macrosetae ( Bareth & Condé 1984). Based on these synapomorphies and other characters, the following taxonomic key is proposed for the current five subgenera and 69 species.

1. Femoral macrosetae absent...............................................................................................................2

– Femoral macrosetae present..............................................................................................................7

2. Lateral posterior mesonotal macrosetae absent ................................................................................... ........................................................ ... Plusiocampa (Stygiocampa) christiani Condé & Bareth, 1996

– Lateral posterior mesonotal macrosetae present..............................................................................3

3. 2+2 lateral posterior mesonotal macrosetae.....................................................................................4

– 1+1 lateral posterior mesonotal macrosetae.....................................................................................6

4. Lateral expansions on posterior side of first urosternite absent in adult males; males without g 1 glandular setae; urosternal appendages in adult males enlarged, thick and subcylindrical ................. .............................................................................. Plusiocampa (Stygiocampa) nivea ( Joseph, 1882)

– Lateral expansions on posterior side of first urosternite present in adult males; males with g 1 glandular setae; urosternal appendages in males subcylindrical.........................................................5

5. 1+1 medial anterior mesonotal and metanotal macrosetae; 2+2 lateral anterior mesonotal macrosetae; 1+1 lateral anterior metanotal macrosetae ....................................................................... .......................................................................... Plusiocampa (Stygiocampa) dalmatica Condé, 1959

– Medial anterior mesonotal and metanotal macrosetae absent; 1+1 lateral anterior mesonotal macrosetae; lateral anterior metanotal macrosetae absent ................................................................... ................................................................................. Plusiocampa (Stygiocampa) remyi Condé, 1947

6. 2 ventral tibial macrosetae; unequal pretarsal claws (1.2–1.6); males with g 1 glandular setae; urosternal appendages in adult males subcylindrical ......................................................................... ................................................................................. Plusiocampa (Stygiocampa) denisi Condé, 1947

– Ventral tibial macrosetae absent; slightly unequal pretarsal claws (1.1–1.2); males without g 1 glandular setae; urosternal appendages in adult males enlarged and subcylindrical ........................... ........................................................................ Plusiocampa (Stygiocampa) bureschi Silvestri, 1931

7. 5 dorsal femoral macrosetae ............................................................................................................... .............................................. Plusiocampa (Pentachaetocampa) inopinata Sendra & Weber, 2018

– 1 or 2 dorsal femoral macrosetae.......................................................................................................8

8. 2 dorsal femoral macrosetae............................................................................................................9

– 1 dorsal femoral macroseta.............................................................................................................11

9. 1+1 medial posterior macrosetae on mesonotum and metanotum; pretarsal claws subequal (1.0– 1.1); 5+5 macrosetae on urosternites II–VII ......................................................................................... ......................................................... Plusiocampa (Dydimocampa) alhamae Condé & Sendra, 1989

– Medial posterior mesonotal and metanotal macrosetae absent; pretarsal claws unequal (1.2–1.6); 6+6 macrosetae on urosternites II–VII..........................................................................................10

10. 3 ventral tibial macrosetae; 2+2 lateral anterior mesonotal macrosetae; 4+4 posterior macrosetae on urotergites IV–V .......................................... Plusiocampa (Dydimocampa) euxina Condé, 1996

– 2 ventral tibial macrosetae; 1+1 lateral anterior mesonotal macrosetae; 2+2 posterior macrosetae on urotergites IV–V .............................. Plusiocampa (Dydimocampa) evallonychia Silvestri, 1949

11. Pretarsal claws very unequal (1.6–2.5)..........................................................................................12

– Pretarsal claws subequal, slightly unequal or unequal (1.0–1.6)..................................................15

12. Male urosternal appendages enlarged, thick and subcylindrical...................................................13

– Male urosternal appendages enlarged and subcylindrical, or subcylindrical...............................14

13. 2+2 lateral posterior metanotal macrosetae; 1+1 lateral anterior metanotal macrosetae; 6+6 macrosetae on urosternite I; posterior macrosetae on urotergite VI absent ..................................... ........................................................... Plusiocampa (Venetocampa) ferrani Sendra & Delić sp. nov.

