Eurythenes gryllus (Lichtenstein, 1822)
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https://doi.org/10.5281/zenodo.5395773 |
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https://treatment.plazi.org/id/2D09EC23-E90E-FFAD-FFDD-FDE9FF75FAA6 |
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treatment provided by |
Marcus (2021-08-30 12:02:35, last updated by Plazi 2023-11-05 23:43:32) |
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scientific name |
Eurythenes gryllus |
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Eurythenes gryllus View in CoL (Lichtenstein in Mandt, 1822) ( Figs 1-11 View FIG View FIG View FIG View FIG View FIG View FIG View FIG View FIG View FIG View FIG View FIG )
Gammarus gryllus Lichtenstein in Mandt, 1822: 34.
Lysianassa magellanica Milne Edwards 1848: 398 . — Lucas 1857: 13, pl. 1 fig. 3. — Bate 1862: 66, pl. 10 fig. 5; 1866: 330. — Lilljeborg 1865a: 3; 1865b: 2. — Bethune 1869: 431. — M. Sars 1869: 260.
Eurytenes magellanicus – Lilljeborg 1865a: 11, pls 1-3; 1865b: 6. — Sars 1891: pl. 30.
Lysianassa gryllus – Goes 1866: 517, pl. 36 fig. 1. — Bate 1867: 229.
Eurytenes gryllus – Boeck 1871: 105; 1872: 144. — Stuxberg 1880: 62. — Stephensen 1912b: 528.
Eurythenes gryllus View in CoL – Smith 1884a: 54. — Hansen 1888: 67. — Chevreux 1889: 298, fig.; 1935: 50, pl. 1, figs 6, 7 — Stebbing 1893: 80; 1906: 73. — Chilton 1911: 563. — Stephensen 1925: 110; 1932: 356 (in part, part = E. obesus View in CoL ); 1933: 12, figs 4, 5 (in part, part = E. obesus View in CoL ); 1935: 91; 1942: 474 (in part, part = E. obesus View in CoL ); 1949: 3 (?in part, part may = E. obesus View in CoL ). — Schellenberg 1927: 679, fig. 70; 1955: 192. — K. H. Barnard 1937: 144. — Shoemaker 1945: 186 (in part, part = E. obesus View in CoL ); 1956: 177. — Gorbunov 1946: 43. — Gurjanova 1951: 265, fig. 134; 1962: 340; 1964: 277. — Dahl 1954: 3. — Birstein & Vinogradov 1955: 225; 1958: 228; 1960: 183; 1962: 36; 1970: 420 (table 1). — Hurley 1957: 2 (in part, part may = E. obesus View in CoL , part probably = E. thurstoni View in CoL ). — J. L. Barnard 1958: 92; 1961: 35 (in part), fig. 5 (material from South Pacific and figs 6, 7 = E. thurstoni View in CoL ). — Oldevig 1959: 19. — Templeman 1967: 215, figs 1-3. — Beck 1969: 34, 35. — Bowman & Manning 1972: 193, figs 2-5. — Paul 1973: 289. — Rannou & Nouguier 1974: 142. — Shulenberger & Hessler 1974: 185. — Dahl et al. 1976: 75, 78. — Lowry & Bullock 1976: 90. — Shulenberger & Barnard 1976: 241. — Griffiths 1977: 97. — Intes 1978: 4, fig. 4. — Andres 1979: 96; 1983: 186; 1987: 96. — Dahl 1979: 168, fig. 6. — George 1979a: 283; 1979b: 63. — Hessler et al. 1979: 704. — Ortiz 1979: 19. — Smith et al. 1979: 57. — Thurston 1979: 56; 1990: 262; 1994: 14, unnumbered fig.; 2000: 684 (table 2), 688, 690. — Hallberg et al. 1980: 280. — Just 1980: 164 (table 2), 167, 171. — Prince 1980: 63 (table 4). — Hessler 1981: 397. — Kamenskaya 1981: 95, figs 1, 2. — Umezu 1982: 2; 1984: 128. — Ingram & Hessler 1983: 683; 1987: 1889. — Lampitt et al. 1983: 73. — Petter 1983: 177, fig. 1. — Smith & Present 1983: 183. — Wickins 1983: 83. — Smith & Baldwin 1984: 1179. — Desbruyères et al. 1985: 233. — Hargrave 1985: 443. — Hopkins 1985: 202, 206, 210. — Laver et al. 1985: 1136. — Rauschert 1985: 319. — Wilson et al. 1985: 1248. — Ainley et al. 1986: 848. — Hasegawa et al. 1986: 70. — Reinhardt & Van Vleet 1986: 151, 157. — Baldwin & Smith 1987: 425. — Bucklin et al. 1987: 1795. — Charmasson & Calmet 1987: 1509; 1989: 159; 1990: 227. — Sainte- Marie & Hargrave 1987: 436. — Calmet & Charmasson 1989: 163. — Costello et al. 1989: 32. — Christiansen et al. 1990: 35; 2001: 2409. — Rauschert 1990: 454. — Vinogradov 1990: 42, 92; 1997: 77. — Wakabara et al. 1990: 5; 1996: 355 (table 1), 360 (table 2). — Barnard & Karaman 1991: 486. — Boudrias 1991: 13, figs 1, 2; 2002: 581. — Gage & Tyler 1991: 78. — Gonzalez 1991: 59. — Palerud & Vader 1991: 35. — Rauschert 1991: 37, 39. — Sainte-Marie 1991: 217; 1992: 105. — Steele & Steele 1991: 1250, figs 3, 4. — Hargrave et al. 1992a: 37; 1992b: 41; 1994: 1489; 1995: 1905. — Smith 1992: 1040. — Smith et al. 1992: 669, 671 (table 3). — Britton & Morton 1993: 369; 1994: 391. — Christiansen & Thiel 1993: 12. — De Broyer & Jazdzewski 1993: 67. — Heinrich et al. 1993: 6. — Lopes et al. 1993: 209. — Ishimaru 1994: 58. — Kaufmann 1994: 54. — Thurston & Bett 1995: 201. — Christiansen 1996: 345. — France & Kocher 1996a: 633 (in part, part = E. thurstoni View in CoL ); 1996b: 304. — Vinogradov et al. 1996: 8. — Gebruk et al. 1997: 116. — Jones et al. 1998: 1124. — Matsumasa et al. 1998: 686. — Takeuchi & Watanabe 1998: 285. — Creasey & Rogers 1999: 28, 32, 42, 43, 50, 78, 81, 87, 96, fig. 11. — Witte 1999: 142. — Christiansen & Martin 2000: 3027. — Janssen et al. 2000: 3011 (table 5). — Bluhm et al. 2001: 642. — Bühring & Christiansen 2001: 369. — Dauby et al. 2001: 81. — Johnson et al. 2001: 198 (table 3). — Klages et al. 2001: 293. — Takeuchi et al. 2001: 653. — Cherel et al. 2002a: 272 (table 6); 2002b: 288 (table 3), 292 (table 4). — Thurston et al. 2002: 205. — Treude et al. 2002: 1281. — Premke et al. 2003: 283.
Euryporeia gryllus View in CoL – Sars 1891: 86, [pl. 30 as Eurytenes gryllus ]. — Della Valle 1893: 848, pl. 60 fig. 58. — Chevreux 1895: 426; 1899a: 147, 148; 1899b: 152; 1900: 24, pl. 14 fig. 4; 1903: 96; 1905: 7; 1910: 4. — Holmes 1903: 277. — Brüggen 1907: 215. — Grieg 1925: 21.
Eurythenes magellanicus View in CoL – K. H. Barnard 1932: 59. — J. L. Barnard 1958: 92. — Beck 1969: 35. — Conroy 1972: 56. — Arnaud 1974: 592 (list).
Eurysthenes gryllus – Smith 1884b: 181 (lapsus calami). Eurythenes grillus – Gilchrist & MacDonald 1980: 35 (lapsus calami).
Eurythennes gryllus – Poupin et al. 1990: 94, pl. 2 fig. g. — Poupin 1994: 16; 1996: 42, fig. h. (lapsus calami).
Eurythenes cf. gryllus View in CoL – Lowry & Stoddart 1994: 158 (in part, part = E. thurstoni View in CoL ).
Unnamed specimen in photograph – Yayanos 1978: 1056, fig. 1.
Not Eurythenes gryllus View in CoL – Murdoch 1885 (= Anonyx nugax View in CoL ). — K. H. Barnard 1940: 440, 515 (list) (= E. obesus View in CoL ). — Springer & Bullis 1956: 6 (= E. obesus View in CoL ). — Lowry & Stoddart 1993: 72 (= E. thurstoni View in CoL ).
DIAGNOSIS. — Anterodorsal margin of head smooth. Gnathopod 1 parachelate; basis length 2 to 2.5 times breadth; propodus slightly tapering distally. Pereopods 3 to 7 dactyli short. Pereopod 4 coxa as deep as wide. Pereopod 7 basis, length of anterior margin subequal to breadth; posteroventral margin straight. Pleonite 3 with anterodorsal notch. Epimeron 3 posteroventral corner broadly rounded. Urosomite 1 not dorsodistally produced over urosomite 2. Uropod 3 peduncle, medial face without robust setae.
LICHTENSTEIN’ S SPECIMEN ( FIGS 1-3 View FIG View FIG View FIG )
Holotype (of Gammarus gryllus Lichtenstein in Mandt, 1822), female, 81 mm, with well developed setose oostegites, from stomach of Northern Fulmar , Greenland Sea ( ZMB 1265 View Materials ).
This specimen was described by Lichtenstein (in Mandt 1822), but not illustrated. The illustrations in Goes (1866) are the first (and only) of this specimen. The whole mouthpart bundle (including the upper lip and epistome) has been removed from the specimen and all these parts are now lost. This had probably occurred before Goes saw the specimen, as he did not illustrate any mouthparts. Other parts of the specimen are also missing, and may have been missing when it was first collected.
Description
Head much deeper than long, not extending much below insertion of antenna 2, anterodorsal margin smooth; lateral cephalic lobe small, narrowly rounded; rostrum absent; eyes not apparent. Antenna 1 peduncular article 1 short, length subequal to breadth, distal margin without significant protrusions; peduncular article 2 short, 0.35 times article 1; peduncular article 3 short, 0.13 times article 1; accessory flagellum 3+-articulate [broken], article 1 long, 5.3 times article 2; flagellum with strong two-field callynophore, 5+- articulate [broken], calceoli absent. Antenna 2 peduncle without brush setae; peduncular article 1 greatly enlarged, covering article 2 laterally; article 3 short, about 0.6 times article 4; flagellum 14+-articulate [broken]; calceoli absent.
Stoddart H. E. & Lowry J. K.
Mouthparts unknown.
Pereonites 1 to 5 dorsally smooth; pereonites 6 and 7 with weak dorsal ridge. Gnathopod 1 subchelate or parachelate [dactyl missing]; coxa vestigial; basis long, slender, length 2.3 times breadth, anterior margin smooth, with tufts of slender setae; ischium short, length 1.4 times breadth; merus with tuft of long slender setae near posterodistal corner; carpus wedge-shaped, produced anteriorly, short, length 1.5 times breadth, shorter than (0.9 times) propodus, with tuft of long slender setae on anterodistal corner and long slender setae near posterodistal corner; propodus large, subrectangular, length 1.9 times breadth, slightly tapering distally, posterior margin subtly sinusoidal, smooth, with some setae [most missing], palm transverse, margin straight, minutely serrate, posterodistal corner with at least one medial robust seta [broken, but socket visible; lateral area too damaged to assess]; dactylus unknown. Gnathopod 2 minutely chelate; coxa small, shorter than coxa 3; ischium long, length 3.6 times breadth; carpus long, length 4.3 times breadth, posterior margin straight; propodus subrectangular, long, length 3.1 times breadth; palm obtuse with convex rugose margin, with three medial and one lateral robust setae; dactylus not quite reaching corner of palm, posterior margin smooth. Pereopod 3 coxa large; merus expanded anteriorly; propodus with at least eight sets of setae on posterior margin [all broken] and one posterodistal robust seta; dactylus short, slender. Pereopod 4 coxa as deep as wide (length 1.03 times breadth), with large posteroventral lobe, anterior margin slightly round- ed; merus expanded anteriorly; propodus and dactylus missing. Pereopod 5 coxa with anterior and posterior lobes subequal; basis expanded posteriorly, posterior margin minutely crenate; merus expanded posteriorly, with curved posteri- or margin; propodus with nine groups of robust setae along anterior margin and two anterodistal robust setae; dactylus short, slender. Pereopod 6 coxa small, not lobate posteriorly; merus expand- ed posteriorly, with curved posterior margin; propodus with 10 groups of robust setae along anterior margin and two anterodistal robust setae; dactylus short, slender. Pereopod 7 basis expanded posteriorly, anterior length 1.1 times breadth, posterior margin slightly rounded, minutely crenate, posteroventral corner rounded, posteroventral margin straight; merus expanded posteriorly, with curved posterior margin; propodus with 10 groups of robust setae along anterior margin, two anterodistal robust setae and three pairs of robust setae along posterior margin; dactylus short, slender.
