Sphiximorpha euprosopa ( Loew, 1869 )

Steenis, Jeroen Van, Ricarte, Antonio, Vujić, Ante, Birtele, Daniele & Speight, Martin C. D., 2016, Revision of the West-Palaearctic species of the tribe Cerioidini (Diptera, Syrphidae), Zootaxa 4196 (2), pp. 151-209 : 191-193

Sphiximorpha euprosopa ( Loew, 1869)

Figs 118 , 126 , 132 , 148–150 , 152 , 169

Ceria euprosopa Loew, 1869: 256 . Type locality: Turkey, Izmir [HT ♂ ZMHB]. Cerioides euprosopa: Sack (1932) comb. nov.

Sphiximorpha euprosopa: Peck (1988) comb. nov., Dirickx (1994), Vujić et al. (2009), Sarıbıyık (2014).

Redescription. MALE ( Figs 118 , 126 ). Body length: 9.8 mm; wing length: 7.2 mm. Head ( Fig. 132 ). Face slightly protruding antero-ventrally with very weakly demarcated tubercle and strongly concave below antennae. Head 1.7 times wider than face just below the antennae; length of eye contiguity 0.56 times as long as length of frons; angle of eyes at eye contiguity 140o. Face with yellow and black colour pattern; genae, mouth edge and hypostomal bridge black; medial black vitta tridentate with yellow medial area; a large black triangular macula around antennal insertion. Frons yellow. Vertical triangle and dorsal surface of head capsule yellow. Ocellar triangle black. Frontal prominence 0.38 times as long as wide; relative length of pedicel is as 1.5: 1: 1.4. Antennae brown-yellow to black coloured; arista white pilose. Thorax. Scutum black with yellow macula on postpronotum and notopleuron. Pleuron black with two yellow maculae, one each on posterior part of posterior anepisternum and on dorsal part of katepisternum. Pile long and white, ventral half of anterior anepisternum with pile about 2/3 as long as pile on adjacent part of dorsal surface of head capsule. Scutellum yellow with very narrow posterior margin black. Legs. Coxae and trochanter black; pro- and mesofemur predominantly black, only some yellow basally and apically; basal 1/8 and apical 1/10 of metafemur yellow; apico-medial 1/4–1/2 of tibiae black; pro- and mesotarsi darkyellow to dark-brown; metatarsus dark-brown to black. Femora with black setulae apicodorsally; metatrochanter without setulae. Mesofemur with small flattened area antero-basally; metatrochanter relatively narrow with weak sulcus and normal rim laterally. Metafemur elongate and metatibia without appendix on apico-ventral part, elongate, narrow basally and clearly widening medially ( Fig. 152 ).

Wing. Hyaline except for anterior 1/2 to spurious vein and along vein CuA. Vein R4+5 straight with short appendix into cell r4+5. Cross-vein r-m slightly curved. Membrane covered with microtrichia; alula with anterior margin densely microtrichose, other part bare or with very scattered microtrichia; anterobasal 1/3 of cell cup and a narrow area along anterobasal 1/5 of anal lobe bare; alula relatively narrow, 3.7 times longer than wide. Abdomen. Black and yellow coloured. Length of tergite I: II: III: IV is as 1: 2.7: 2.7: 3.4. Width of yellow fascia medially: length of tergite of respectively tergite II, III and IV as 1: 5.4, 1: 5.4 and 1: 3.8. Tergite II slightly longer than wide, anteromedial part most narrow; length of tergite II: width of tergite II at respectively anterior: narrowest: posterior as 1: 0.74: 0.63: 1.0. Tergite II with triangular maculae on antero-lateral corner, rather widely separated medially, ratio of black between maculae and width of tergite anteriorly 1: 2.0; fascia on tergite II medially broad, narrowed towards lateral margin; fascia on tergite III medially concave and slightly but gradually narrowed towards lateral margin; fascia on tergite IV straight, gradually narrowed towards lateral margin. Tergite IV weakly emarginated. Tergite IV with short bow formed grey-white pollinosity medio-laterally. Sternites I–III posterior with straight yellow fascia. Tergite VIII white pilose.