– 1+1 lateral posterior metanotal macrosetae; lateral anterior metanotal macrosetae absent; 7+7 macrosetae on urosternite I; 2+2 posterior macrosetae on urotergite VI ........................................... ........................................................... Plusiocampa (Venetocampa) paolettii Bareth & Condé, 1984

14. Male urosternal appendages enlarged and subcylindrical .................................................................. ............................................... Plusiocampa (Plusiocampa) dublanskii Sendra & Turbanov sp. nov.

– Male urosternal appendages subcylindrical ........................................................................................ .................................................... Plusiocampa (Plusiocampa) ternovensis Sendra & Borko sp. nov.

15. Pretarsal claws subequal (1.0–1.1)................................................................................................16

– Pretarsal claws unequal or slightly unequal (1.1–1.6).................................................................35

16. 1+1 medial posterior mesonotal macrosetae..................................................................................17

– Medial posterior mesonotal macrosetae absent..............................................................................30

17. Lateral anterior metanotal macrosetae present...............................................................................18

– Lateral anterior metanotal macrosetae absent................................................................................26

18. 2+2 lateral anterior metanotal macrosetae......................................................................................19

– 1+1 lateral anterior metanotal macrosetae......................................................................................21

19. Pretarsal lateral crests small; posterior claw without backward overhang; 2+2 lateral anterior macrosetae on urotergite IV; shape of body and appendages non-troglomorphic ............................... ....................................................... Plusiocampa (Plusiocampa) suspiciosa Condé & Mathieu, 1957

– Pretarsal lateral crests large; posterior claw with backward overhang; 1+1 lateral anterior macrosetae on urotergite IV; troglomorphic body and appendages.................................................20

20. 5+5 posterior macrosetae on urotergites IV–VII ................................................................................ ...................................................................... Plusiocampa (Plusiocampa) provincialis Condé, 1949

– 2+2 posterior macrosetae on urotergites IV–V; 4+4 posterior macrosetae on urotergites VI–VII ... ................................................................................ Plusiocampa (Plusiocampa) socia Condé, 1983

21. Pretarsal lateral crests small ......................... Plusiocampa (Plusiocampa) notabilis Silvestri, 1912

– Pretarsal lateral crests large.............................................................................................................22

22. 8+8 macrosetae on urosternite I.....................................................................................................23

– 7+7 macrosetae on urosternite I.....................................................................................................24

23. Posterior pretarsal claw without backward overhang; 2+2 posterior macrosetae on urotergite III; 7+7 posterior macrosetae on urotergite VIII; 1+1 lateral anterior macrosetae on urotergite IV .. ............................................................................. Plusiocampa (Plusiocampa) lagoi Silvestri, 1932

– Posterior pretarsal claw with backward overhang; 1+1 posterior macrosetae on urotergite III; 6+6 posterior macrosetae on urotergite VIII; 2+2 lateral anterior macrosetae on urotergite IV .. ............................................................................. Plusiocampa (Plusiocampa) breuili Condé, 1955

24. 3+3 lateral anterior mesonotal macrosetae.....................................................................................25

– 2+2 lateral anterior mesonotal macrosetae ......................................................................................... ............................................................... Plusiocampa (Plusiocampa) lagari Sendra & Condé, 1987

25. 5+5 posterior macrosetae on urotergite IV ..... Plusiocampa (Plusiocampa) bonadonai Condé, 1948

– 3+3 posterior macrosetae on urotergite IV ........................................................................................... ..................................................... Plusiocampa (Plusiocampa) tinoamorei Sendra & Nicolosi, 2019

26. Lateral anterior mesonotal macrosetae present..............................................................................27

– Lateral anterior mesonotal macrosetae absent ... Plusiocampa (Plusiocampa) glabra Condé, 1984

27. 3+3 lateral anterior mesonotal macrosetae....................................................................................28

– 1+1 or 2+2 lateral anterior mesonotal macrosetae.........................................................................29

28. 6+6 posterior macrosetae on urotergites V–VII ................................................................................... ........................................... Plusiocampa (Plusiocampa) apollo Sendra, Giachino & Vailati sp. nov.

– 5+5 posterior macrosetae on urotergites V–VII ... Plusiocampa (Plusiocampa) festae Silvestri, 1932

29. 2+2 lateral anterior mesonotal macrosetae ....... Plusiocampa (Plusiocampa) rudnica Blesić, 1992

– 1+1 lateral anterior mesonotal macrosetae ......................................................................................... .................................................. Plusiocampa (Plusiocampa) chiosensis Sendra & Gasparo sp. nov.