Oostegites present from gnathopod 2 to pereopod 5, long, setose [but with most setae missing]. Gills present from gnathopod 2 to pereopod 7.
Pleonites 1 to 3 with weak dorsal ridge; pleonite 3 with anterodorsal notch. Epimeron 1 anteroventral corner rounded, with a few slender setae. Epimeron 2 ventral margin lined with short fine setae, posteroventral corner produced into sharp spine. Epimeron 3 ventral margin lined with short fine setae, posteroventral corner broadly rounded. Urosomite 1 with anterodorsal notch. Uropod 1 peduncle with 26 dorsolateral, one apicolateral, 18 dorsomedial and two apicomedial robust setae, without plumose setae; rami subequal in length; outer ramus with 17 lateral and at least five medial robust setae; inner ramus with seven lateral and eight medial robust setae. Uropod 2 peduncle with 20 dorsolateral, two apicolateral, 11 dorsomedial and one apicomedial robust setae; rami subequal in length; outer ramus with 21 lateral and eight or nine medial robust setae; inner ramus with eight lateral and 11 medial robust setae, without constriction. Uropod 3 peduncle short, length 1.8 times breadth, without dorsolateral flange, with four apicomedial robust setae, without midmedial setae, with 10 distoventral robust setae, with lightly plumose setae on dorsomedial margin; rami paddle-like, subequal in length, inner ramus about 0.9 times outer ramus; outer ramus twoarticulate, article 2 short, article 1 with four lateral and seven medial robust setae; inner ramus with 11 medial robust setae; slender plumose setae present on both rami. Telson longer than broad, length 2.6 times breadth, deeply cleft (65%), without dorsal setae, distally tapered, without apical setae.
H. MILNE EDWARDS’ SPECIMEN ( FIGS 4-7 View FIG View FIG View FIG View FIG ) Holotype (of Lysianassa magellanica H. Milne Edwards, 1848 ), female, c. 85 mm, with well developed setose oostegites, collected by d’Orbigny from stomach of a large fish caught off Cape Horn, Drake Passage, [c. 56°S, 67°W] (MNHN-Am 3148). GoogleMaps
H. Milne Edwards (1848) gave only a brief description of a few characters and no illustrations of his new species, Lysianassa magellanica . Lucas (1857) repeated some of Milne Edwards’ description and provided a whole animal figure. Bate (1862: 66) gave a brief description of Lysianassa magellanica , and a whole animal figure taken from a tracing of Lucas’ plate which he said he had “corrected from a specimen – the original type, I believe, of M. Milne-Edwards”. Bate’s figure is somewhat stylised compared to that of Lucas, and at least two of Bate’s “corrections” are wrong. Firstly, Lucas’ figure shows antenna 2 about twice the length of antenna 1, and he states that the first antennae are “courtes, […] ne dépassent pas la partie médiane du premier anneau thoracique” but the second antennae “atteignent le quatrième anneau du thorax”; Bates’ figure shows antennae 1 and 2 subequal, and his description states “Antennae subequal”. Lilljeborg (1865a: 3, footnote 1) wrote that in Bate’s (1862) British Museum Catalogue, “neither the description nor the drawing is good”. Bate (1866: 331) responded in defence of his illustration: “[…] we feel assured that both the short description and the figure may be depended
Stoddart H. E. & Lowry J. K.
upon, except perhaps that of one of the inferior antennae, which was broken off when we saw it […] but which, if our memory is not treacherous, was ascertained by comparison with the same organ on the opposite side; or it may have been hypothetically inserted”. Unfortunately both second antennae are now missing from the specimen, but there seems no reason to doubt Lucas’ description and figure; all specimens of Eurythenes that we have seen have the second antenna about twice the length of the first antenna. Secondly, the deep anterodorsal notches on pleonite 3 and urosomite 1 are clearly shown in Lucas’ figure, but have been omitted entirely in Bate’s figure. Bate’s (1862: 66) description of uropod 3 (the “posterior pair of pleopoda”) as “having the rami styliform, clean, scarcely longer than the base” is puzzling, wrong, and probably in part a misreading of both Milne Edwards’ and Lucas’ description of the telson as “deux appendices styliformes très forts représentent le septième anneau”. Bate’s confusion about uropods and telson is compounded in his 1866 report. He did not accept Lilljeborg’s genus Eurytenes and suggested that Lilljeborg’s material would fit just as well to Anonyx , “into which the species described as Lysianassa magellanica , with its squamiform, undivided telson, cannot enter” ( Bate 1866: 332). Bate further states that “the superior antennae […] carry a secondary appendage that itself reaches beyond that of the primary branch in L. magellanica ” ( Bate 1866: 331) . This statement was presumably based on Lucas’ figure, which actually shows both left and right antenna 1, not antenna 1 with its accessory flagellum. K. H. Barnard (1932) noted some of Bate’s confusing statements and re-published Milne Edwards’ original description. The specimen has never been adequately described or figured.
When we received this specimen it was mounted with glue onto a slab of glass. It was photographed in situ and then carefully removed from the glass. It is in relatively good condition, considering its history. The right gnathopod 1 and pereopod 5 were missing from the specimen and both antennae 2 were broken off at the end of the peduncle, the flagella missing. Many margins are abraded or worn, particularly those of the pereopods and uropods, so that many setae are missing, represented only by their insertion points or broken stumps. The apparently stocky dactyls of pereopods 3 and 6 are probably an artefact; the extremity of the cuticle seems abrad- ed or maybe digested.
Description
Head much deeper than long, not extending much below insertion of antenna 2, anterodorsal
Stoddart H. E. & Lowry J. K.
margin smooth; lateral cephalic lobe small, narrowly rounded; rostrum absent; eyes not apparent. Antenna 1 short, 0.1 times body; peduncular article 1 short, length 1.2 times breadth, distal margin without significant protrusions; peduncular article 2 short, 0.22 times article 1; peduncular article 3 short, 0.19 times article 1; accessory flagellum medium length (about 0.5 times primary flagellum), 15-articulate, article 1 long, 4.3 times article 2; flagellum 30+- articulate [broken], with strong two-field callynophore, calceoli absent. Antenna 2 relative length not known [both flagella missing]; peduncle without brush setae; peduncular article 1 greatly enlarged, covering article 2 laterally; peduncular article 3 short, 0.7 times article 4; [flagellum missing].
Mouthpart bundle subquadrate. Epistome and upper lip separate; epistome produced, angular; upper lip not produced, slightly rounded. Mandible incisors symmetrical, large, with smooth slightly convex margins; lacinia mobilis a long slender distally-cuspidate robust seta; accessory setal row left row with 13, right row with 15, short, slender, robust setae; intermediate setae pappose; molar large, setose with vestigial distal triturating patch; palp attached midway, article 1 short, length 1.3 times breadth, article 2 slender, length 2.6 times breadth, once article 3, with many submarginal posterodistal A2-setae, without D2-setae, article 3 falcate, long, length 3.2 times breadth, with one proximal A3-seta, with 46 D3-setae along most of posterior margin, with eight to 10 apical and subapical E3-setae. Maxilla 1 inner plate narrow, with 15 apical pappose setae; outer plate narrow, with 11 setal-teeth in 8/3 crown arrangement, with ST1 to ST3 large, slender, smooth to weakly cuspidate, ST4 to ST6 large, slender, four- to six-cuspidate, ST7 slightly displaced from ST6, long, slender, curved, nine-cuspidate medially; STA to STD large, broad, two- to six-cuspidate; palp large, two-articulate, with three apical and four subapical long robust setae and four subapical slender setae, flag seta absent, distomedial margin smooth. Maxilla 2 inner and outer plates broad; inner plate length 0.7 times outer plate. Maxilliped inner plate large, subrectangular, with four apical nodular robust setae and seven distal robust setae on lateral face near inner margin, oblique setal row well developed, with 14 pappose setae; outer plate medium size, subovate, with 18 apical pappose setae, with three apical robust setae, medial setae small, bead-shaped, submarginal setae absent; palp large, four-articulate, article 2 very broad, length 1.5 times breadth, 1.3 times article 3, article 3 long, slen- der, length 2.6 times breadth, dactylus well developed, with 10 subapical setae, unguis present.
Pereonites 1 and 2 dorsally smooth, pereonites 3 to 7 with mid-dorsal ridge. Gnathopod 1 parachelate; coxa vestigial; basis long, slender, length 2.2 times breadth, anterior margin smooth, with many slender setae; ischium short, length 1.4 times breadth; merus posterior margin lined with long slender setae; carpus wedge-shaped, not produced anteriorly, short, length 1.4 times breadth, shorter than (0.85 times) propodus, with long slender setae near posterodistal corner; propodus large, subrectangular, length twice breadth, tapering distally, posterior margin subtly sinusoidal, smooth, with slender setae, palm transverse, margin convex (posterior to small robust seta near insertion of dactylus), smooth, posterodistal corner with one medial and one lateral robust seta; dactylus simple, without subapical spines or slender setae. Gnathopod 2 minutely subchelate; coxa small, shorter than coxa 3; ischium long, length 3.6 times breadth; carpus long, length 4.2 times breadth, posterior margin straight; propodus subrectangular, long, length 3 times breadth, palm transverse, with straight, smooth margin, with two medial and four lateral robust setae; dactylus reaching corner of palm, posterior margin smooth. Pereopod 3 coxa large; merus expanded anteriorly; propodus with many slender setae along posterior margin and one posterodistal robust seta; dactylus short, stocky. Pereopod 4 coxa about as deep as wide (length 1.1 times breadth), with large posteroventral lobe, anterior margin slightly rounded; merus expanded anteriorly; dactylus short, slender. Pereopod 5 coxa with anterior and posterior lobes subequal; basis expanded posteriorly, posterior margin minutely crenate; merus expanded posteriorly, with curved posterior margin; propodus with nine groups of robust setae along anteri- or margin and two anterodistal robust setae; dactylus short, slender. Pereopod 6 coxa small, not lobate posteriorly; basis expanded posteriorly with minutely crenate posterior margin; merus expanded posteriorly, with slightly curved posteroproximal margin and slightly curved posterodistal margin; propodus with eight groups of robust setae along anterior margin and two anterodistal robust setae; dactylus short, stocky. Pereopod 7 basis expanded posteriorly, anterior length 1.2 times breadth, posterior margin slightly rounded, minutely crenate, posteroventral corner rounded, posteroventral margin straight; merus expanded posteriorly, with sloping straight posteroproximal margin and straight posterodistal margin; propodus with nine groups of robust setae along anterior margin, two anterodistal robust setae and three pairs of robust setae along posterior margin; dactylus short, stocky.
Oostegites present from gnathopod 2 to pereopod 5, long, setose. Gills present from gnathopod 2 to pereopod 7.
Pleonites 1 to 3 with mid-dorsal ridge, increasing in size posteriorly; pleonite 3 with anterodorsal notch. Epimeron 1 anteroventral corner rounded with a few short slender setae. Epimeron 2 ventral margin lined with short fine setae; posteroventral corner produced into sharp spine. Epimeron 3 ventral margin lined with short fine setae; posteroventral corner broadly rounded. Urosomite 1 with anterodorsal notch. Uropod 1 peduncle with 35 dorsolateral, 19 dorsomedial and three apicomedial robust setae; rami subequal in length; outer ramus with 20 lateral and seven medial robust setae; inner ramus with 11 lateral and 11 medial robust setae. Uropod 2 peduncle with 21 dorsolateral, 12 dorsomedial and one apicomedial robust setae; rami subequal in length; outer ramus with 21 lateral and eight medial robust setae; inner ramus with 13 lateral and 16 medial robust setae, without constriction. Uropod 3 peduncle short, length 1.7 times breadth, without dorsolateral flange, with three apicomedial robust setae, without midmedial slender or robust setae, with at least 10 distoventral robust setae, with lightly plumose setae on dorsomedial margin; rami paddle-like, subequal in length, inner ramus about 0.9 times outer ramus; outer ramus two-articulate; both rami with a few small robust setae on distolateral margin, slender plumose setae present on both rami; [most of the distal setae are missing, only insertion points remain]. Telson longer than broad, length twice breadth, deeply cleft (80%), without dorsal setae, distally tapered, tips of both lobes damaged.
Remarks
The mid-dorsal ridge is distinct but low from pereonite 3 to pleonite 1; from pleonite 2 to urosomite 1 the ridge is strong enough to be described as a slight carina, strongest on pleonite 3. On all segments the ridge is slightly more pronounced posteriorly.
