Poecilosomella borboroides ( WALKER, 1860 )
publication ID |
https://doi.org/ 10.5281/zenodo.12587600 |
persistent identifier |
https://treatment.plazi.org/id/5B5787B6-5F77-FFEB-407E-7EFAFB12312F |
treatment provided by |
Felipe |
scientific name |
Poecilosomella borboroides ( WALKER, 1860 ) |
status |
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Poecilosomella borboroides ( WALKER, 1860) View in CoL
( Figs 1–5 View Figs 1–5 )
Ephydra borboroides : WALKER 1860: 171. Holotype female (BMNH): Celebes, Makessar [now Sulawesi, Ujungpandang].
The original description says: “Length of the body 1 1/2 line; of the wings 3 lines” [that would be 7.62 mm], which must be an overstatement.
Mr NIGEL WYATT (The Natural History Museum London) examined the only type specimen and he wrote: “I have examined the holotype of Poecilosomella borboroides WALKER and it seems to belong to your ‘species 2’ [my term to differentiate the two forms I saw]. There are no ventral or anteroventral setae on the mid tibia except for the ventral seta at the apex.” This piece of information is critically important in arbitration of the closely related P. ornata , which was formerly erroneously placed into synonymy with P. borboroides (see below).
Material studied: 3 males, 7 females: MALAYSIA: Sabah, G. F. Hevel & W. E. Steiner 1983 ( NMNH, 1 male, 1 female in HNHM): 1 km S Kundasang, el. 1530 m, 6 Aug; Kinabalu National Park , Poring , el. 570, 8 Sept., ibid., Summit trail below Carson Camp , 1980–2700 m, 16 Sept., Telipok , 12 August. 1 male ( ROM): INDONESIA, Sumatra, W. Sumatra: Kerinci-Seblat Nat. Pk., trail to L. Tujuh, 14 SEP 1989. B Hubley, DC Darling, ROM 893074 – 1 About ROM ° rainforest, 1640 m, screen sweep, 1°42’N, 101°19’E. 1 male, 2 females ( HNHM): India, leg. Topál. GoogleMaps
Male abdominal sternite 5 strongly asymmetrical, sternite 6 contiguous with 2 asymmetrical median lobes and a broad lobe on the left side ( Fig. 1 View Figs 1–5 ). Subepandrial sclerite ( Fig. 2 View Figs 1–5 ) with a broad paired, apically rounded process (characteristic for this species group), whose hairlike setae are longer than in the species from the Philippines. Surstylus ( Figs 4–5 View Figs 1–5 ) apically with 6 thick black teeth (prensisetae). Postgonite ( Fig. 3 View Figs 1–5 ) less broad medially than in the Philippines form, apex more demarcated. There are slight variations in the fine details of the surstylus among specimens from different localities ( Fig. 4 View Figs 1–5 , vs Fig. 5 View Figs 1–5 ) but I think this normal.
2–5
It is useless to say that any specimens identified as P. borboroides before (incl. by myself before 2000) are doubtful. I hope colleagues will not regard my decision to select this biological species for the name P. borboroides (WALKER) just as an arbitrary decision. The form fixed by Figs 1–5 View Figs 1–5 seems to be the widespread one, and I also found specimens collected rather close to its type locality (Sulawesi (Celebes) vs. Borneo, Sabah). A dissection of the female holotype of P. borboroides (being a female) would not get closer to a better solution.
mm for Figs 6–8 View Figs 6–9
In the last phase of work on the Oriental Poecilosomella , I managed to study the type of P. picturata ( MALLOCH, 1913), described from the Philippines and formerly regarded as a junior synonym of P. borboroides . Based on a study of its genitalia, I would like to remove it from synonymy (see also below)
Poecilosomella ornata (DE MEIJERE, 1908) View in CoL , sp. restit.
( Figs 10–13 View Figs 10–15 )
Limosina ornata View in CoL : DE MEIJERE 1908: 177.
Originally two syntypes were known from Java, Semarang. It was proposed as synonymous erroneously on two different occasions for two different species (DE MEIJERE 1918, HACKMAN 1977)
Dr. HERMAN DE JONG (Amsterdam) sent me the following information: “ DE MEIJERE (1918; Tijdschrift voor Entomologie 60: 323) regarded his ornata as a variety of venalicia OSTEN SACKEN. According to DE MEIJERE, both should differ in the extent of the dark coloration of the wing membrane.
The single remaining type specimen of ornata was placed under ‘ venalicia ’ in our collection. It has the legs folded and closely appressed to the body. Nevertheless it is clear that the right t2 carries a strong (antero) ventral bristle near midlength. I could not find a ventro-apical bristle on either t2, although the view of both legs is quite obscured.”
Material studied: 2 males ( HNHM) : TAIWAN: Pingtung Hsien, Kenting, Heng-Chun Trop. Botanical Garden – forest undergrowth, Oct 5, 2000, leg. L. PAPP, No. 19. 2 males, 1 female ( ROM, 1 male in HNHM) : INDONESIA: Sumatra, Aceh, Gunung Leuser Nat. Pk., Ketambe Res. Sta., 9–21 SEP 1989, DC Darling, ROM 893087 – 1 About ROM ° rainforest. Young forest , Terrace 3, closed canopy, 350 m, 3°41’N, 97°39’E, Malaise traphead; 1 female ( ROM) GoogleMaps : ibid., 3–10 SEP, B Hubley, DC Darling, ROM 893027 About ROM ; 1 female ( ROM) GoogleMaps : ibid., 7 SEP, ROM 893059 – 1 About ROM ° rainforest, 350 m, 3°41’N, 97°39’E. Screen sweep; 1 female GoogleMaps : INDONESIA: E. Kalimantan, Kac. Pujungan, Kayan-Mentarang Nat. Reserve, Jun-Aug. 1993, DC Darling, Rosichon U. IIS 930596 – lowland diptero.[Dipterocarpus] forest, WWF Station, Lalut Birai, vicinity of Base Camp. Log emergence traps, 2°52’N, 115°49’E, 378 m. GoogleMaps
Male sternite 5 ( Fig. 10 View Figs 10–15 ) less asymmetrical than in its relatives, there are 3 wide (long) rows of comparatively long hairs on the sternite 5 & 6 complex medially ( Fig. 10 View Figs 10–15 ), setae on sternite 5 long and comparatively thick. Ventral process of subepandrial sclerite ( Fig. 11 View Figs 10–15 ) long, comparatively thin, finger-like. Sursylus ( Fig. 12 View Figs 10–15 ) with only 1 thick black tooth but there is a dentate process behind it. Postgonite ( Fig. 13 View Figs 10–15 ) rather curved with definitely demarcated dentiform apex. See more in the key below.
postgonite, lateral view. Scales: 0.2 mm for Figs 10–11 View Figs 10–15 , 0.1 mm for Figs 12–15 View Figs 10–15
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Genus |
Poecilosomella borboroides ( WALKER, 1860 )
Papp, L. 2002 |
Limosina ornata
MEIJERE, DE 1908: 177 |
Ephydra borboroides
WALKER, F. 1860: 171 |