30. 2 ventral tibial macrosetae; 1+1 lateral anterior macrosetae on urotergites IV–V.....................31

– 1 ventral tibial macroseta; lateral anterior macrosetae on urotergites IV–V absent ....................... ................................................... Plusiocampa (Plusiocampa) dolichopoda Bareth & Condé, 1984

31. Pretarsal lateral crests small...........................................................................................................32

– Pretarsal lateral crests large.............................................................................................................34

32. 3+3 lateral anterior mesonotal macrosetae; 1+1 lateral anterior metanotal macrosetae..............33

– 2+2 lateral anterior mesonotal macrosetae; lateral anterior metanotal macrosetae absent .............. ......................................................................... Plusiocampa (Plusiocampa) cyrnensis Condé, 1953

33. 5+5 posterior macrosetae on urotergites IV–VII ............................................................................... .......................................................................... Plusiocampa (Plusiocampa) exsulans Condé, 1947

– 3+3 posterior macrosetae on urotergite IV; 4+4 posterior macrosetae on urotergites V–VII ......... ..................................................................... Plusiocampa (Plusiocampa) corcyraea Silvestri, 1912

34. Male urosternal appendages enlarged and subcylindrical; 2+2 lateral anterior mesonotal macrosetae; lateral anterior metanotal macrosetae absent; posterior claw without backward overhang ... .......................................................................... Plusiocampa (Plusiocampa) lindbergi Condé, 1956

– Male urosternal appendages subcylindrical; 3+3 lateral anterior mesonotal macrosetae; 1+1 lateral anterior metanotal macrosetae; posterior claw with backward overhang ........................................ ................................................ Plusiocampa (Plusiocampa) arbanasiensis Bareth & Condé, 2001

35. Pretarsal claws unequal (1.2–1.6)..................................................................................................36

– Pretarsal claws slightly unequal (1.1–1.2).....................................................................................67

36. 1+1 medial posterior mesonotal macrosetae...................................................................................37

– Medial posterior mesonotal macrosetae absent.............................................................................47

37. Lateral anterior metanotal macrosetae present.............................................................................38

– Lateral anterior metanotal macrosetae absent ................................................................................... ...................................................................... Plusiocampa (Plusiocampa) hoelzeli ( Neuherz, 1984)

38. 2+2 lateral anterior metanotal macrosetae.....................................................................................39

– 1+1 lateral anterior metanotal macrosetae.....................................................................................41

39. 8+8 macrosetae on urosternite I; urosternal appendages in females enlarged and subcylindrical; 4–5 olfactory chemoreceptors; setae on frontal process slightly tuberculated .................................. ........................................................................ Plusiocampa (Plusiocampa) elongata Ionescu, 1955

– 7+7 macrosetae on urosternite I; urosternal appendages in females subcylindrical; more than 5 olfactory chemoreceptors; setae on frontal process tuberculated...................................................40

40. 5+5 posterior macrosetae on urotergites IV–VII ................................................................................. ....................................................................... Plusiocampa (Plusiocampa) bulgarica Silvestri, 1931

– 4+4 posterior macrosetae on urotergites IV–VII ................................................................................. ......................................................... Plusiocampa (Plusiocampa) vodniensis Bareth & Condé, 2001

41. 3 ventral tibial macrosetae............................................................................................................42

– 2 ventral tibial macrosetae............................................................................................................45

42. 3+3 lateral anterior mesonotal macrosetae.....................................................................................43

– 2+2 lateral anterior mesonotal macrosetae.....................................................................................44

43. Male urosternal appendages enlarged and subcylindrical; 2+2 posterior macrosetae on urotergite III; slightly troglomorphic body and appendages .............................................................................. .............................................................................. Plusiocampa (Plusiocampa) caprai Condé, 1950

– Male urosternal appendages subcylindrical; 1+1 posterior macrosetae on urotergite III; troglomorphic body and appendages ...................... Plusiocampa (Plusiocampa) strouhali Silvestri, 1933