LILLJEBORG’ S SPECIMEN ( FIGS 8-10 View FIG View FIG View FIG )
Material of Lilljeborg 1865a (as Eurytenes magellanicus ), immature female, 55 mm, with short non-setose oostegites, Bear Island, Arctic Ocean, [c. 74°N, 19°E], from stomach of Scymnus borealis (MNHN-Am 4479).
The label with this specimen reads: “ Eurythenes magellanicus . Île Boeren. Mer Glaciale. Lilljeborg. 133-67”. Lilljeborg (1865a: 3, 4), recorded three specimens of Lysianassa magellanica brought to the Zoological Museum by Mr Th. M. Fries, who “had received them from a shipper in Hammerfest, who had found them in the belly of an ‘Haaskierding’ […], on the banks by Beeren Island”. Bate (1867), writing in the Zoological Record for 1866, commented that he had received a letter from M. Alphonse Milne-Edwards saying that “J’ai reçu dernièrement de M. le Prof. Lilljeborg un exemplaire de son Eurytenes magellanicus […]”. With the agreement of Drs Saint Laurent and Defaye of MNHN, we believe the specimen we illustrate here is the one sent to Paris by Lilljeborg, and one of the three specimens he used when establishing the genus Eurytenes .
Description
Head much deeper than long, not extending much below insertion of antenna 2, anterodorsal margin smooth; lateral cephalic lobe small, narrowly rounded; rostrum absent; eyes not apparent. Antenna 1 short; peduncular article 1 short, length equals breadth, distal margin with small midmedial swelling; peduncular article 2 short, 0.37 times article 1; peduncular article 3 short, 0.19 times article 1; accessory flagellum 10+- articulate [broken], article 1 long, 4.4 times article 2; flagellum 10+-articulate [broken], with strong two-field callynophore, flagellum without robust setae, calceoli absent. Antenna 2 length not known; peduncle with weak brush setae [setae missing but insertion points visible]; peduncular article 1 enlarged, covering article 2 laterally; peduncular article 3 short, subequal to article 4; flagellum missing.
Mouthpart bundle subquadrate. Epistome and upper lip separate, epistome produced, rounded; upper lip not produced, slightly rounded. Mandible incisors large, with slightly convex margins; lacinia mobilis a long slender distallycuspidate robust seta; accessory setal row left and right with nine short, slender, robust setae; intermediate setae pappose; molar large, setose with vestigial distal triturating patch; palp attached midway; article 1 short, length 1.4 times breadth; article 2 slender, length 2.9 times breadth, 1.1 times article 3, with many submarginal posterodistal A2-setae, without D2-setae; article 3 falcate, long, length 3.1 times breadth, with one proximal A3-seta, with about 50 D3-setae along most of posterior margin, with two apical and six subapical E3-setae. Maxilla 1 inner plate narrow, with eight apical and subapical pappose setae; outer plate narrow, with 11 setal-teeth in 8/3 crown arrangement; ST1 to ST6 large, slender, weakly cuspidate, ST7 slightly displaced from ST6, elongate, slender, curved, eight-cuspidate medially; STA large, broad, two-cuspidate; STB and STC large, broad, five-cuspidate; STD large, broad, six-cuspidate; palp large, two-articulate, with five apical and one apicolateral long robust setae, with three subapical setae, flag setae absent, distomedial margin smooth. Maxilla 2 inner and outer plates broad; inner plate length 0.6 times outer plate. Maxilliped inner plate large, subrectangular, with three apical nodular robust setae; oblique setal row well developed, with at least 16 pappose setae; outer plate medium sized, subovate, with 18 apical pappose setae and three apical robust setae, medial setae small, bead-shaped, submarginal setae vestigial; palp large, four-articulate, article 2 very broad, length 1.8 times breadth (when completely flattened), 1.4 times article 3, article 3 long, slender, length 2.8 times breadth, dactylus well developed, with seven subapical setae, unguis present.
Pereonites 1 to 4 dorsally smooth; pereonites 5 to 7 with slight mid-dorsal ridge. Gnathopod 1 parachelate; coxa vestigial; basis long, slender, length 2.3 times breadth, anterior margin smooth, with tufts of slender setae; ischium short, length 1.3 times breadth; merus posterior margin lined with long slender setae; carpus wedge-shaped, produced anteriorly, short, length 1.5 times breadth, shorter than (0.79 times) propodus, with long slender setae near posterodistal corner; propodus large, subrectangular, length twice breadth, slightly tapering distally, posterior margin subtly sinusoidal, smooth, with tufts of slender setae, palm transverse, margin straight, minutely serrate, posterodistal corner with one medial and one lateral robust seta; dactylus simple, without subapical spines or slen- der setae, reaching beyond palm. Gnathopod 2 minutely subchelate; coxa large, shorter than coxa 3; ischium long, length 3.4 times breadth; carpus long, length 4.3 times breadth, posterior margin straight; propodus subrectangular, long, length 3.1 times breadth, palm slightly obtuse, with convex, rugose margin, with two medial and two lateral robust setae; dactylus reaching corner of palm, posterior margin smooth. Pereopod 3 coxa large; merus expanded anteriorly; propodus with seven tufts of slender setae along posterior margin [some setae missing, insertion points visible] and one posterodistal robust seta; dactylus short, slen- der. Pereopod 4 coxa as deep as wide (length 1.02 times breadth), with large posteroventral lobe, anterior margin slightly rounded; merus expanded anteriorly; propodus with five tufts of slender setae along posterior margin and one posterodistal robust seta; dactylus short, slender. Pereopod 5 coxa anterior and posterior lobes subequal; basis expanded posteriorly, posterior margin minutely crenate; merus expanded posteriorly, with curved posterior margin; propodus with eight groups of two or three short robust setae along anterior margin and two anterodistal robust setae; dactylus short, slender. Pereopod 6 coxa small, not lobate posteriorly; basis expanded posteriorly, posterior margin minutely crenate; merus expanded posteriorly, with curved posteri- or margin; propodus with eight groups of three or four short robust setae along anterior margin and two anterodistal robust setae; dactylus short, slender. Pereopod 7 basis expanded posteriorly, anterior length 1.1 times breadth, posterior margin slightly rounded, minutely crenate, posteroventral corner rounded, posteroventral margin straight; merus expanded posteriorly, with curved posterior margin; propodus with eight groups of two to five short robust setae along anterior margin and two anterodistal robust setae, with three pairs of robust setae along posterior margin; dactylus short, slender.
Oostegites present from gnathopod 2 to pereopod 5, short, non-setose. Gills present from gnathopod 2 to pereopod 7.
Pleonites 1 to 3 with slight but distinct middorsal ridge. Pleonite 3 with a deep anterodorsal notch. Epimeron 1 anteroventral corner rounded with a few short slender setae. Epimeron 2 ventral margin lined with short fine setae; posteroventral corner produced into sharp spine. Epimeron 3 ventral margin lined with short fine setae; posteroventral corner broadly rounded. Urosomite 1 with anterodorsal notch. Uropod 1 peduncle with 33 dorsolateral, at least 18 dorsomedial and one apicomedial robust setae; outer ramus with at least eight lateral and at least three medial robust setae; inner ramus with five lateral and at least eight medial robust setae [both rami missing distal portion]. Uropod 2 peduncle with 17 dorsolateral and about 10 dorsomedial robust setae; outer ramus with about 15 lateral and seven medial robust setae; inner ramus with at least two lateral and six medial robust setae [both rami missing distal portion]. Uropod 3 peduncle short, length 1.9 times breadth, without dorsolateral flange, with three apicomedial robust setae, without midmedial setae, with four distoventral robust setae, with lightly plumose setae on dorsomedial margin; rami paddle-like, subequal in length, inner ramus about 0.9 times outer ramus; outer ramus two-articulate, article 2 short, article 1 with six lateral and two medial robust setae; inner ramus with one lateral and two medial robust setae, slender plumose setae present on both rami. Telson longer than broad, length 2.3 times breadth, deeply cleft (79%), without dorsal setae, distally tapered, distal margins without setae [but both slightly damaged].
COMPARISON OF THE THREE SPECIMENS
These three specimens differ in many points, most of which are very slight and some of which may be due to difference in overall size. Differences in numbers of setae – for example in the accessory setal row of the mandible, on the inner plate of maxilla 1, on the inner and outer plates of the maxilliped, on the pereopods and on the uropods – are most likely size-related. So too is the development of the dorsal ridge. There are other small differences which do not appear to be size-related – for example the length to breadth of antenna 1 peduncular article 1 in the Milne Edwards specimen is slightly greater than in the other two specimens; the lateral cephalic lobe is slightly more rounded in the Lichtenstein specimen; the epistome is rounded in the Lilljeborg specimen but slightly angular in the Milne Edwards specimen; coxa 4 of the Milne Edwards specimen is slightly longer (relative to its breadth) than in the other two specimens.
The gnathopodal palms vary more than do other characters. Unfortunately neither of the gnathopod 1 dactyli is present in the Lichtenstein specimen, so it is not known if the dactyl overhung the palm or not. The size and shape of the gnathopod 1 palm in the Lichtenstein specimen is very similar to that of the Lilljeborg specimen (small, straight), but that of the Milne Edwards specimen is quite different (very small, convex). It is similar to that illustrated by Bowman & Manning (1972) for material from Guadalupe Island, eastern North Pacific Ocean. No other Eurythenes material that we have seen has this type of gnathopod 1, but it may be similar to that of the material recorded by Thurston et al. (2002) from the Atacama Trench, off northern Chile. The geographically closest specimens of the material recorded below – the large female from Macquarie Island and specimens from French Polynesia – have a small but straight, transverse palm on gnathopod 1, similar to that of the Lichtenstein and Lilljeborg specimens. The gnathopod 2 palm is different in all three specimens: obtuse in the Lichtenstein specimen, slightly obtuse in the Lilljeborg specimen and transverse in the Milne Edwards specimen. There are five illustrations of E. gryllus gnathopods in the literature: Lilljeborg (1865a), Goes (1866), Sars (1891), Bowman & Manning (1972) and Rauschert (1985). They also show a diversity of size and shape in the propodus/palm of both gnathopods. There is no clear pattern to this variation.
It is possible that “ Lysianassa magellanica ” represents a valid species, but until additional material of this form is recognised the name remains a junior synonym.
OTHER MATERIAL EXAMINED. — Australia. E of Flynn Reef, Queensland, RV Sunbird , SEAS, stn QLD-930, 16°37.81’S, 146°23.08’E, 1000 m, baited trap, bottom temperature 5.2°C, 6-7.VI.1993, J. K. Lowry, P. Freewater & W. Vader, 1 immature with short non-setose oostegites, 48 mm GoogleMaps ; 2 immature without calceoli, 48 and 51 mm; 3 juveniles 31, 36 and 38 mm ( AM P61826). — Same data, SEAS, stn QLD-931, 20 immature and juvenile specimens 16- 30 mm ( AM P50237). — Same data, SEAS, stn QLD-932, 3 juveniles 16-17 mm ( AM P57605) ; 10 juveniles 16-25 mm ( AM P50242). — Same data, SEAS, stn QLD-948, 23 immature and juvenile specimens 16-48 mm ( AM P50290). — Same data, SEAS, stn QLD-949, 4 juveniles 17-24 mm ( AM P50296). — Same data, SEAS, stn QLD-950, 7 immature specimens 22-28 mm ( AM P61827). — E of Cairns, Queensland, RV Franklin , CIDARIS-1, stn 35.4, 16°54.4’S, 147°14.35’E, 1473-1590 m, beam trawl, 14.V.1986, M. Pichon, P. W. Arnold & R. A. Birtles, 1 without calceoli, 50 mm GoogleMaps ; 1 with calceoli, 65 mm ( QM 2684 ) .
Western South Pacific Ocean (material recorded as Eurythenes cf. gryllus in Lowry & Stoddart 1994): New Caledonia. RV Vauban, stn CA-1, 20°44’S, 166°27’E, 1000 m, baited fish trap, A. Intes, 19.II.1977, 2 with calceoli, 52 and 64 mm GoogleMaps ; 1 immature specimen 46 mm ( AM P28855) .
Loyalty Islands. W of Lifou, 20°48.12’S, 166°53’E, 1620-1630 m, beam trawl, 1.V.1987, BIOGEOCAL, stn CP 317, 1 without calceoli, 50 mm (MNHN- Am 4403).
French Polynesia. Fangataufa Atoll , Tuamotu Archipelago, 22°16’S, 138°42’W, 900-1000 m, baited trap, J. Poupin, 1 with setose oostegites, 90 mm GoogleMaps ; 2 with calceoli, 62 and 78 mm ( AM P42139). — Just off Rapa, Austral Isles, RV Marara, SMCB , stn FRP-54, 27°35.3’S, 144°15.5’W, 870 m, baited trap, J. K. Lowry & J. Poupin, 17-18.VIII.1991, 1 without calceoli, 50 mm ( AM P42138) GoogleMaps .
