Coniopterygidae, Burmeister, 1839

Sarmiento-Cordero, Mariza Araceli & Contreras-Ramos, Atilano, 2019, Two new species and a key to the dustywings of Mexico (Neuroptera, Coniopterygidae), Zootaxa 4671 (1), pp. 551-563 : 558-561

publication ID

https://doi.org/10.11646/zootaxa.4671.4.6

publication LSID

lsid:zoobank.org:pub:E4FC17A7-79B2-42BB-BB30-3B367B8A8BA3

persistent identifier

https://treatment.plazi.org/id/4D02878E-FFDF-FFA9-C4D0-399AFA4CFEA8

treatment provided by

Plazi

scientific name

Coniopterygidae
status

 

Key to adult males of Coniopterygidae from Mexico (modified from Sziráki 2011).

1. Abdomen with plicaturae, only lateral plicaturae present on the abdominal sternites ( Fig. 1B View FIGURE 1 ; Meinander 1972, fig. 16C; Johnson 1977, fig. 4)… Aleuropteryginae ...................................................................... 2

1’. Abdomen without plicaturae… Coniopteryginae ............................................................ 11

2. Hindwing with Radial cross-vein bit R 2+3 ( Meinander 1972, figs. 88E, 89G; Johnson 1977, fig. 3); wings in some cases shortened or absent… Fontenelleini ........................................................................... 3

............................................................................ Bidesmida morrisoni Johnson

3’. Sternite 7 of males without plicaturae ( Fig. 1B View FIGURE 1 ; Meinander 1972, fig. 88A); M thickened at the base of seta in the forewing ( Meinander 1972, fig. 88E); sternite 8 of males unsclerotized; parameres not fused with ectoprocts, but fused with the styli ( Figs. 1L, 1M View FIGURE 1 ; Meinander 1972, fig. 86A)… Neoconis ........................................................ 4

4. Forewing membrane hyaline along margin ( Meinander 1972, fig. 88E); ninth abdominal segment with prominent spines ( Meinander 1972, fig. 88B)........................................................ Neoconis inexpectata Meinander

4’. Forewing membrane with dark spots between longitudinal veins along distal edge of the wing ( Fig. 1B View FIGURE 1 ; Meinander 1972, fig. 89G; Monserrat 1985, fig. 1); ninth abdominal segment with one or three spines ( Figs. 1J, I, K View FIGURE 1 ; Monserrat 1985, fig. 2).... 5

5. Ninth abdominal segment with short apophyses with prominent spines ( Monserrat 1985, fig. 2)... Neoconis unam Monserrat

5’. Ninth abdominal segment without apophyses ( Fig. 1K View FIGURE 1 ; Meinander 1972, fig. 89B)................................. 6

6. Styli a dorsal long bar and a shorter ventral one, medially connected by a narrow bridge ( Meinander 1972, fig. 89F); an apparent narrow bridge connects dorsal rods of styli above penis ( Meinander 1972, fig. 89E)........ Neoconis marginata Meinander

6’. Styli undivided, with a small median protuberance, apically hooked in lateral view ( Figs. 1K, M View FIGURE 1 ); styli appearing connected ventrally to distal end of parameres ( Figs. 1L, M View FIGURE 1 ).......................................... Neoconis szirakii sp. n.

7. Penis ending in a single spine ( Meinander 1972, fig. 16F)........................... Aleuropteryx unicolor Meinander

7’. Penis ending in two or three spines ( Fig. 3C View FIGURE 3 ; Meinander 1972, figs. 13C, 14D).................................... 8

8. Penis ending in two short spines ( Meinander 1972, fig. 14C; Meinander 1974, fig. 3B); apohyses of ninth sternite long and slen- der ( Meinander 1972, fig. 14D), if apophyses are short, they are slightly curved inwards in ventral view ( Fig. 3C View FIGURE 3 ; Meinander 1974, figs. 2B, 3B).................................................................................... 9

8’. Penis ending in three spines ( Meinander 1972, fig. 13C); apohyses of ninth sternite short and straight in ventral view ( Fig. 3C View FIGURE 3 ; Meinander 1972, fig. 13C)................................................ Aleuropteryx maculipennis Meinander