44. 6+6 macrosetae on urosternites II–VII; 1+1 posterior macrosetae on urotergite III; 2+2 lateral anterior macrosetae on urotergites V–VI ....... Plusiocampa (Plusiocampa) grandii Silvestri, 1933

– 5+5 macrosetae on urosternites II–VII; 2+2 posterior macrosetae on urotergite III; 1+1 lateral anterior macrosetae on urotergites V–VI ............ Plusiocampa (Plusiocampa) fagei Condé, 1955

45. Subcylindrical male urosternal appendages enlarged; 3+3 lateral anterior mesonotal macrosetae; slightly troglomorphic body and appendages................................................................................46

– Subcylindrical male urosternal appendages not enlarged; 2+2 lateral anterior mesonotal macrosetae; troglomorphic body and appendages .......... Plusiocampa (Plusiocampa) fagei Condé, 1955

46. Olfactory chemoreceptors simple; more than 5 olfactory chemoreceptors ..................................... ........................................................ Plusiocampa (Plusiocampa) friulensis Bareth & Condé, 1984

– Olfactory chemoreceptor complex; 4–5 olfactory chemoreceptors ................................................. ............................................................ Plusiocampa (Plusiocampa) beroni Bareth & Condé, 2001

47. Ventral tibial macrosetae present....................................................................................................48

– Ventral tibial macrosetae absent ............... Plusiocampa (Plusiocampa) dargilani ( Moniez, 1894)

48. 3 ventral tibial macrosetae.............................................................................................................49

– 1 or 2 ventral tibial macrosetae.....................................................................................................51

49. 1+1 medial intermediate mesonotal and metanotal macrosetae; 1+1 lateral intermediate mesonotal and metanotal macrosetae; 4+4 posterior macrosetae on urotergite VI .................................... ............................................................. Plusiocampa (Plusiocampa) lucenti Sendra & Condé, 1986

– Medial and lateral intermediate macrosetae absent on mesonotum and metanotum; 5+5 posterior macrosetae on urotergite VI............................................................................................................50

50. 2+2 lateral anterior macrosetae on urotergites V–VII; olfactory chemoreceptors laminar ................ ................................................................ Plusiocampa (Plusiocampa) dallai Bareth & Condé, 1984

– 1+1 lateral anterior macrosetae on urotergites V–VII; olfactory chemoreceptors simple ............... ....................................................................... Plusiocampa (Plusiocampa) djurdjurae Condé, 1984

51. 2 ventral tibial macrosetae.............................................................................................................52

– 1 ventral tibial macroseta...............................................................................................................66

52. 3+3 lateral posterior metanotal macrosetae ........................................................................................ .................................................................... Plusiocampa (Plusiocampa) pouadensis ( Denis, 1930)

– 1+1 or 2+2 lateral posterior metanotal macrosetae.......................................................................53

53. 2+2 lateral posterior metanotal macrosetae...................................................................................54

– 1+1 lateral posterior metanotal macrosetae....................................................................................65

54. Urosternal appendages in adult males subtrapezoidal or subtriangular.....................................55

– Urosternal appendages in adult males subcylindrical, enlarged and subcylindrical, or enlarged, thick and subcylindrical..................................................................................................................56

55. 3+3 lateral anterior mesonotal macrosetae; 5+5 macrosetae on urosternites II–VII; 2+2 posterior macrosetae on urotergite III; 5+5 posterior macrosetae on urotergite IV ............................................. .............................................................................. Plusiocampa (Plusiocampa) isterina Condé, 1993

– 2+2 lateral anterior mesonotal macrosetae; 6+6 macrosetae on urosternites II–VII; 1+1 posterior macrosetae on urotergite III; 2+2 posterior macrosetae on urotergite IV ............................................. .............................................................................. Plusiocampa (Plusiocampa) balsani Condé, 1947

56. Urosternal appendages in adult males enlarged, thick and subcylindrical ........................................ ............................................................................ Plusiocampa (Plusiocampa) isterina Condé, 1993

– Urosternal appendages in adult males subcylindrical, or enlarged and subcylindrical.................57

57. Urosternal appendages in adult males enlarged and subcylindrical...............................................58

– Urosternal appendages in adult males subcylindrical.....................................................................63