Southern Ocean: Macquarie Island. Bauer Bay, [c. 54°33’S 158°58’E], regurgitated by chick of Lightmantled Sooty Albatross, 3.I.1964, K. G. Simpson, Ser. No. M/64/Ar/1, 1 mature with setose oostegites, 120 mm ( AM P62147).
REMARKS
None of the specimens from trap samples on the east coast of Australia were mature. The largest female (48 mm) was only beginning to develop oostegites. However, the deepest trapset used in the project in which these specimens were caught was at 1000 m depth. This is probably the very top of the species’ bathymetric range in this area. Traps set at shallower depths in the same locality did not collect any Eurythenes .
The very large (120 mm) female from Macquarie Island has a well developed oostegite on gnathopod 1. It is the only specimen in this collection to have five pairs of oostegites. The next largest (90 mm) female, from French Polynesia, has four pairs of long, well developed, setose oostegites but no trace of an oostegite on gnathopod 1.
Eurythenes obesus ( Chevreux, 1905) ( Figs 12-15 View FIG View FIG View FIG View FIG )
Katius obesus Chevreux, 1905: 1 View in CoL , figs 1-3; 1935: 63. — Stebbing 1906: 721. — Tattersall 1906: 29; 1909: 210. — Stephensen 1912a: 89; 1912b: 614; 1913: 123; 1915: 37; 1925: 126; 1933: 12, 13, 18, figs 6, 7. — Shoemaker 1920: 8 E; 1956: 177. — Schellenberg 1926: 217, fig. 26d; 1927: 681, fig. 72; 1931: 16. — K. H. Barnard 1932: 56, fig. 21, pl. 1 fig. 1. — Belloc 1960: 7. — Costello et al. 1989: 32.
Eurythenes obesus View in CoL – Schellenberg 1955: 183, 192. — Shoemaker 1956: 178. — J. L. Barnard 1958: 92; 1961: 38, fig. 8. — Birstein & Vinogradov 1960: 184; 1962: 36; 1964: 163; 1970: 420 (table 1). — Hurley 1963: 59. — Brusca 1967: 384. — Imber 1973: 652. — Bellan-Santini & Ledoyer 1974: 681, pl. 25. — Griffiths 1975: 145; 1976: 56, 100. — Lowry & Bullock 1976: 89. — Ortiz 1979: 19. — Andres 1983: 186; 1987: 96, 97. — Umezu 1984: 128. — Hopkins 1985: 202. — Costello et al. 1989: 32. — Barnard & Karaman 1991: 486. — Boudrias 1991: 13. — Palerud & Vader 1991: 35. — De Broyer & Jazdzewski 1993: 68. — Kaufmann 1994: 54. — Piatkowski et al. 1994: 19 (table 1). — Thurston & Bett 1995: 201. — Johnson et al. 2001: 198 (table 3). — Cherel et al. 2002a: 268 (table 3); 2002b: 288 (table 3), 292 (table 4).
Eurythenes gryllus View in CoL – K. H. Barnard 1940: 440, 515 (list). — Springer & Bullis 1956: 6.
Eurythenes gryllus View in CoL – Stephensen 1932: 356 (in part).
TYPE MATERIAL. — Chevreux’s type specimen of E. obesus , a 12 mm? taken near the Azores, eastern North Atlantic Ocean, is now lost, according to Dr Christian Carpine of the Musée océanographique, Monaco. Because of the increasing complexity of species concepts in the genus Eurythenes it is important to fix the identity of E. obesus and establish the state of characters which are now considered important for delimiting species. Dr M. Thurston of the Southampton Oceanography Centre has made available a mature female specimen, taken near the Cape Verde Islands, eastern North Atlantic Ocean, which we here designate as the neotype for E. obesus . This is the specimen recorded and studied by Thurston & Bett (1995).
Neotype: 48 mm, with setose oostegites and hatchlings ( BMNH 2003.1059 ), RRS Discovery , stn 9541#30, NE of Cape Verde Islands, eastern North Atlantic Ocean, 20°1.8’N, 21°19.8’W- 20°1.3’N, 21°20.0’W, 995-1500 m over bottom depth 3800-3850 m, rectangular midwater trawl RMT 8, 1921-2321 hours, 22.IV.1977. GoogleMaps
ADDITIONAL MATERIAL EXAMINED. — Eastern North Atlantic Ocean. RRS Discovery, stn 8508#13, 44°4.3’N, 12°44.7’W- 44°4.8’N, 12°47.9’W, 590- 610 m, rectangular midwater trawl RMT 8, 0551- 0651 hours, 14.IV.1974, with calceoli, 18 mm ( BMNH 2003.1060).
Australia. Western Front, Eddy J, E of Sydney, NSW, 33°33’S, 152°34’E, 271 m, 1835-1935 hours, 2.X.1979, stn SP 10-79, B. Jeffries & R. Fritz, 1 with oostegite buds, 25 mm ( AM P31001). — SE of Montague Island, NSW, FRV Kapala , stn K77-19-01, 36°24’S, 150°18’E- 36°30’S, 150°17’E, 128 m over bottom depth 146 m, midwater trawl, 0900- 1125 hours, 1.XI.1977, 1 with setose oostegites, 80 mm ( AM P30498). — S of Point Hicks, Victoria, RV Franklin, stn SLOPE 38, 38°39.23’S, 149°15.00’E, 1500 m over bottom depth 2900-3200 m, 8 m rectangular midwater trawl, 24.VII.1986, M. F. Gomon et al., 1 with oostegite buds, 24 mm ( NMV J 52806 View Materials ). — Off Freycinet Peninsula, Tasmania, RV Franklin, stn SLOPE 42, 42°03.30’S, 148°57.90’E, 900-1000 m over bottom depth 2100-2300 m, 8 m rectangular midwater trawl, 26.VII.1986, M. F. Gomon & L. Hamond, 1 juvenile 16 mm ( NMV J 52807 View Materials ). — Off Freycinet Peninsula, Tasmania, RV Franklin, stn SLOPE 44, 42°13.00’S, 148°46.10’E, 500-1000 m over bottom depth 1700 m, 8 m rectangular midwater trawl, 27.VII.1986, M. F. Gomon et al., 1 juvenile 22 mm ( NMV J 52808 View Materials ). — 61 km ENE of Cape Tourville, Tasmania, RV Franklin, stn SLOPE 75, 41°58.49’S, 149°04.41’E, 3 m Isaacs-Kidd midwater trawl over bottom depth 1685-2524 m, 28.X.1988, G. C. B. Poore et al., 1 ovigerous 34 mm GoogleMaps ; 1 with oostegite buds, 24 mm ( NMV J 52805 View Materials ) .
Macquarie Island. Southern Ocean, [c. 54°35’S, 158°55’E], regurgitated by Northern Giant Petrel chick, Macronectes halli, 1970 , G. W. Johnstone, 1 with setose oostegites, 53 mm ( AM P20352).
Southern Ocean. 55°01.9’S, 99°53.7’E, to 1000 m, stn 125, rectangular midwater trawl, 0815-0900 hours, 15.III.1981, J. Kirkwood, 1 with setose oostegites, 50 mm ( AM P62437). — 64°18.3’S, 78°43.9’E, to 980 m, stn 80, rectangular midwater trawl, 1742- 1812 hours, 18.II.1981, J. Kirkwood, 1 juvenile, 16 mm ( AM P62438).
DIAGNOSIS. — Anterodorsal margin of head smooth. Gnathopod 1 parachelate; basis length 2 to 2.5 times breadth; propodus slightly tapering distally. Pereopods 3 to 7 dactyli long. Pereopod 4 coxa as deep as wide. Pereopod 7 basis length of anterior margin subequal to breadth; posteroventral margin rounded. Pleonite 3 with anterodorsal notch. Epimeron 3 posteroventral corner subquadrate. Urosomite 1 not dorsodistally produced over urosomite 2. Uropod 3 peduncle, medial face with robust setae.
DESCRIPTION
Based on neotype female. Head much deeper than long, not extending much below insertion of antenna 2, anterodorsal margin smooth; lateral cephalic lobe small, subacute; rostrum absent; eyes not apparent. Antenna 1 short; peduncular article 1 short, length 1.1 times breadth, distal margin without midmedial tooth; peduncular article 2 short, 0.2 times article 1; peduncular article 3 short; accessory flagellum 11-articulate, article 1 long, 3.3 times article 2; flagellum 25- articulate, with strong two-field callynophore, calceoli absent. Antenna 2 medium length, 1.6 times antenna 1; peduncle without brush setae; peduncular article 1 enlarged, covering article 2 laterally; peduncular article 3 short, 1.3 times article 4; flagellum 46-articulate, calceoli absent.
Mouthpart bundle subquadrate. Epistome and upper lip separate, epistome produced, rounded; upper lip not produced, slightly rounded. Mandible incisors large, symmetrical, with slightly convex smooth margins; lacinia mobilis a long slender distally-cuspidate robust seta; accessory setal row left and right with 10 short, slender, robust setae; intermediate setae pappose; molar large, setose, with vestigial distal triturating patch; palp attached midway; article 1 short, length 1.4 times breadth; article 2 slender, length 3.2 times breadth, 1.1 times article 3, with many submarginal posterodistal A2-setae, without D2- setae; article 3 falcate, long, length 3.2 times breadth, with one proximal A3-seta, with about 44 D3-setae along most of posterior margin, with two apical and four subapical E3-setae. Maxilla 1 inner plate narrow, with eight apical and subapical pappose setae; outer plate narrow, with 11 setal-teeth in 8/3 crown arrangement; ST1 to ST3 large, slender, three- to four-cuspidate, ST4 to ST6 large, slender, four-cuspidate, ST7 slightly displaced from ST6, elongate, slender, ninecuspidate medially; STA large, broad, two-cuspidate; STB to STD large, broad, five- to eight-cuspidate; palp large, two-articulate, with five apical and three apicolateral short robust setae, and four subapical slender seta, flag seta absent, distomedial margin smooth. Maxilla 2 inner and outer plates broad; inner plate length 0.7 times outer plate. Maxilliped inner plate large, subrectangular, with three apical nodular robust setae and three robust setae on lateral face; oblique setal row well developed, with at least 30 pappose setae; outer plate medium size, subovate, with 10 apical pappose setae and three apical robust setae, medial setae small, bead-shaped, submarginal setae vestigial; palp large, four-articulate, article 2 very broad, length 1.8 times breadth, 1.3 times article 3, article 3 long, slen- der, length 2.7 times breadth, dactylus well developed, with five subapical setae, unguis present.
Pereonites 1 and 2 dorsally smooth; pereonites 3 to 7 with sharp dorsal ridge. Gnathopod 1 parachelate; coxa vestigial; basis long, slender, length 2.3 times breadth, anterior margin smooth, with tufts of slender setae; ischium short, length 1.4 times breadth; merus posterior margin lined with long slender setae; carpus wedge-shaped, slightly produced anteriorly, short, length 1.4 times breadth, shorter than (0.7 times) propodus, with long slender setae near posterodistal corner; propodus large, subrectangular, length 2.1 times breadth, slightly tapering distally, posterior margin subtly sinusoidal, smooth, with tufts of slender setae, palm transverse, margin slightly convex, minutely serrate, posterodistal corner with one medial and one lateral robust seta; dactylus simple, without subapical spines or slender setae, reaching beyond corner of palm. Gnathopod 2 minutely subchelate; coxa small, shorter than coxa 3; ischium long, length 3.9 times breadth; carpus very long, length 5.2 times breadth, posterior margin straight; propodus subrectangular, long, length 4.0 times breadth, palm transverse, with convex, minutely serrate margin, with one medial and one lateral robust seta; dactylus not reaching corner of palm, posterior margin smooth. Pereopod 3 coxa large; merus weakly expanded anteriorly; propodus with four groups of slender setae along posterior margin and one posterodistal robust seta; dactylus long, slender. Pereopod 4 coxa deeper than wide (length 1.1 times breadth), with large posteroventral lobe, anterior margin curved; merus weakly expanded anteriorly; propodus with four groups of slender setae along posterior margin and one posterodistal robust seta; dactylus long, slender. Pereopod 5 coxa with anterior and posterior lobes subequal; basis expanded posteriorly, posterior margin smooth; merus expand- ed posteriorly, with sloping posteroproximal margin and straight posterodistal margin; propodus with five short robust setae along anterior margin and two anterodistal robust setae; dactylus long, slender. Pereopod 6 coxa small, not lobate posteriorly; basis expanded posteriorly, posterior margin minutely crenate; merus expanded posteriorly, with curved posterior margin; propodus with five groups of short robust setae along anterior margin and two anterodistal robust setae; dactylus long, slender. Pereopod 7 basis expanded posteriorly, anterior length subequal to breadth, posterior margin slightly rounded, minutely crenate, posteroventral corner subquadrate, posteroventral margin straight; merus expanded posteriorly, with curved posterior margin; propodus with five groups of four to eight short robust setae along anterior margin, two anterodistal robust setae and two pairs of robust setae along posterior margin; dactylus long, slender.