Genitalia. Epandrium with narrow ventral rim ( Figs 148, 149 ); in dorsal view, cerci elongate semi-circular shaped ( Fig. 149 ), pile about as long as width of cerci; surstylus bi-lobed, dorsal lobe rectangular elongate basally broad and with ventro-basal squarish flange, ventral lobe small semi-circular with membranous basal part ( Fig. 148 ); surstylar apodeme rectangular, separated in two broad elongate sclerotized parts ( Fig. 150 ); hypandrium ( Fig. 148 ) very broad, nearly squarish; with short, broad ventral lobe and rugged area basally; superior lobe balloon shaped, articulating with hypandrium; aedeagus with short and broad baso-ventral projection, apex with three projections, dorsally and ventrally short and rounded, medially somewhat longer and rectangular. FEMALE: no description available.

Material examined. Holotype of Ceria euprosopa : 1 Ƌ, " Smyrna / 28-3-[18]63", "Coll. / H. Loew ", " Typus " [faded red label], " Ceria / euprosopa / Lw. " [handwritten], Zool . Mus. / Berlin ", " Holotype Ƌ / Ceria euprosopa / Loew, 1869 / det. J. van Steenis, 2016" [red label] ( ZMHB) . Additional material: 1 Ƌ, 1 ♀ (FSUNS); not studied by the senior author and hence no description of the female is available.

Distribution ( Fig. 169 ). Greece, Turkey.

Biology. Adults fly from late March to April.

Figures

FIGURES 118 – 121. Adults of Sphiximorpha species, overall appearance, dorsal view. 118 Sphiximorpha euprosopa male, holotype 119 Sphiximorpha petronillae female, lectotype 120 Sphiximorpha subsessilis male, the Netherlands 121 Sphiximorpha subsessilis female, Greece. Scale line = 1 mm.

FIGURES 122 – 129. Adults of Primocerioides and Sphiximorpha species, overall appearance, lateral view. 122 Primocerioides regale male (holotype of Sphiximorpha hiemalis) 123 Primocerioides regale female, Cyprus 124 Sphiximorpha garibaldii male, holotype 125 Sphiximorpha garibaldii female, (holotype of Ceria tridens) 126 Sphiximorpha euprosopa male, holotype 127 Sphiximorpha petronillae female, lectotype 128 Sphiximorpha subsessilis male, Greece 129 Sphiximorpha subsessilis female, Greece. Scale line = 1 mm.

FIGURES 130 – 137. Head of Primocerioides and Sphiximorpha species, frontal view. 130 Primocerioides regale male (holotype of Sphiximorpha hiemalis) 131 Primocerioides regale female, Cyprus 132 Sphiximorpha euprosopa male, holotype 133 Sphiximorpha garibaldii male, holotype 134 Sphiximorpha garibaldii female, (holotype of Ceria tridens) 135 Sphiximorpha petronillae female, lectotype 136 Sphiximorpha subsessilis male, Greece 137 Sphiximorpha subsessilis female, Greece. Scale line = 1 mm.

FIGURES 148 – 150. Male genitalia of Sphiximorpha euprosopa, holotype 148 Epandrium and hypandrium, lateral view 149 Epandrium, dorsal view 150 Epandrium, ventral view. Scale line = 0.5 mm.

FIGURES 151 – 156. Metafemur and tibia of Primocerioides and Sphiximorpha species, lateral view. 151 Primocerioides regale male (holotype of Sphiximorpha hiemalis) 152 Sphiximorpha euprosopa male, holotype 153 Sphiximorpha garibaldii male, holotype 154 Sphiximorpha petronillae female, lectotype 155 Sphiximorpha subsessilis male, The Netherlands 156 Sphiximorpha subsessilis female, Greece. Scale line = 1 mm.

FIGURES 169 – 172. Distribution maps of the West-Palaearctic Cerioidini. 169 Ceriana glaebosa (red square), Sphiximorpha euprosopa (green circle) and Sphiximorpha petronillae (blue triangle) 170 Ceriana vespiformis (red circle) 171 Primocerioides regale (red circle) and Sphiximorpha garibaldii (blue triangle) 172 Sphiximorpha subsessilis (yellow circle).