58. 1+1 lateral intermediate mesonotal and metanotal macrosetae; 1+1 medial intermediate mesonotal and metanotal macrosetae; 1+1 lateral anterior macrosetae on urotergite III ......................... ............................................................. Plusiocampa (Plusiocampa) lucenti Sendra & Condé, 1986

– Medial and lateral intermediate macrosetae absent on mesonotum and metanotum; lateral anterior macrosetae absent on urotergite III................................................................................................59

59. Posterior macrosetae present on urotergite III.............................................................................60

– Posterior macrosetae absent on urotergite III .................................................................................... ..................................................... Plusiocampa (Plusiocampa) gueorguievi Bareth & Condé, 2001

60. 2+2 posterior macrosetae on urotergite III....................................................................................61

– 1+1 posterior macrosetae on urotergite III....................................................................................62

61. 5+5 posterior macrosetae on urotergite IV; pretarsal lateral processes long; non-troglomorphic body and appendages; olfactory chemoreceptors simple .................................................................... ............................................................................. Plusiocampa (Plusiocampa) isterina Condé, 1993

– 2+2 posterior macrosetae on urotergite IV; pretarsal lateral processes short; troglomorphic body and appendages; olfactory chemoreceptors complex .......................................................................... ................................................................................ Plusiocampa (Plusiocampa) dobati Condé, 1975

62. Urosternal appendages in adult females enlarged and subcylindrical; olfactory chemoreceptors complex ......................................................... Plusiocampa (Plusiocampa) schweitzeri Condé, 1947

– Urosternal appendages in adult females subcylindrical; olfactory chemoreceptors simple ............ ....................................................................... Plusiocampa (Plusiocampa) djurdjurae Condé, 1984

63. 5+5 posterior macrosetae on urotergite IV ...... Plusiocampa (Plusiocampa) isterina Condé, 1993

– 2+2 or 3+3 posterior macrosetae on urotergite IV..........................................................................64

64. 3+3 posterior macrosetae on urotergite IV ....... Plusiocampa (Plusiocampa) latens Condé, 1948

– 2+2 posterior macrosetae on urotergite IV ...... Plusiocampa (Plusiocampa) affinis Condé, 1947

65. 1+1 medial anterior metanotal macrosetae; 3+3 posterior macrosetae on urotergites VI–VII; 6+6 posterior macrosetae on urotergite VIII ........ Plusiocampa (Plusiocampa) vedovinii Condé, 1981

– Medial anterior metanotal macrosetae absent; 4+4 posterior macrosetae on urotergites VI–VII; 5+5 posterior macrosetae on urotergite VIII .... Plusiocampa (Plusiocampa) hoelzeli ( Neuherz, 1984)

66. 2+2 lateral posterior metanotal macrosetae; urosternal appendages in adult males subtrapezoidal or subtriangular; 1+1 medial anterior metanotal macrosetae; a 2 glandular setae present in adult males ................................................................... Plusiocampa (Plusiocampa) balsani Condé, 1947

– 1+1 lateral posterior metanotal macrosetae; urosternal appendages in adult males enlarged and subcylindrical; medial anterior metanotal macrosetae absent; a 2 glandular setae absent in adult males ............................................................ Plusiocampa (Plusiocampa) hoelzeli ( Neuherz, 1984)

67. 1+1 medial posterior mesonotal and metanotal macrosetae.........................................................68

– Medial posterior mesonotal and metanotal macrosetae absent......................................................79

68. Lateral anterior metanotal macrosetae present...............................................................................69

– Lateral anterior metanotal macrosetae absent ... Plusiocampa (Plusiocampa) rhea Sendra sp. nov.

69. 2+2 lateral anterior metanotal macrosetae....................................................................................70

– 1+1 lateral anterior metanotal macrosetae....................................................................................72

70. 4+4 posterior macrosetae on urotergite III ...... Plusiocampa (Plusiocampa) romana Condé, 1954

– 2+2 or 3+3 posterior macrosetae on urotergite III........................................................................71

71. 3+3 posterior macrosetae on urotergite III .......................................................................................... ..................................................... ... Plusiocampa (Plusiocampa) hystricula Bareth & Condé, 1984

– 2+2 posterior macrosetae on urotergite III .... Plusiocampa (Plusiocampa) sardiniana Condé, 1981

72. 4 ventral tibial macrosetae ........................ Plusiocampa (Plusiocampa) magdalenae Condé, 1957

– 2 or 3 ventral tibial macrosetae.....................................................................................................73

73. 3 ventral tibial macrosetae............................................................................................................74

– 2 ventral tibial macrosetae............................................................................................................76

74. 7+7 posterior macrosetae on urotergite VIII; 1+1 lateral anterior macrosetae on urotergite III ... ....................................... Plusiocampa (Plusiocampa) hoffmanni Sendra & Paragamanm sp. nov.