Oostegites present from gnathopod 2 to pereopod 5, long, setose. Gills present from gnathopod 2 to pereopod 7.
Pleonites 1 to 3 with sharp dorsal ridge. Pleonite 3 with an anterodorsal notch. Epimeron 1 anteroventral corner subquadrate with a few short slender setae. Epimeron 2 ventral margin lined with short fine setae; posteroventral corner produced into sharp spine. Epimeron 3 ventral margin lined with short fine setae; posteroventral corner subquadrate. Urosomite 1 with anterodorsal notch, with a dorsal ridge. Uropod 1 peduncle with 34 dorsolateral, 12 dorsomedial and one apicomedial robust setae, without plumose setae; rami subequal in length; outer ramus with 14 lateral and five medial robust setae; inner ramus with seven lateral and eight medial robust setae. Uropod 2 peduncle with 23 dorsolateral, two apicolateral, seven dorsomedial and one apicomedial robust setae; rami subequal in length; outer ramus with 16 lateral and five medial robust setae; inner ramus with nine lateral and nine medial robust setae. Uropod 3 peduncle short, length 1.4 times breadth, without dorsolateral flange, with four apicomedial, three midmedial and six distoventral robust setae, with lightly plumose setae on dorsomedial margin; rami paddle-like, subequal in length, inner ramus about 0.9 times outer ramus; outer ramus two-articulate, article 2 short, article 1 with three lateral and 11 distomedial robust setae; inner ramus with 11 distomedial robust setae; slender plumose setae present on both rami. Telson longer than broad, length 2.4 times breadth, deeply cleft (83%), without dorsal robust setae, distally tapered, without apical setae.
REMARKS
The 80 mm female from SE Australia ( AM P30498) is the largest specimen of E. obesus so far recorded. It has long, setose oostegites but was not carrying eggs or young. The oostegites were flat against the sternum; the gonopore is large and the encircling rim of tissue is soft; this female may have been close to producing a brood. There is no trace of an oostegite on gnathopod 1. This specimen is also notable for where it was captured: at 128 m depth over a bottom depth of 146 m. Although E. gryllus has occasionally been taken at relatively shallow depths such captures have been made over deep water (e.g., three records at 200 m or less recorded by Andres [1983], but over bottom depths of 3000- 4000 m). The previously shallowest recorded bottom depth over which E. obesus has been captured is 950 m ( Andres 1983).
Very little is known of the life habits of E. obesus . It has never been taken in baited traps; has frequently been taken in midwater trawls; and has once been recorded as burrowing into a salp ( Stephensen 1915), once as having coelenterate remains in the stomach ( Hopkins 1985), once with possibly siliceous sponge spicules in the stomach ( Brusca 1967), and once as attacking fish taken in midwater trawls ( Thurston & Bett 1995).
AINLEY D. G., FRASER W. R., SULLIVAN C. W., TORRES J. J., HOPKINS T. L. & SMITH W. O. 1986. - Antarctic mesopelagic micronekton: evidence from seabirds that pack ice affects community structure. Science 232: 847 - 849.
ANDRES H. G. 1979. - Gammaridea (Amphipoda, Crustacea) der Antarktis-Expedition 1975 / 76: Auswertung der Dauerstation sudlich von Elephant Island. Meeresforschung 27 (2): 88 - 102.
ANDRES H. G. 1983. - Die Gammaridea (Crustacea: Amphipoda) der Deutschen Antarktis-Expeditionen 1975 / 76 und 1977 / 78. 3. Lysianassidae. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut 80: 183 - 220.
ANDRES H. G. 1987. - Die Gammaridea der 76. Reise von FFS Walther Herwig mit Beschreibung von Parachevreuxiella lobata gen. n. und sp. n. (Crustacea: Amphipoda). Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut 84: 95 - 103.
ARNAUD P. M. 1974. - Contribution a la bionomie marine benthique des regions antarctiques et subantarctiques. Tethys 6 (3): 465 - 656.
BALDWIN R. J. & SMITH K. L. 1987. - Temporal variation in the catch rate, length, color and sex of the necrophagous amphipod, Eurythenes gryllus, from the central and eastern North Pacific. Deep- Sea Research Part A: Oceanographic Research Papers 34 (3): 425 - 439.
BARNARD K. H. 1932. - Amphipoda. Discovery Reports 5: 1 - 326, pl. 1.
BARNARD K. H. 1937. - Amphipoda. Scientific Reports of the John Murray Expedition 4 (6): 131 - 201.
BARNARD K. H. 1940. - Contributions to the crustacean fauna of South Africa. 12. Further additions to the Tanaidacea, Isopoda, and Amphipoda, together with keys for the identification of the hitherto recorded marine and fresh-water species. Annals of the South African Museum 32: 381 - 543.
BARNARD J. L. 1958. - Index to the families, genera, and species of the gammaridean Amphipoda (Crustacea). Allan Hancock Foundation Publications, Occasional Paper 19: 1 - 145.
BARNARD J. L. 1961. - Gammaridean Amphipoda from depths of 400 to 6000 meters. Galathea Report 5: 23 - 128.
BARNARD J. L. & KARAMAN G. S. 1991. - The families and genera of marine gammaridean Amphipoda (except marine gammaroids). Records of the Australian Museum, Supplement 13 (1 & 2): 1 - 866.
BATE C. S. 1862. - Catalogue of the Specimens of Amphipodous Crustacea in the Collection of the British Museum. Trustees, British Museum, London, 399 p.
BATE C. S. 1866. - Crustacea. The Record of Zoological Literature, 1865 2: 306 - 366.
BATE C. S. 1867. - Crustacea. The Record of Zoological Literature, 1866 3: 216 - 250.
BECK J. R. 1969. - Food, moult and age of first breeding in the Cape Pigeon, Daption capensis Linnaeus. British Antarctic Survey Bulletin 21: 33 - 44.
BELLOC G. 1960. - Catalogue des types d'amphipodes du Musee oceanographique de Monaco. Bulletin de l'Institut oceanographique, Monaco 57 (1170): 1 - 28.
BETHUNE A. 1869. - Remarks on the distribution of animal life in the depths of the sea. Annals and Magazine of Natural History ser. 4, 3: 423 - 441 (translation of Sars 1869).
BIRSTEIN J. A. & VINOGRADOV M. E. 1955. - [Pelagic gammarideans (Amphipoda-Gammaridea) from the Kurile-Kamchatka Trench]. Akademiya Nauk SSSR, Trudy Instituta Okeanologii 12: 210 - 287 (in Russian).
BIRSTEIN J. A. & VINOGRADOV M. E. 1958. - [Pelagic gammarids (Amphipoda, Gammaridea) from the northwestern part of the Pacific Ocean]. Akademiya Nauk SSSR, Trudy Instituta Okeanologii 27: 219 - 257 (in Russian).
BIRSTEIN J. A. & VINOGRADOV M. E. 1960. - [Pelagic gammarids from the tropical Pacific Ocean]. Akademiya Nauk SSSR, Trudy Instituta Okeanologii 34: 165 - 241 (in Russian).
BIRSTEIN J. A. & VINOGRADOV M. E. 1962. - [Pelagic Gammaridea (Amphipoda, Gammaridea) collected by the Soviet Antarctic Expedition on the M / V Ob, south of 40 ° S], in ANDRIYASHEV A. P. & USHAKOV P. V. (eds), [Biological reports of the Soviet Antarctic Expedition (1955 - 1958)]. Akademiya Nauk SSSR, Issledovaniya Fauny Morei 1 (10): 33 - 56 (in Russian).
BIRSTEIN J. A. & VINOGRADOV M. E. 1964. - [Pelagic gammarid amphipods of the northern part of the Indian Ocean]. Akademiya Nauk SSSR, Trudy Instituta Okeanologii 65: 152 - 195 (in Russian).
BIRSTEIN J. A. & VINOGRADOV M. E. 1970. - [On the fauna of pelagic gammaridean amphipods from the Kurile-Kamchatka region of the Pacific Ocean]. Akademiya Nauk SSSR, Trudy Instituta Okeanologii 86: 401 - 419 (in Russian).
BLUHM B. A., BREY T., KLAGES M. & ARNTZ W. E. 2001. - Occurrence of the autofluorescent pigment, lipofuscin, in polar crustaceans and its potential as an age marker. Polar Biology 24 (9): 642 - 649.
BOECK A. 1871. - Crustacea Amphipoda borealia et arctica. Forhandlinger i Videnskabs-Selskabet i Christiania 1870: 83 - 280, i-viii [index].
BOECK A. 1872. - De Skandinaviske og Arktiske Amphipoder. Part 1. A. W. Brogger, Christiania, 160 p., 7 pls.
BOUDRIAS M. A. 1991. - Methods for the study of amphipod swimming: behavior, morphology, and fluid dynamics, in WATLING G. (ed.), Proceedings of the VIIth International Colloquium on Amphipoda held in Walpole, Maine, USA, 14 - 16 September 1990. Hydrobiologia 223: 11 - 25.
BOWMAN T. E. & MANNING R. B. 1972. - Two arctic bathyal crustaceans: the shrimp Bythocaris cryonesus new species, and the amphipod Eurythenes gryllus, with in situ photographs from Ice Island T- 3. Crustaceana 23 (2): 187 - 201, pl. 1.
BRITTON J. C. & MORTON B. 1993. - Are there obligate marine scavengers?, in MORTON B. (ed.), The Marine Biology of the South China Sea. Proceedings of the First International Conference on the Marine Biology of Hong Kong and the South China Sea. Hong Kong, 28 October- 3 November 1990. Hong Kong University Press, Hong Kong: 357 - 391.
BRITTON J. C. & MORTON B. 1994. - Marine carrion and scavengers. Oceanography and Marine Biology Annual Review 32: 369 - 434.
BRUGGEN E. 1907. - Zoologische Ergebnisse der russischen Expedition nach Spitzbergen. Amphipoda. Annuaire du Musee zoologique de l'Academie imperiale des Sciences de St-Petersbourg 11: 214 - 245.
BRUSCA G. J. 1967. - The ecology of pelagic Amphipoda. I. Species accounts, vertical zonation and migration of Amphipoda from the waters off Southern California. Pacific Science 21 (3): 382 - 393.
BUCKLIN A., WILSON R. R. & SMITH L. S. 1987. - Genetic differentiation of seamount and basin populations of the deep-sea amphipod Eurythenes gryllus. Deep-Sea Research Part A: Oceanographic Research Papers 34 (11): 1795 - 1810.
BUHRING S. I. & CHRISTIANSEN B. 2001. - Lipids in selected abyssal benthopelagic animals: links to the epipelagic zone? Progress in Oceanography 50: 369 - 382
CHARMASSON S. S. & CALMET D. P. 1987. - Distribution of scavenging Lysianassidae amphipods Eurythenes gryllus in the northeast Atlantic: comparison with studies held in the Pacific. Deep- Sea Research Part A: Oceanographic Research Papers 34 (9): 1509 - 1523.
CHARMASSON S. S. & CALMET D. P. 1989. - Distribution of Lysianassidae amphipods Eurythenes gryllus in the North-East Atlantic, in NYFFELER F. (ed.), Interim Oceanographic Description of the North-East Atlantic Site For the Disposal of Low-Level Radioactive Waste. Vol. 3. OECD, Paris: 159 - 162.
CHEREL Y., BOCHER P., DE BROYER C. & HOBSON K. A. 2002 a. - Food and feeding ecology of the sympatric thin-billed Pachyptila belcheri and Antarctic P. desolata prions at Iles Kerguelen, Southern Indian Ocean. Marine Ecology Progress Series 228: 263 - 281.
CHEVREUX E. 1889. - Quatrieme campagne de l' Hirondelle, 1888. Sur la presence d'une rare et interessante espece d'amphipode, Eurythenes gryllus Mandt, dans les eaux profondes de l'ocean, au voisinage des Acores. Bulletin de la Societe zoologique de France 14: 298 - 300.
CHEVREUX E. 1895. - Les amphipodes des premieres campagnes de la Princesse-Alice. Memoires de la Societe zoologique de France 8: 424 - 435.
CHEVREUX E. 1899 a. - Sur quelques interessantes especes d'amphipodes provenant de la derniere campagne du yacht Princesse-Alice. Bulletin de la Societe zoologique de France 24: 147 - 152.
CHEVREUX E. 1905. - Description d'un amphipode (Katius obesus, nov. gen. et sp.), suivie d'une liste des amphipodes de la tribu des Gammarina ramenes par le filet a grande ouverture pendant la derniere campagne de la Princesse-Alice en 1904. Bulletin du Musee oceanographique de Monaco 35: 1 - 7.
