Chelipoda tainuia, Published, 2007

Published, First, 2007, The Hemerodromiinae (Diptera: Empididae) of New Zealand II. Chelipoda Macquart, Zootaxa 1537 (1), pp. 1-88 : 15-16

4. Chelipoda tainuia sp. n.

[ Figs. 60, 61 , 73 , 111 ]

Type material. Holotype ♂: New Zealand, North Is. , CL, Little Barrier Is., swept in forest on summit track, 500–650m, 11/xi/1985, ARP [ NZAC] . Paratypes: North Is : 1♂, WO, Pirongia, 22/ix/1985, ARP [ NZAC] ; 1♂, 2♀, TO, Pureora Forest , 29/xii/1986, ARP , [ NZAC] ; 2♀, AK, Titirangi, ex light trap, 8/ii/1953 and 7/vii/1953, CRT [ NZAC] ; 5♀ same data as holotype [ NMWC] ; 2♂, WO, Kawhia, Hauturu Road North, 13/ix/1985, ARP [ NMWC] ; 1♂, BP, Kaimai range, Wairere Falls track, 350m, 22/iii/1988, ARP [ NMWC] .

Etymology: The specific epithet refers to the Tainui people, the Maori tribe whose territory is embraced by the range of this species.

Description. Male: length 2.0– 2.5mm (wet specimens).

Head: blackish brown, dusted greyish; setae black, only lpo pale; eyes touching on upper face, widening towards mouth edge; ocl strong, diverging, as strong as verticals which are arranged as inner (vt1) and outer (vt2) pairs; line of upo congruent with vt1; lpo approximately biserial, hair-like. Palpi very small, rather elongate, yellowish with 1 or 2 pale apical setae. Antenna ( Fig. 111 ) yellowish to greyish (basal segments sometimes appearing paler), rather long, about 2X as long as head is deep; postpedicel conical, apically pointed, 2.5–3X as long as wide, arista 3X as long.

Thorax: ground colour orange-yellow often slightly darker on mesonotum medially and laterally, on metanotum and on katepisternum anteriorly; dusted greyish, more strongly so on pleura, without conspicuous dusting stripes and not markedly obscuring ground colour. Chaetotaxy similar to C. modica ; all setae black, dc3 about 0.6–0.7X as long as dc2; pprn, ph and lnp of equally small, hair-like; prothoracic episternum with a few minute hairs at side; Anatergite with one strong and 2 weaker setae.

Legs: yellow, ‘knees’ and apical tarsal segments dusky; C 1 0.75X as long as thorax, 5.2–5.4X as long as wide with minute pile and minute pale hairs, a fine bristle at side apically distinct from anteroapical fan of hairs; C 2 and C 3 with short pale bristles anteroapically; F 1 more slender than in C. modica , slightly longer than C 1, 3.6X as long as wide, a row of fine hairs dorsally more conspicuous than other hairs; femoral formula 4(4– 4)/19(18–20)/10(9–11)/4(4–5) with av and pv spines yellowish and basal 3–4 denticles in pv series black and point-like, abruptly becoming yellowish and bristle-like distally; 2–4 basal bristles contiguous basally with lines of av and pv spines. F 2 simple, short haired, F 3 and T 3 with dorsal bristles rather stronger and more erect than other hairs; at2 0.8X at3; pt1 slightly longer than pt2 + pt3 + pt4.

Abdomen: yellowish dorsally, paler ventrally, tergites with posterior marginal hairs strongest on tergite 5; vent shorter haired, longer on apical sternites. Hypopygium ( Fig. 60 ) yellowish, reflexed anteriorly; epandrium divided, the two lobes clearly separated from large rather keel shaped hypandrium; epandrial lobes bluntly pointed, a row of short flattened black setae distally and more normal setae proximally on posterior margin; cerci small, forming a blunt inwardly directed prominence at their tip where fused with epandrial lobes; postgonite (?) bearing a thick mat of short blunt rod-like setae. Phallus narrow, gently curved; phallic sheath apically with a pair of small blunt ended lateral processes and a longer pointed ventral process ( Fig. 61 ).

Wing: extending beyond abdomen, veins brownish yellow, membrane faintly tinged yellow; cell cup complete, rounded apically; vein A 1 strong to end of cell cup, continued weakly for a short distance beyond end of cell. Halteres whitish yellow.

Description. Female: length 2.5–3.0mm.

Very similar to male. F 1 with more numerous bristles, femoral formula 4(4–5)/22.5(18–24)/11(8–13)/4(4– 5). Pregenital sternite apically narrowed ( Fig. 73 ).

Comments. An orange-yellow species of the C. modica -group, similar to C. atrocitas but with epandrial lobes bluntly pointed with distinctive chaetotaxy and female pregenital sternite apically narrowed. It is currently known only from large relatively intact forests of the northern half of the North Island at altitudes between sea level and about 600 m. Capture dates were from November to March inclusive and in July.

Figures

FIGURES 56–62. Male genitalia of Chelipoda spp. 56. C. rakiuraensis sp. n. posterior view. 57–59. C. secreta Collin. 57. anterior view. 58. posterior view. 59. apex of postgonite. 60–61. C. tainuia sp. n. 60. lateral view. 61. apex of phallic structure. 62. C. trepida Collin, posterior view.

FIGURES 71–75. Female pregenital sternite of 71. C. atrocitas sp. n. 72. C. macrostigma sp. n. 73. C. tainuia sp. n. 74. C. modica Collin. 75. C. mediolinea sp. n. 76. head of ♂ C. aritarita sp. n. 77. C. ultraferox sp. n. ♂ scutum, lateral view. 78. C. consignata Collin, ♂ head. 79–80. thoracic profile. 79. C. rangopango sp. n. 80. C. cornigera sp. n. 81. C. gracilis sp. n. ♂ head & front of scutum, dorsal view. Abbreviations: see text.

FIGURES 104–116. Antennae of Chelipoda spp. 104–105. C. monorhabdos sp. n. 104. ♀. 105. ♂. 106–107. C. puhihiroa sp. n. 106. ♀. 107. ♂. 108. C. secreta Collin ♀. 109. C. rakiuraensis sp. n. ♂. 110. C. secreta Collin, ♂. 111. C. tainuia sp. n. ♂. 112. C. trepida Collin ♂. 113. C. venatrix sp. n. ♂. 114. C. tangerina sp. n. ♂. 115–116. C. didhami sp. n. 115. ♀. 116. ♂.