9. Two short spines of penis situated side by side ( Fig. 3E View FIGURE 3 ; Meinander 1972, fig. 14C)....... Aleuropteryx simillima Meinander

9’. One spine of penis situated above the other ( Meinander 1974, fig. 2A).......................................... 10

10. Spines of penis directed caudally in lateral view ( Meinander 1974, fig. 3C); wing membrane distinctly spotted ( Meinander 1974, fig. 3D).............................................................. Aleuropteryx punctata Meinander

10’. Spines of penis directed upwards in lateral view ( Meinander 1974, fig. 2A); wing membrane without spots, unicolored grey ( Meinander 1974, fig. 2D).................................................. Aleuropteryx longipennis Meinander

11. M of hindwing unforked ( Fig. 2A View FIGURE 2 ; Meinander 1972, fig. 166A)… Coniopteryx .................................... 12

11’. M of hindwing forked ( Meinander 1972, fig. 214E; Meinander 1974, fig. 13A)................................... 25

12. Gonarcus and hypandrium separated ( Meinander 1972, figs. 128A, 168B)....................................... 13

12’. Gonarcus and hypandrium synscleritous ( Fig. 2D View FIGURE 2 ; Meinander 1972, fig. 149A; Meinander 1975, fig. 2A)… C. (Scotoconiopteryx).............................................................................................. 14

13. Stylus arising from the caudal end of gonarcus ( Meinander 1972, figs. 163A, 166A); flagellomeres with scale-like hairs (except in C. delta ) ( Meinander 1972, figs. 165A, 166D; Johnson 1977, fig. 1F)… C. ( Coniopteryx s. str.)..................... 16

13’. Stylus arising well before the caudal end of gonarcus ( Meinander 1972, figs. 128A, 129A); flagellomeres (in Mexican species) without scale-like hairs ( Meinander 1972, fig. 129F)... C. ( Xeroconiopteryx )… Coniopteryx diversicornis group.......... 23

14. Tips of processus terminalis on both sides of the median incision long, narrow and pointed ( Fig. 2G View FIGURE 2 ; Meinander 1972, figs. 149A, E)........................................................................................... 15

14’. Tips of processus terminalis on both sides of the median incision more or less broad, short and fairly blunt ( Meinander 1975, figs. 2A, C)................................................................ Coniopteryx (S.) flinti Meinander

15. Distal end of paramere abruptly bent downward in lateral view ( Meinander 1972, fig. 149C)............................................................................................... Coniopteryx (S.) isthmicola Meinander

15’. Distal end of paramere directed caudally in lateral view ( Figs. 2C, D View FIGURE 2 ) ...................... Coniopteryx (S.) josephus sp. n.

16. Segment four (some cases also segment three) of maxillary palpi modified ( Fig. 4C View FIGURE 4 ); frons modified with projections with bristles ( Meinander 1972, fig. 166F)........................................... Coniopteryx westwoodi group…17

16’. Segments of maxillary palpi not modified; frons without any modifications … Coniopteryx tineiformis group.............20

17. Ventral line of hypandrium together with the processus terminalis straight in lateral view ( Meinander 1972, fig. 163A).................................................................................. Coniopteryx (C.) fitchi Banks

17’. Ventral line of hypandrium distinctly bent upwards ( Fig. 4D View FIGURE 4 ; Meinander 1972, fig. 166A)........................... 18

18. Third segment of maxillary palpi with a large, forwards directed thorn ( Fig. 4C View FIGURE 4 ; Meinander 1972, fig. 166F).................................................................................... Coniopteryx (C.) palpalis Meinander

18’. Third segment of maxillary palpi without forwards directed thorn.............................................. 19

19. Ventral part of processus apicalis hooky and sharp ( Fig. 5B View FIGURE 5 ; Meinander 1972, fig. 168E)...................................................................................................... Coniopteryx (C.) westwoodi (Fitch)

19’. Ventral part of processus apicalis blunt, with an inwards directed dentiform projection ( Meinander 1974, figs. 10B, 10C)......................................................................... Coniopteryx (C.) mexicana Meinander