CHILTON C. 1911. - The Crustacea of the Kermadec Islands. Transactions and Proceedings of the New Zealand Institute 43: 544 - 573.
CHRISTIANSEN B., PFANNKUCHE O. & THIEL H. 1990. - Vertical distribution and population structure of the necrophagous amphipod Eurythenes gryllus in the West European Basin. Marine Ecology Progress Series 66 (1 - 2): 35 - 45.
CHRISTIANSEN B. & THIEL H. 1993. - Megafauna und benthopelagisches Nekton: Regionale Unterschiede zwischen JGOFS-Stationen des Nordostatlantiks. Berichte aus dem Institut fur Meereskunde an der Christian-Albrechts Universitat Kiel 242: 10 - 18.
CHRISTIANSEN B. 1996. - Bait-attending amphipods in the deep-sea: a comparison of three localities in the north-eastern Atlantic. Journal of the Marine Biological Association of the United Kingdom 76: 345 - 360.
CHRISTIANSEN B. & MARTIN B. 2000. - Observations on deep-sea benthopelagic nekton at two stations in the northern Arabian Sea: links to organic matter supply? Deep-Sea Research Part II: Topical Studies in Oceanography 47 (14): 3027 - 3038.
CONROY J. W. H. 1972. - Ecological aspects of the biology of the Giant Petrel, Macronectes giganteus (Gmelin), in the maritime antarctic. British Antarctic Survey Scientific Reports 75: 1 - 74, pls 1 - 7.
COSTELLO M. J., HOLMES J. M. C., MCGRATH D. & MYERS A. A. 1989. - A review and catalogue of the Amphipoda (Crustacea) in Ireland. Irish Fisheries Investigations, ser. B, Marine 33: 1 - 70.
CREASEY S. S. & ROGERS A. D. 1999. - Population genetics of bathyal and abyssal organisms. Advances in Marine Biology 35: 1 - 151.
DAHL E. 1954. - The distribution of deep sea Crustacea. International Union of Biological Sciences, ser. B, 16: 43 - 48.
DAHL E., LAUBIER L., SIBUET M. & STROMBERG J. - O. 1976. - Some quantitative results on benthic communities of the deep Norwegian Sea. Astarte 9: 61 - 79.
DAHL E. 1979. - Deep-sea carrion feeding amphipods: evolutionary patterns in niche adaptation. Oikos 33 (2): 167 - 175.
DAUBY P., SCAILTEUR Y. & DE BROYER C. 2001. - Trophic diversity within the eastern Weddell Sea amphipod community. Hydrobiologia 443 (1 - 3): 69 - 86.
DE BROYER C. & JAZDZEWSKI K. 1993. - Contribution to the marine biodiversity inventory. A checklist of the Amphipoda (Crustacea) of the Southern Ocean. Documents de Travail de l'Institut royal des Sciences naturelles de Belgique 73: 1 - 154.
DELLA VALLE A. 1893. - Gammarini del Golfo di Napoli. Fauna und Flora des Golfes von Neapel 20: 1 - 948, pls 1 - 61.
DESBRUYERES D., GEISTDORFER P., INGRAM C. L., KHRIPOUDOFF A. & LAGARDERE J. P. 1985. - Repartition des populations de l'epibenthos carnivore, in LAUBIER L. & MONNIOT C. (eds), Peuplements profonds du Golfe de Gascogne. IFREMER, Paris: 233 - 251.
FRANCE S. C. & KOCHER T. D. 1996 a. - Geographic and bathymetric patterns of mitochondrial 16 s rRNA sequence divergence among deep-sea amphipods, Eurythenes gryllus. Marine Biology 126: 633 - 643.
GAGE J. D. & TYLER P. A. 1991. - Deep-Sea Biology: A Natural History of Organisms at the Deep-Sea Floor. Cambridge University Press, Cambridge, 504 p.
GEBRUK A. V., GALKIN S. V., VERESCHAKA A. L., MOSKALEV L. I. & SOUTHWARD A. J. 1997. - Ecology and biogeography of the hydrothermal vent fauna of the Mid-Atlantic Ridge. Advances in Marine Biology 32: 93 - 144.
GEORGE R. Y. 1979 a. - Behavioural and metabolic adaptations of polar and deep-sea crustaceans: a hypothesis concerning physiological basis for evolution of cold adapted crustaceans. Bulletin of the Biological Society of Washington 3: 283 - 296.
GEORGE R. Y. 1979 b. - What adaptive strategies promote immigration and speciation in deep-sea environment. Sarsia 64: 61 - 65.
GILCHRIST I. & MACDONALD A. G. 1980. - Hydraulic-decompression neurological syndrome in deep-sea animals. Journal of Physiology (Cambridge) 305: 35 P- 36 P.
GOES A. 1866. - Crustacea Amphipoda maris Spetsbergiam alluentis, cum speciebus aliis Arcticis enumerat. Ofversigt af Kongliga Svenska Vetenskaps- Akademiens Forhandlingar 22: 517 - 36, pls 36 - 41.
GONZALEZ E. 1991. - Actual state of gammaridean amphipoda taxonomy and catalogue of species from Chile. Hydrobiologia 223: 47 - 68.
GORBUNOV G. 1946. - [Bottom life of the Novosiberian shoalwaters and the central part of the Arctic Ocean], [Works of Drifting Ice Expedition in the Central Arctic Ocean in Ice Breaking Steamer T. Sedov]: 30 - 128, 1 pl. (in Russian).
GRIEG J. A. 1925. - Evertebrater fra bankerne ved Spitsbergen indsamlet av m. k. Blaafjeld og m. k. Tovik somrene 1923 og 1924. (Fiskenaering og bundfauna). Bergens Museums Aarbok 9: 1 - 33.
GRIFFITHS C. L. 1975. - The Amphipoda of southern Africa. Part 5. The Gammaridea and Caprellidea of the Cape Province west of Cape Agulhas. Annals of the South African Museum 67 (5): 91 - 181.
GRIFFITHS C. L. 1976. - Guide to the Benthic Marine Amphipods of Southern Africa. Trustees, South African Museum, Cape Town, 106 p.
GRIFFITHS C. L. 1977. - Deep-sea amphipods from west of Cape Point, South Africa. Annals of the South African Museum 73 (4): 93 - 104.
GURJANOVA E. F. 1951. - [Amphipods of the seas of the USSR and surrounding waters (Amphipoda- Gammaridea)]. Akademiya Nauk SSSR, Opredeliteli po Faune SSSR 41: 1 - 1029 (in Russian).
HALLBERG E., NILSSON H. L. & ELOFSSON R. 1980. - Classification of amphipod compound eyes. The fine structure of the ommatidial units (Crustacea, Amphipoda). Zoomorphologie 94: 279 - 306.
HANSEN H. J. 1888. - Malacostraca marina Groenlandiae occidentalis. Oversight over det vestlige Gronlands fauna af malakostrake havkrebsdyr. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kjobenhavn 1887: 5 - 226, pls 2 - 7.
HARGRAVE B. T. 1985. - Feeding rates of abyssal scavenging amphipods (Eurythenes gryllus) determined in situ by time-lapse photography. Deep-Sea Research Part A: Oceanographic Research Papers 32 (4): 443 - 450.
HARGRAVE B. T., GERMAIN P., PHILIPPOT J. C., HEMON G. & SMITH J. N. 1992 a. - Stable elements and 210 Po in the deep-sea amphipod Eurythenes gryllus. Deep-Sea Research Part A: Oceanographic Research Papers 39 (1): 37 - 44.
HARGRAVE B. T., HARDING G. C., VASS W. P., ERICKSON P. E., FOWLER B. R. & SCOTT V. 1992 b. - Organochlorine pesticides and polychlorinated biphenyls in the Arctic Ocean food web. Archives of Environmental Contamination and Toxicology 22 (1): 41 - 54.
HARGRAVE B. T., PROUSE N. J., PHILLIPS G. A. & CRANFORD P. J. 1994. - Meal size and sustenance time in the deep-sea amphipod Eurythenes gryllus collected from the Arctic Ocean. Deep-Sea Research Part I: Oceanographic Research Papers 41 (10): 1489 - 1508.
HARGRAVE B. T., PHILLIPS G. A., PROUSE N. J. & CRANFORD P. J. 1995. - Rapid digestion and assimilation of bait by the deep-sea amphipod Eurythenes gryllus. Deep-Sea Research Part I: Oceanographic Research Papers 42 (11 / 12): 1905 - 1921.
HASEGAWA M., KUROHIJI Y., TAKAYANAGI S., SAWADAISHI S. & YAO M. 1986. - [Collection of fish and Amphipoda from abyssal sea-floor at 30 ° N- 147 ° E using traps tied to 10000 m wire of research vessel]. Bulletin of the Tokai Regional Fisheries Research Laboratory 119: 65 - 75 (in Japanese).
HEINRICH A. K., PARIN N. V., RUDYAKOV Y. A. & SAZHIN A. F. 1993. - [Dwellers of the ocean nearbottom layer]. Akademiya Nauk SSSR, Trudy Instituta Okeanologii 128: 6 - 25 (in Russian).
HESSLER R. R., INGRAM C. L. & SMITH C. R. 1979. - Benthic biological studies. Amphipods. Seabed Disposal Program Annual Report January to December 1977, Vol. III Appendices, Appendix N, Sandia Laboratories: 704 - 714.
HESSLER R. R., 1981. - Biological oceanography model development. Benthic biological studies. Subseabed Disposal Program Report, January to December 1979, Volume II, Part 2, Sandia National Laboratories Report: 397 - 419.
HOLMES S. J. 1903. - Synopses of North-American invertebrates. XVIII. The Amphipoda. American Naturalist 37: 267 - 292.
HOPKINS T. L. 1985. - Food web of an Antarctic midwater ecosystem. Marine Biology 89: 197 - 212.
HURLEY D. E. 1957. - Some Amphipoda, Isopoda and Tanaidacea from Cook Strait. Zoology Publications from Victoria University College 21: 1 - 20.
HURLEY D. E. 1963. - Amphipoda of the family Lysianassidae from the west coast of North and Central America. Allan Hancock Foundation Publications, Occasional Paper 25: 1 - 160.
IMBER M. J. 1973. - The food of Grey-faced Petrels (Pterodroma macroptera gouldi (Hutton )), with special reference to diurnal vertical migration of their prey. Journal of Animal Ecology 42: 645 - 662.
INGRAM C. L. & HESSLER R. R. 1983. - Distribution and behavior of scavenging amphipods from the central North Pacific. Deep-Sea Research Part A: Oceanographic Research Papers 30 (7): 683 - 706.
INGRAM C. L. & HESSLER R. R. 1987. - Population biology of the deep-sea amphipod Eurythenes gryllus: inferences from instar analysis. Deep-Sea Research Part A: Oceanographic Research Papers 34 (12): 1889 - 1919.
INTES A. 1978. - Peche profonde aux casiers en Nouvelle-Caledonie et iles adjacentes. Essais preliminaires. ORSTOM Rapports scientifiques et techniques 2: 1 - 10, figs 1 - 10.
ISHIMARU S. 1994. - A catalogue of gammaridean and ingolfiellidean Amphipoda recorded from the vicinity of Japan. Reports of the Sado Marine Biological Station, Niigata University 24: 29 - 86.
JANSSEN F., TREUDE T. & WITTE U. 2000. - Scavenger assemblages under differing trophic conditions: a case study in the deep Arabian Sea. Deep-Sea Research Part II: Topical Studies in Oceanography 47 (14): 2999 - 3026.
JOHNSON W. S., STEVENS M. & WATLING L. 2001. - Reproduction and development of marine peracaridans. Advances in Marine Biology 39: 105 - 260.
JONES E. M., COLLINS M. A., BAGLEY P. M., ADDISON S. & PRIEDE I. G. 1998. - The fate of cetacean carcasses in the deep sea: observations on consumption rates and succession of scavenging species in the abyssal north-east Atlantic Ocean. Proceedings of the Royal Society, London 265: 1119 - 1127
JUST J. 1980. - Abyssal and deep bathyal Malacostraca (Crustacea) from the Polar Sea. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kjobenhavn 142: 161 - 177.
KAMENSKAYA O. E. 1981. - [The amphipods (Crustacea) from deep-sea trenches in the western part of the Pacific Ocean]. [Deep Sea Bottom Fauna of the Pacific Ocean]. Transactions of the P. P. Shirshov Institute of Oceanology 115: 94 - 107 (in Russian).
KAUFMANN R. S. 1994. - Structure and function of chemoreceptors in scavenging lysianassoid amphipods. Journal of Crustacean Biology 14 (1): 54 - 71.
