Dolichognatha deelemanae , Smith, HELEN M., 2008

Smith, HELEN M., 2008, Synonymy of Homalopoltys (Araneae: Araneidae) with the genus Dolichognatha (Araneae: Tetragnathidae) and descriptions of two new species, Zootaxa 1775, pp. 1-24: 13-15

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Dolichognatha deelemanae

sp. n.

Dolichognatha deelemanae  sp. n.

Figs 29-37

Holotype. INDONESIA: Kalimantan: ♂, RMNH. ARA.11299 (ex coll. CLD), SE Kalimantan, Meratus Mtn, E of Lake Riam Kanan, 3.vii.1980, C.L. & P.R. Deeleman, primary forest.  Paratypes. INDONESIA: Kalimantan: 1 ♂, RMNH. ARA.11300,  1 ♀, RMNH.ARA.11301, data as holotype. 

Etymology. The specific epithet honours Christa Deeleman-Reinhold, who collected the specimens which originally sparked interest in this group.

Diagnosis. Male. Palpal bulbus short and rounded (Figs 33, 34), abdomen with apical ‘drip’ point (Fig. 31, may vary), palpal tibia almost as wide as long, MEA broadens only slightly distally.

Female. Abdomen apically with point, copulatory openings of epigynum set almost on posterior margin in ventral view, ducts shorter than in D. albida  (Fig 35 c.f. Fig. 26), internal genitalia delicate (Fig. 37, c.f. D. albida, Fig.  28).

Description. Male (holotype). Carapace (Fig. 29): length 0.78 (range 0.78-0.80), width 0.61, height 0.39; caput strongly humped in lateral view; similar to D. mandibularis  in dorsal view; caput margins produced to distinct projections anterolaterally over cheliceral bases (arrowed in Fig. 32). Chelicerae (Figs 29, 32): paturon length shorter than in other species (c. 0.8 × carapace length), basal ledge strongly produced, two distinct promarginal cheliceral teeth and one large intermediate, which is the largest tooth, three retromarginal teeth, basal mound absent; cheliceral fangs shorter than other species, and not sinuous, tips arced. Labium: with distinct, but rather recumbent, spur (Fig. 29). Sternum: distinctly convex (Fig. 29). Eyes (Figs 29, 32). AME: 0.10, PME: 0.08, ALE: 0.065, PLE: 0.065, AME–AME: 0.04, AME–ALE: 0.025, PME–PME: 0.04, PME– PLE: 0.06, ALE–PLE: 0.025; no reflective tapeta visible in either male. Legs: I: 3.76, II: 3.12, III: 1.67, IV: 2.24; femur I with strong prolateral macroseta at mid point, femur II with single retrolateral macroseta at c. ¾ length; front femurs with at least one distal dorsal macroseta as in other species. Abdomen: length 1.94, width 0.86; apex produced into drip-shaped apex (Fig. 31); post-genital mound strongly developed but without coloration (Fig. 30). Palpal organ (Figs 33, 34): palpal femur 0.29 × carapace length; patella without macroseta, tibia short compared to other species, distally almost as wide as long; embolus a curved rod; MEA relatively slender, only slightly broadened distally; overall shape of bulbus similar to D. mandibularis,  width: length = 1:1.5. Colour in alcohol: almost entirely creamy-white; median eyes with black around margins; faint trace of colour on dorsal carapace.

Female ( RMNH.ARA.11301). Carapace: length 0.86, width c. 0.55 (slightly squashed), height c. 0.41; caput strongly domed, sides bulging. Chelicerae: promargin with two definite teeth plus large intermediate where rows meet; retromargin with one large and 4 small teeth grouped distally (rather indistinct). Labium: mounded on anterior face. Sternum: deep, strongly convex. Eyes. AME: 0.10, PME: 0.07, ALE:0.065, PLE: 0.07, AME–AME: 0.02, AME–ALE: 0.03, PME–PME: 0.04, PME–PLE: 0.065, ALE–PLE: 0.03; no reflective tapeta visible (specimen not in good condition). Legs: I: 3.29, II: 2.94, III: 1.57, IV: 2.12; macrosetae near midpoint of prolateral femur I and 4/5 length of retrolateral femur II. Abdomen: length 1.14, width N/A (squashed); attenuated anterior tip not very well developed, moderately developed post-genital mound. Epigyne: copulatory openings set almost on posterior margin in ventral view, ducts viewed through cuticle short and curved into a flattened bowl shape (Fig. 35); in posterior view the internal margins of the lateral plates diverge sharply towards the copulatory openings (Fig. 36). Internal genitalia (Fig. 37): as D. albida  but distinctly delicate. Colour in alcohol: creamy-white, traces of dark colouring on dorsal carapace and around eyes. Abdomen pale.

Distribution. Dolichognatha deelemanae  is only recorded from south-eastern Kalimantan.


Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]


USA, Florida, Gainesville, University of Florida, Florida Museum of Natural History, Allyn Museum