20. Most antennal segments with scale-like hairs ( Meinander 1972, fig. 162F)....................................... 21

20’. Antenal segments without scale-like hairs ( Johnson 1981, fig. 1F)....................... Coniopteryx (C.) delta Johnson

21. Anterior apodema of hypandrium ventrally complete, and curving backwards as far as the median incision ( Meinander 1972, fig. 165B), or it has a backwards directed median branch ( Meinander 1972, fig. 167D)............................. 22

21’. Anterior apodema of hypandrium often incomplete, and not curving backwards as far as median incision, without a backwards directed median branch ( Meinander 1972, fig. 162C)........................... Coniopteryx (C.) callangana Enderlein

22’. Median incision of processus terminalis moderately wide, V-shaped ( Meinander 1972, fig. 165C).............................................................................................. Coniopteryx (C.) minuta Meinander

23. Gonarcus long ( Fig. 5D View FIGURE 5 ; Meinander 1972, fig. 128B); tips of processus terminalis pointed on both sides of the median incision ( Fig. 5D View FIGURE 5 ; Meinander 1972, fig. 128D)..................................... Coniopteryx (X.) diversicornis Meinander

23’. Gonarcus short ( Meinander 1972, fig. 129B; Johnson 1981, fig. 4B); tips of processus terminalis very small or absent.... 24

24. A short processus terminalis of hypandrium present, with a narrow V-shaped median incision ( Johnson 1981, fig. 4C)............................................................................ Coniopteryx (X.) meinanderi Johnson

24’. Processus terminalis of hypandrium absent, consequently median incision also absent ( Meinander 1972, figs. 129A, C)............................................................................. Coniopteryx (X.) texana Meinander

25. Cross-vein M-Cu1 of both wings at right angle ( Meinander 1972, figs. 182G, 191E), striking the stem of M well before its fork, at least in forewing ( Meinander 1972, fig. 192E)........................................................... 26

25’. Cross-vein M-Cu1 of both wings oblique, striking the branch M 3+4 or the fork of M ( Meinander 1972, fig. 214E; Meinander 1980, fig. 11F) Semidalis .............................................................................. 27

26. Hindwing shortened or not shortened ( Meinander 1972, fig. 192E), in latter case Rs originates at the middle of this wing ( Meinander 1972, fig. 191E); ectoprocts ventrally with large digit-like outer processes ( Meinander 1972, fig. 192B) Conwentzia .. .................................................................................................. 40

26’. Hindwing normally developed, Rs originates basally of the middle of this wing ( Fig. 6A View FIGURE 6 ; Meinander 1972, fig. 182G); ectoprocts without outer processes ( Fig. 6B View FIGURE 6 ; Meinander 1972, fig. 182A)… Parasemidalis ( Parasemidalis s. str.)................................................................................... Parasemidalis (P.) fuscipennis (Reuter)

27. Middle or distal part of paramere much wider than proximal part (apart from the possible club of the latter); wide part plate-like or lobiform, or divided into hooks or broad teeth ( Meinander 1972, figs. 209B, 218B; Johnson 1980, fig. 7D)........... 28

27’. Paramere not or only moderately widened distally; when widened, not plate-like or lobiform, but swollen; in Mexican species divided into narrow and, at least partly, acute branches running caudally or dorso-caudally ( Meinander 1972, fig. 214C; Meinander 1975, fig. 3G; Monserrat 1985, fig. 18) … Semidalis boliviensis group...................................... 29

28. Widened part of paramere (before the hooked ending) rectangular and situated dorsally ( Meinander 1972, fig. 209B); hypandrium (in Mexican species) caudally terminating in a long spine ( Meinander 1972, fig. 209A)… Semidalis angusta group................................................................................... Semidalis angusta (Banks)

28’. Widened part of paramere not rectangular or trapezoid ( Meinander 1972, figs. 217C, 218B); hypandrium in most instances without distinct long spine ( Meinander 1972, fig. 218A; Meinander 1975, fig. 4A)… Semidalis aleyrodiformis group..... 35

29. Paramere slightly swollen distally; distal end consists of a dorso-caudally directed acute thorn, a caudally directed, slightly obtuse tooth and a dorso-caudally directed membranous branch ( Meinander 1972, figs. 214B, C)................................................................................................ Semidalis boliviensis (Enderlein)