– 6+6 posterior macrosetae on urotergite VIII; lateral anterior macrosetae on urotergite III absent................................................................................................................................................75

75. 5+5 posterior macrosetae on urotergite IV; 2+2 lateral anterior macrosetae on urotergite IV; 3+3 lateral anterior macrosetae on urotergites V–VI ................................................................................. ..................................................................... Plusiocampa (Plusiocampa) magdalenae Condé, 1957

– 3+3 posterior macrosetae on urotergite IV; 1+1 lateral anterior macrosetae on urotergites IV–VI ................................................... Plusiocampa (Plusiocampa) tinoamorei Sendra & Nicolosi, 2019

76. Posterior pretarsal claw without backward overhang ....................................................................... ...................................................................... Plusiocampa (Plusiocampa) humicola Ionescu, 1951

– Posterior pretarsal claw with backward overhang........................................................................77

77. 2+2 posterior macrosetae on urotergite III; non-troglomorphic body and appendages; 4–5 simple olfactory chemoreceptors .................................. Plusiocampa (Plusiocampa) spelaea Stach, 1930

– 1+1 posterior macrosetae on urotergite III; troglomorphic body and appendages; more than 5 complex olfactory chemoreceptors.................................................................................................78

78. Urosternal appendages in adult males enlarged, thick and subcylindrical; 7+7 posterior macrosetae on urotergite VIII; 1+1 lateral anterior macrosetae on urotergite III; setae on frontal process tuberculated ................... Plusiocampa (Plusiocampa) hoffmanni Sendra & Paragamian sp. nov.

– Urosternal appendages in adult males subcylindrical; 6+6 posterior macrosetae on urotergite VIII; lateral anterior macrosetae on urotergite III absent; setae on frontal process slightly tuberculated ................................................... Plusiocampa (Plusiocampa) tinoamorei Sendra & Nicolosi, 2019

79. Lateral anterior mesonotal macrosetae present...............................................................................80

– Lateral anterior mesonotal macrosetae absent ... Plusiocampa (Plusiocampa) baetica Sendra, 2004

80. 3+3 lateral anterior mesonotal macrosetae ..... Plusiocampa (Plusiocampa) bonneti Condé, 1948

– 2+2 lateral anterior mesonotal macrosetae.....................................................................................81

81. 2+2 lateral anterior metanotal macrosetae; 5+5 posterior macrosetae on urotergites VI–VII; posterior macrosetae on urotergite I absent .... Plusiocampa (Plusiocampa) gadorensis Sendra, 2001

– 1+1 lateral anterior metanotal macrosetae; 4+4 posterior macrosetae on urotergites VI–VII; 1+1 posterior macrosetae on urotergite I .... Plusiocampa (Plusiocampa) solerii Silvestri, 1932

Figures

Fig. 210. Distribution map of Plusiocampinae Paclt, 1957 and tachycampoid phyletic group genera in the Euro-Mediterranean area: Cestocampa Condé, 1956 (green squares), Hystrichocampa Condé, 1948 (grey squares), Jeannelicampa Condé, 1952 (brown squares), Oncinocampa Condé, 1982 (red squares), Paratachycampa Wygodzinsky, 1944 (black squares), Patrizicampa Condé, 1956 (yellow squares), Plusiocampa Silvestri, 1912 (navy blue squares), Tachycampa Silvestri, 1936 (violet squares) and Vandelicampa Condé, 1955 (light blue squares).

Fig. 211. Distribution map of subgenera of Plusiocampa Silvestri, 1912: Dydimocampa Paclt, 1957 (blue squares), Pentachaetocampa Sendra & Weber, 2018 (black square), Plusiocampa s. str. (red squares), Stygiocampa Silvestri, 1934 (green squares) and Venetocampa Bareth & Condé, 1984 (yellow squares).