KLAGES M., VOPEL K., BLUHM H., BREY T., SOLTWEDEL T. & ARNTZ W. E. 2001. - Deep-sea food falls: first observation of a natural event in the Arctic Ocean. Polar Biology 24 (4): 292 - 295.
LAMPITT R. S., MERRETT N. R. & THURSTON M. H. 1983. - Inter-relations of necrophagous amphipods, a fish predator, and tidal currents in the deep sea. Marine Biology 74: 73 - 78.
LAVER M. B., OLSSON M. S., EDELMAN J. L. & SMITH K. L. 1985. - Swimming rates of scavenging deepsea amphipods recorded with a free-vehicle video camera. Deep-Sea Research Part A: Oceanographic Research Papers 32 (9): 1135 - 1142.
LILLJEBORG W. 1865 a. - On the Lysianassa magellanica H. Milne Edwards, and on the Crustacea of the Suborder Amphipoda and Subfamily Lysianassina found an [sic] the Coast of Sweden and Norway. Royal Academic Press, Uppsala, 38 p.
LILLJEBORG W. 1865 b. - Bidrag till kannedomen om underfamilien Lysianassina inom underordningen Amphipoda bland kraftdjuren. Uppsala Universitets Arsskrift 1865: 1 - 25.
LOPES M. F. R., MARQUES J. C. & BELLAN- SANTINI D. 1993. - The benthic amphipod fauna of the Azores (Portugal): an up-to-date annotated list of species, and some biogeographic considerations. Crustaceana 65 (2): 204 - 217.
LOWRY J. K. & BULLOCK S. 1976. - Catalogue of the marine gammaridean Amphipoda of the Southern Ocean. Royal Society of New Zealand Bulletin 16: 1 - 187.
LOWRY J. K. & STODDART H. E. 1993. - Crustacea Amphipoda: Lysianassoids from Philippine and Indonesian waters, in CROSNIER A. (ed.), Resultats des campagnes MUSORSTOM, vol. 10. Memoires du Museum national d'Histoire naturelle A, 156: 55 - 109.
LOWRY J. K. & STODDART H. E. 1994. - Crustacea Amphipoda: Lysianassoids from the tropical western South Pacific Ocean, in CROSNIER A. (ed.), Resultats des campagnes MUSORSTOM, vol. 12. Memoires du Museum national d'Histoire naturelle A, 161: 127 - 223.
LUCAS H. 1857. - Entomologie, in Expedition dans les parties centrales de l'Amerique du Sud, de Rio de Janeiro a Lima, et de Lima au Para; executees par ordre du gouvernement francais pendant les annees 1843 a 1847, sous la direction du comte Francis de Castelnau. Part 7 Zoologie, Volume 3. P. Bertrand, Paris: 204 p., 19 pls.
SAINTE- MARIE B. & HARGRAVE B. T. 1987. - Estimation of scavenger abundance and distance of attraction to bait. Marine Biology 94: 431 - 443.
MATSUMASA M., KIKUCHI S. & TAKEUCHI I. 1998. - Specialized ion-transporting epithelium around the blood vessel of the coxal gills in a deep-sea amphipod, Eurythenes gryllus. Journal of Crustacean Biology 18 (4): 686 - 694.
MILNE EDWARDS H. 1848. - Sur un crustace amphipode, remarquable par sa grande taille. Annales des Sciences naturelles ser. 3, 9: 398.
MURDOCH J. 1885. - Marine invertebrates (exclusive of Mollusks), in RAY P. H. (ed.), Report of the International Polar Expedition to Point Barrow, Alaska. Government Printing Office, Washington DC: 136 - 176, pls 1, 2.
OLDEVIG H. 1959. - Arctic, subarctic and Scandinavian amphipods in the collections of the Swedish Natural History Museum in Stockholm. Goteborgs Kungliga Vetenskaps- och Vitterhets- Samhalles Handlingar ser. B, 8 (2): 1 - 132, 4 pls.
ORTIZ M. 1979. - Lista de especes y bibliografia de los anfipodos (Crustacea: Amphipoda) del Mediterraneo Americano. Ciencias (La Habana) ser. 8, Investigaciones Marinas 43: 1 - 40.
PALERUD R. & VADER W. 1991. - Marine Amphipoda Gammaridea in north-east Atlantic and Norwegian Arctic. Tromura, Naturvitenskap 68: 1 - 97.
PAUL A. Z. 1973. - Trapping and recovery of living deep-sea amphipods from the Arctic Ocean floor. Deep-Sea Research and Oceanographical Abstracts 20 (3): 289 - 290.
PETTER A. J. 1983. - Description d'un nouveau genre de Benthimermithidae (Nematoda) presentant des uterus munis de glandes annexes. Annales de Parasitologie humaine et comparee 58 (2): 177 - 184.
PIATKOWSKI U., RODHOUSE P. G., WHITE M. G., BONE D. G. & SYMON C. 1994. - Nekton community of the Scotia Sea as sampled by the RMT 35 during austral summer. Marine Ecology Progress Series 112 (1 - 2): 13 - 28.
POUPIN J., TAMARII T. & VANDENBOOMGAERDE A. 1990. - Peches profondes aux casiers sur les pentes oceaniques des iles de Polynesie francaise. (N / O Marara - 1986 / 1989). Notes et Documents d'Oceanographie du Centre ORSTOM de Tahiti 42: 1 - 97, pls 1 - 3.
POUPIN J. 1994. - Faune marine profonde des Antilles francaises. Recoltes du navire Polka faites en 1993. ORSTOM, Paris, 79 p.
POUPIN J. 1996. - Atlas des crustaces marins profonds de Polynesie francaise. Recoltes du navire Marara (1986 / 1996). Service mixte de Surveillance radiologique et biologique, Montlhery, 59 p.
PREMKE K., MUYAKSHIN S., KLAGES M. & WEGNER J. 2003. - Evidence for long-range chemoreceptive tracking of food odour in deep-sea scavengers by scanning sonar data. Journal of Experimental Marine Biology and Ecology 285: 283 - 294.
PRINCE P. A. 1980. - The food and feeding ecology of Blue Petrel (Halobaena caerulea) and Dove Prion (Pachyptila desolata). Journal of Zoology (London) 190 (1): 59 - 76.
RANNOU M. & NOUGUIER J. 1974. - Peches abyssales aux casiers. Annales de l'Institut oceanographique, Monaco 50 (2): 139 - 143.
RAUSCHERT M. 1985. - Eurythenes gryllus (Lichtenstein) (Crustacea, Amphipoda) in der marinen Fauna von King George (Sudshetlandinseln, Antarktis). Milu, Berlin 6: 319 - 324.
RAUSCHERT M. 1990. - New amphipods from the sublittoral of King George Island. Faunistic contribution to ecological investigations. Geodatische und geophysikalische Veroffentlichungen, Reihe I, Berlin 16: 447 - 458.
RAUSCHERT M. 1991. - Ergebnisse der faunistischen Arbeiten im Benthal von King George Island (Sudshetlandinseln, Antarktis). Berichte zur Polarforschung 76: 1 - 75.
REINHARDT S. B. & VAN VLEET E. S. 1986. - Lipid composition of twenty-two species of Antarctic midwater zooplankton and fish. Marine Biology 91: 149 - 159.
SARS M. 1869. - Fortsaette Bemaerkninger over det dyriske Livs Udbredning i Havets Dybder. Forhandlinger i Videnskabs-Selskabet i Christiania 1868: 246 - 275.
SARS G. O. 1891. - An Account of the Crustacea of Norway, with Short Descriptions and Figures of all the Species. Vol. I. Amphipoda. Parts 4 - 9. Alb. Cammermeyer, Christiana: 69 - 212, pls 25 - 72.
SCHELLENBERG A. 1926. - Amphipoda 3: Die Gammariden der Deutschen Tiefsee-Expedition. Wissenschaftliche Ergebnisse der Deutschen Tiefsee- Expedition auf dem Dampfer Valdivia 1898 - 1899 23 (5): 193 - 243, pl. 5.
SCHELLENBERG A. 1927. - Amphipoda des nordischen Plankton. Nordisches Plankton 6: 589 - 722.
SCHELLENBERG A. 1955. - Amphipoda. Reports of the Swedish Deep-Sea Expedition ser. 2, Zoology, 14: 181 - 195.
SHOEMAKER C. R. 1920. - The amphipods of the Canadian Arctic Expedition, 1913 - 18. Report of the Canadian Arctic Expedition, 1913 - 18 7 (E): 3 E- 30 E.
SHOEMAKER C. R. 1945. - The Amphipoda of the Bermuda Oceanographic Expeditions, 1929 - 1931. Zoologica, Scientific Contributions of the New York Zoological Society 30 (4): 185 - 266.
SHOEMAKER C. R. 1956. - Notes on the amphipods Eurythenes gryllus (Lichtenstein) and Katius obesus Chevreux. Proceedings of the Biological Society of Washington 69: 177 - 178.
SHULENBERGER E. & HESSLER R. R. 1974. - Scavenging abyssal benthic amphipods trapped under oligotrophic central North Pacific gyre waters. Marine Biology 28: 185 - 187.
SHULENBERGER E. & BARNARD J. L. 1976. - Amphipods from an abyssal trap set in the North Pacific gyre. Crustaceana 31 (3): 241 - 258.
SMITH S. I. 1884 a. - Crustacea of the Albatross dredgings in 1883. American Journal of Science ser. 3, 28: 53 - 56.
SMITH S. I. 1884 b. - Crustacea of the Albatross dredgings in 1883. Annals and Magazine of Natural History ser. 5, 14: 179 - 183 (reprinted from the American Journal of Science).
SMITH K. L., WHITE G. A., LAVER M. B., MCCONNAUGHEY R. R. & MEADOR J. P. 1979. - Free vehicle capture of abyssopelagic animals. Deep- Sea Research Part A: Oceanographic Research Papers 26 (1): 57 - 64.
SMITH C. R. & PRESENT T. M. C. 1983. - In vivo marking of shallow-water and deep-sea amphipods by ingestion of bait mixed with fast green. Marine Biology 73: 183 - 192.
SMITH K. L. & BALDWIN R. J. 1984. - Vertical distribution of the necrophagous amphipod, Eurythenes gryllus, in the North Pacific: spatial and temporal variation. Deep-Sea Research Part A: Oceanographic Research Papers 31 (10): 1179 - 1196.
SMITH K. L., KAUFMANN R. S., EDELMAN J. L. & BALDWIN R. J. 1992. - Abyssopelagic fauna in the central North Pacific: comparison of acoustic detection and trawl and baited trap collections to 5800 m. Deep-Sea Research Part A: Oceanographic Research Papers 39 (3 / 4): 659 - 685.
SPRINGER S. & BULLIS H. R. 1956. - Collections by the Oregon in the Gulf of Mexico. List of crustaceans, mollusks, and fishes identified from collections made by the exploratory fishing vessel Oregon in the Gulf of Mexico and adjacent seas 1950 through 1955. United States Department of the Interior, Special Scientific Report: Fisheries 196: 1 - 134.
STEBBING T. R. R. 1893. - A History of Crustacea Recent Malacostraca. Kegan Paul, Trench, Trubner & Company, London, 466 p.
STEBBING T. R. R. 1906. - Amphipoda. I. Gammaridea. Das Tierreich 21: 1 - 806.
STEELE D. H. & STEELE V. J. 1991. - The structure and organization of the gills of gammaridean Amphipoda. Journal of Natural History 25: 1247 - 1258.
STEPHENSEN K. 1912 b. - Report on the Malacostraca Pycnogonida and some Entomostraca collected by the Danmark Expedition to north-east Greenland. Meddelelser om Gronland 45: 501 - 630, pls 39 - 43.
STEPHENSEN K. 1912 a. - Report on the Malacostraca collected by the Tjalfe - Expedition under the direction of cand. mag. Ad. S. Jensen, especially at W. Greenland. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening 64: 57 - 134.
STEPHENSEN K. 1913. - Gronlands Krebsdyr og Pycnogonider (Conspectus Crustaceorum et Pycnogonidorum Groenlandiae). Meddelelser om Gronland 22: 1 - 479.
STEPHENSEN K. 1915. - Isopoda, Tanaidacea, Cumacea, Amphipoda (excl. Hyperiidea). Report on the Danish Oceanographical Expeditions 1908 - 10 to the Mediterranean and Adjacent Seas 2, Biology (D 1): 1 - 53.
STEPHENSEN K. 1925. - Crustacea Malacostraca, VI: (Amphipoda, II). Danish Ingolf-Expedition 3 (9): 101 - 178.
STEPHENSEN K. 1932. - The Tanaidacea and Amphipoda of the Arctic. Fauna Arctica 6: 343 - 378.
STEPHENSEN K. 1933. - The Godthaab Expedition 1928. Amphipoda. Meddelelser om Gronland 79 (7): 1 - 88.