29’. Paramere slender, distally divided into acute branches ( Meinander 1972, fig. 216B; Meinander 1980, fig. 11D; Monserrat 1985, fig. 18)............................................................................................ 30

30. Ectoproct with well-developed caudal projection directed inwards and/or backwards ( Monserrat 1985, figs. 9, 16, 17).... 31

30’. Ectoproct without distinct caudal projection ( Fig. 7D View FIGURE 7 ; Meinander 1972, fig. 216A; Meinander 1975, fig. 3A)........... 33

31. Caudal projection of ectoproct an acute hook ( Monserrat 1985, figs. 15, 17)...................................... 32

31’. Caudal projection of ectoproct a long, duplicate spine ( Monserrat 1985, fig. 9)................. Semidalis soleri Monserrat

32. A short dorsal branch anterior to the forked main branch of the distal part of paramere ( Meinander 1980, fig. 11D)................................................................................. Semidalis manausensis Meinander

32’. No dorsal branch anterior to the forked main branch of the distal part of paramere ( Monserrat 1985, fig. 18)....................................................................................... Semidalis problematica Monserrat

33. Processus terminalis of hypandrium rounded conic and snouted, with a minute median incision ( Meinander 1972, figs. 216A, D)......................................................................... Semidalis mexicana Meinander

33’. Processus terminalis of hypandrium tapering tongue-like ( Fig. 7E View FIGURE 7 ; Meinander 1975, figs. 3A, E)..................... 34

34. Median incision of processus terminalis of hypandrium deep V-shaped ( Fig. 7E View FIGURE 7 ; Meinander 1975, fig. 3D)....... Semidalis hidalgoana Meinander

34’. Median incision of processus terminalis of hypandrium shallow ( Meinander 1975, fig. 3E)... Semidalis sonorana Meinander

35. Paramere swollen distally, expansion of distal part oviform ( Meinander 1972, fig. 210B; Meinander 1972, fig. 218B; Johnson 1980, fig. 7D).................................................................................. ……..36

35’. Paramere slender distally, distal part only slightly wider than proximal one ( Meinander 1972, fig. 217C; Meinander 1974, fig. 13D).............................................................................................. 39

36. Uncini fused into a single transverse plate ( Meinander 1972, figs. 210A, C; Johnson 1980, figs. 7E, F)................ 37

36’. Uncini not fused, long or claw-like in lateral view (in Mexican species) ( Meinander 1972, fig. 218B; Meinander 1975, fig. 4C)............................................................................................... 38

37. Paramere apically with one upwards directed hook terminally and one backwards directed hook in the middle ( Fig. 7B View FIGURE 7 ; Meinander 1972, figs. 210B)......................................................... Semidalis arnaudi Meinander

37’. Paramere apically with a deep dorsal incision formed by body of paramere and dorsally directed hook terminally, and a large dorsally directed spine in the middle (Johnson 1980, fig. 7D).............................. Semidalis tricornis Johnson

38. Widened part of paramere with a proximal hook or spine ( Meinander 1972, fig. 218B).......... Semidalis flinti Meinander

38’. Widened part of paramere without a proximal hook or spine ( Meinander 1975, fig. 4C)................................................................................................ Semidalis frommeri sensu Meinander 1975

39’. Preapical tooth of distal part of paramere absent, the apical one is minute in lateral view ( Meinander 1974, fig. 12D).................................................................................... Semidalis deserta Meinander

40. Hindwing normally developed ( Fig. 6C View FIGURE 6 ; Meinander 1972, fig. 191E); outer process of ectoprocts short and blunt, about twice as broad as long ( Fig. 6D View FIGURE 6 ; Meinander 1972, figs. 191A, B, C); tips of parameres parallel, close to each other in caudal view ( Fig. 6D View FIGURE 6 ; Meinander 1972, fig. 191B)................................................... Conwentzia barretti (Banks)

40’. Hindwing shortened or reduced ( Meinander 1972, fig. 192E); outer process of ectoprocts tapering, about as long as breadth at base ( Meinander 1972, figs. 192A, B); tips of parameres diverging in caudal view ( Meinander 1972, fig. 192C)................................................................................... Conwentzia californica Meinander