Cimeliomorpha Diakonoff, 1966

Pinkaew, Nantasak & Horak, Marianne, 2019, Revision of the genus Cimeliomorpha Diakonoff (Lepidoptera: Tortricidae), Zootaxa 4615 (3), pp. 457-480 : 458

Cimeliomorpha Diakonoff, 1966

Cimeliomorpha Diakonoff, 1966: 50 , fig. 1. Obraztsov, 1968: 185, figs 9‒13; Robinson et al., 1994: 104, plate 17 fig. 5; Kuznetsov, 1997: 801, fig. 7; Brown et al., 2005: 177.

Type species: Copromorpha cymbalora Meyrick, 1907 , by original designation.

Diagnosis. Cimeliomorpha has an unmistakable two-toned forewing pattern with a uniformly yellow or white basal half and a distal half with complex mostly black pattern on red-brown ground with raised silvery lines and spots, and a hindwing that is either white or orange at the base, with a narrow to wide blackish band around its margin. Anthozela Meyrick and some Loboschiza species also have a conspicuously two-toned forewing, but in the former the basal half is yellow and either speckled with black or suffused with orange, and in the latter the hindwing is not white or yellowish in its basal half. The genitalia of both sexes of Cimeliomorpha also indicate a close relationship with Loboschiza , but not with Anthozela . The wing venation with widely separated R s and M 1 and parallel M 2 and M 3 in the hindwing ( Fig. 3 ) is most similar to that of Irianassa Meyrick , but this is due rather to symplesiomorphy than to a close relationship as evidenced by the very different genitalia in both sexes.

Description. Head. Ocellus and chaetosemata well developed. Frons, vertex and labial palpus white in the cymbalora -group and yellow in the egregiana -group, labial palpus rather short, thick and upcurved, second segment slightly widened medially and apical segment triangular; antenna short, not reaching middle of wing, basal segments of flagellum covered with scales and remainder of flagellum only with few black scales dorsally; cilia minute.

Thorax. Without raised posterior scale tufts. Pronotum, tegulae and mesonotum white in the cymbalora -group and yellow in the egregiana -group. Forewing length 5.8−9.9 mm; medium-sized to large. Forewing broadly subtri- angular, costa evenly curved, male costal fold absent, termen rather round, weakly concave below apex; basal 2/5 of wing white in the cymbalora -group and yellow in the egregiana -group, distal 3/5 with light brown to orange brown ground colour, with a broad transverse silvery band along its inner margin, angled near costa, with 4−5 oblique sil- very lines from costa, ocelloid patch a large bean-shaped to roundish area of white-tipped orange brown scales producing fine transverse white lines, with a central band of longitudinal blackish striation scattered with blackish dots, distally delineated by two convergent silvery streaks; with 4 large spots of raised silvery scales, shadowed by black and some orange: an angled series of three behind CuA 1, the fourth at the anterior angle of the discal cell. Hindwing with broad blackish band along termen, basally white in the cymbalora -group and orange in the egregiana -group.

Abdomen. Male genitalia with tegumen rounded dorsally, pedunculus with large anterior subtriangular process, uncus absent, socii broad and hairy, vinculum moderately broad, valva with large ovate to subrectangular sacculus, cucullus variable in shape. Female genitalia with papillae anales elongate and moderately broad, densely setose; sternum VII moderately sclerotised, with patch of scale sockets on lamella postvaginalis, ductus bursae with short to long colliculum, corpus bursae with a small granulate signum or two horn-shaped signa.

Biology. Meyrick (1939) reported for Laspeyresia cymbalora Meyrick “Buitenzorg [Bogor, Java], bred March from larvae feeding on leaves of Derris elliptica Bentham (Fabaceae) (Dr. J. van der Vecht).” To this is added a note by T.B.F. [Thomas Bainbrigge Fletcher] that the species in question is in fact novarana Felder & Rogenhofer.

The cymbalora -group. Forewing length 5.8−7.2 mm. Frons, vertex, labial palpus, pronotal collar, tegulae and mesonotum white. Forewing with basal 2/5 white; hindwing with basal half white; male genitalia with membrane next to juxta without setae, cucullus tip pointed and ending in short strong spine; female genitalia with one small signum, a granulate sclerite.

The egregiana- group. Forewing length 7.2−9.9 mm. Frons, vertex, labial palpus, pronotal collar, tegulae and me- sonotum yellow. Forewing with basal 2/5 yellow; hindwing with basal part orange or at least with large orange patch in centre; male genitalia with membrane next to juxta with numerous short setae, cucullus distally rounded and not ending in a single long spine ( C. jarujini with one strong spine centrally); female genitalia with two horn-shaped signa.

Figures

FIGURE 3. Wing venation of Cimeliomorpha cymbalora male (Thailand, KKIC, np1735).