STUXBERG A. 1880. - Evertebratfaunan i Sibiriens Ishaf. Forelopande studies grundade pa de zoologiska undersokningarna under Prof. A. E. Nordenskiolds Ishafs-expedition 1878 - 79. Bihang till Kunliga Svenska Vetenskaps-Akademiens Handlingar 5 (22): 1 - 76.
TAKEUCHI I. & WATANABE K. 1998. - Respiration rate and swimming speed of the necrophagous amphipod Eurythenes gryllus from Antarctic waters. Marine Ecology Progress Series 163: 285 - 288.
TAKEUCHI I., WATANABE K., TANIMURA A. & FUKUCHI M. 2001. - Assemblages of necrophagous animals off Enderby Land, East Antarctica. Polar Biology 24 (9): 650 - 656.
TATTERSALL W. M. 1906. - The marine fauna of the coast of Ireland. Part VIII. Pelagic Amphipoda of the Irish Atlantic slope. Department of Agriculture and Technical Instruction for Ireland, Fisheries Branch, Scientific Investigations 1905 (4): 3 - 39, pls 1 - 5.
TATTERSALL W. M. 1909. - Amphipoda and Isopoda, with descriptions of two new species, in WOLFENDEN R. N. (ed.), Scientific and biological researches in the North Atlantic, conducted by the author on his yachts The Walwin and The Silver Belle. Memoirs of the Challenger Society 1: 210 - 219, pls 4, 5.
TEMPLEMAN W. 1967. - Predation on living fishes on longline in Baffin Bay by the amphipod Eurythenes gryllus (Lichtenstein), and a new distribution record. Journal of the Fisheries Research Board of Canada 24 (1): 215 - 217, pls 1 - 3.
THURSTON M. H. 1979. - Scavenging abyssal amphipods from the north-east Atlantic Ocean. Marine Biology 51 (1): 55 - 68.
THURSTON M. H. 1990. - Abyssal necrophagous amphipods (Crustacea: Amphipoda) in the northeast and tropical Atlantic Ocean. Progress in Oceanography 24: 257 - 274.
THURSTON M. H. 1994. - Eurythenes gryllus hatchlings. Deep-Sea Newsletter 22: 14 - 15.
THURSTON M. H. & BETT B. J. 1995. - Hatchling size and aspects of biology in the deep-sea amphipod genus Eurythenes (Crustacea: Amphipoda). Internationale Revue der Gesamten Hydrobiologie 80 (2): 201 - 216.
THURSTON M. H., PETRILLO M. & DELLA CROCE N. 2002. - Population structure of the necrophagous amphipod Eurythenes gryllus (Amphipoda: Gammaridea) from the Atacama Trench (south-east Pacific Ocean). Journal of the Marine Biological Association of the United Kingdom 82 (2): 205 - 211.
TREUDE T., JANSSEN F., QUEISSER W. & WITTE U. 2002. - Metabolism and decompression tolerance of scavenging lysianassoid deep-sea amphipods. Deep-Sea Research Part I: Oceanographic Research Papers 49 (7): 1281 - 1289.
UMEZU T. 1982. - Deep sea pelagos. Part 2. Collection by KAIYO-maru for radionuclide analysis. Aquabiology (Tokyo) 4 (1): 2 - 9 (in Japanese with English summary).
UMEZU T. 1984. - Deep-sea biomass in North Pacific Polar Frontal Zone 40 ° N- 150 ° E collected by KOC-net from 150 - 4500 m depth in May 1981. Bulletin of the Tokai Regional Fisheries Research Laboratory 113: 115 - 139 (in Japanese with English summary).
VINOGRADOV G. M. 1990. - [Pelagic amphipods (Amphipoda, Crustacea) from the south-eastern Pacific]. Akademiya Nauk SSSR, Trudy Instituta Okeanologii 124: 27 - 104 (in Russian with English summary).
VINOGRADOV M. E., VERESCHCHAKA A. L. & VINOGRADOV G. M. 1996. - Visual observations from DSRV Mir in location of the Russian submarine Komsomolet wreck. Deep-Sea Newsletter 24: 7 - 8.
WAKABARA Y., TARARAM A. S., VALERIO- BERARDO M. T. & OGIHARA R. M. 1990. - Records of Amphipoda collected during I and III Brazilian Antarctic Expeditions. Relatorio interno do Instituto Oceanografico Universidade de Sao Paulo 23: 1 - 10.
WAKABARA Y., TARARAM A. S. & MIYAGI V. K. 1996. - The amphipod fauna of the West Antarctic region (South Shetland Islands and Bransfield Strait), in JAZDZEWSKI K., DE BROYER C. & STOCK J. H. (eds), Biology and ecology of amphipod crustaceans. Polskie Archiwum Hydrobiologii 42 (4): 347 - 365.
WICKINS J. F. 1983. - Catches of large lysianassid amphipods in baited traps at the Nuclear Energy Authority dumpsite during June 1979. Deep-Sea Research Part A: Oceanographic Research Papers 30 (1): 83 - 86.
WILSON R. R., SMITH K. L. & ROSENBLATT R. H. 1985. - Megafauna associated with bathyal seamounts in the central North Pacific Ocean. Deep-Sea Research Part A: Oceanographic Research Papers 32 (10): 1243 - 1254.
WITTE U. 1999. - Consumption of large carcasses by scavenger assemblages in the deep Arabian Sea: observations by baited camera. Marine Ecology Progress Series 183: 139 - 147.
YAYANOS A. A. 1978. - Recovery and maintenance of live amphipods at a pressure of 580 bars from an ocean depth of 5700 meters. Science 200: 1056 - 1059.
FIG. 2. — Eurythenes gryllus (Lichtenstein in Mandt, 1822), holotype 81 mm (ZMB 1265). Abbreviations: G1, G2, gnathopods 1, 2; P3-P7, pereopods 3-7. Scale bars: 2.0 mm.
FIG. 3. — Eurythenes gryllus (Lichtenstein in Mandt, 1822), holotype 81 mm (ZMB 1265). Abbreviations: A1, A2, antennae 1, 2; EP1-EP3, epimera 1-3; H, head; T, telson; U1-U3, uropods 1-3. Scale bars: U1-3, T, 2.0 mm; A1, A2, 1.0 mm.
FIG. 4. — Eurythenes gryllus (Lichtenstein in Mandt, 1822), holotype of Lysianassa magellanica H. Milne Edwards, 1848, c. 85 mm, (MNHN-Am 3148).
FIG. 5. — Eurythenes gryllus (Lichtenstein in Mandt, 1822), holotype of Lysianassa magellanica H. Milne Edwards, 1848, c. 85 mm (MNHN-Am 3148). Abbreviations: A1, A2, antennae 1, 2; H, head; LMD, left mandible; MP, maxilliped; MX1, MX2, maxillae 1, 2; ★, lateral view of right mandible; ✪, maxilliped palp completely flattened. Scale bars: 1.0 mm.
FIG. 6. — Eurythenes gryllus (Lichtenstein in Mandt, 1822), holotype of Lysianassa magellanica H. Milne Edwards, 1848, c. 85 mm (MNHN-Am 3148). Abbreviations: G1, G2, gnathopods 1, 2; P3-P7, pereopods 3-7. Scale bars: 2.0 mm.
FIG. 7. — Eurythenes gryllus (Lichtenstein in Mandt, 1822), holotype of Lysianassa magellanica H. Milne Edwards, 1848, c. 85 mm (MNHN-Am 3148). Abbreviations: T, telson; U1-U3, uropods 1-3. Scale bars: 1.0 mm.
FIG. 8. — Eurythenes gryllus (Lichtenstein in Mandt, 1822), material reported as Eurytenes magellanicus by Lilljeborg (1865a), immature 55 mm (MNHN-Am 4479).
FIG. 9. — Eurythenes gryllus (Lichtenstein in Mandt, 1822), material reported as Eurytenes magellanicus by Lilljeborg (1865a), immature 55 mm (MNHN-Am 4479). Abbreviations: A1, A2, antennae 1, 2; H, head; MD, mandible; MP, maxilliped; MX1, MX2, maxillae 1, 2; U1-U3, uropods 1-3; T, telson. Scale bars: A1, A2, 0.5 mm; others, 1.0 mm.
FIG. 10. — Eurythenes gryllus (Lichtenstein in Mandt, 1822), material reported as Eurytenes magellanicus by Lilljeborg (1865a), immature 55 mm (MNHN-Am 4479). Abbreviations: G1, G2, gnathopods 1, 2; P3-P7, pereopods 3-7. Scale bars: 1.0 mm.
FIG. 13. — Eurythenes obesus (Chevreux, 1905), neotype 48 mm (BMNH 2003.1059). Abbreviations: A1, A2, antennae 1, 2; H, head; MD, mandible; MP, maxilliped; MX1, MX2, maxillae 1, 2. Scale bars: 0.5 mm.
FIG. 14. — Eurythenes obesus (Chevreux, 1905), neotype 48 mm (BMNH 2003.1059). Abbreviations: EP1-EP3, epimera 1-3; G1, G2, gnathopods 1, 2; P3-P7, pereopods 3-7; T, telson; U1-U3, uropods 1-3. Scale bars: 1.0 mm.
| AM |
Australian Museum |
| NMV |
Museum Victoria |
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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Eurythenes gryllus
| Stoddart, Helen E. & Lowry, James K. 2004 |
Eurythenes cf. gryllus
| LOWRY J. K. & STODDART H. E. 1994: 158 |
Eurythennes gryllus
| POUPIN J. 1996: 42 |
| POUPIN J. 1994: 16 |
| POUPIN J. & TAMARII T. & VANDENBOOMGAERDE A. 1990: 94 |
Eurythenes obesus
| CHEREL Y. & BOCHER P. & DE BROYER C. & HOBSON K. A. 2002: 268 |
| JOHNSON W. S. & STEVENS M. & WATLING L. 2001: 198 |
| THURSTON M. H. & BETT B. J. 1995: 201 |
| KAUFMANN R. S. 1994: 54 |
| PIATKOWSKI U. & RODHOUSE P. G. & WHITE M. G. & BONE D. G. & SYMON C. 1994: 19 |
| DE BROYER C. & JAZDZEWSKI K. 1993: 68 |
| BARNARD J. L. & KARAMAN G. S. 1991: 486 |
| BOUDRIAS M. A. 1991: 13 |
| PALERUD R. & VADER W. 1991: 35 |
| COSTELLO M. J. & HOLMES J. M. C. & MCGRATH D. & MYERS A. A. 1989: 32 |
| ANDRES H. G. 1987: 96 |
| HOPKINS T. L. 1985: 202 |
| UMEZU T. 1984: 128 |
| ANDRES H. G. 1983: 186 |
| ORTIZ M. 1979: 19 |
| GRIFFITHS C. L. 1976: 56 |
| LOWRY J. K. & BULLOCK S. 1976: 89 |
| GRIFFITHS C. L. 1975: 145 |
| IMBER M. J. 1973: 652 |
| BIRSTEIN J. A. & VINOGRADOV M. E. 1970: 420 |
| BRUSCA G. J. 1967: 384 |
| BIRSTEIN J. A. & VINOGRADOV M. E. 1964: 163 |
| HURLEY D. E. 1963: 59 |
| BIRSTEIN J. A. & VINOGRADOV M. E. 1962: 36 |
| BARNARD J. L. 1961: 38 |
| BIRSTEIN J. A. & VINOGRADOV M. E. 1960: 184 |
| BARNARD J. L. 1958: 92 |
| SHOEMAKER C. R. 1956: 178 |
| SCHELLENBERG A. 1955: 183 |
Eurythenes gryllus
| LOWRY J. K. & STODDART H. E. 1993: 72 |
| SPRINGER S. & BULLIS H. R. 1956: 6 |
| BARNARD K. H. 1940: 440 |
Eurythenes gryllus
| SPRINGER S. & BULLIS H. R. 1956: 6 |
| BARNARD K. H. 1940: 440 |
Eurythenes magellanicus
| ARNAUD P. M. 1974: 592 |
| CONROY J. W. H. 1972: 56 |
| BECK J. R. 1969: 35 |
| BARNARD J. L. 1958: 92 |
| BARNARD K. H. 1932: 59 |
Eurythenes gryllus
| STEPHENSEN K. 1932: 356 |
Katius obesus
| COSTELLO M. J. & HOLMES J. M. C. & MCGRATH D. & MYERS A. A. 1989: 32 |
| BELLOC G. 1960: 7 |
| SHOEMAKER C. R. 1956: 177 |
| STEPHENSEN K. 1933: 12 |
| BARNARD K. H. 1932: 56 |
| SCHELLENBERG A. 1926: 217 |
| STEPHENSEN K. 1925: 126 |
| SHOEMAKER C. R. 1920: 8 |
| STEPHENSEN K. 1915: 37 |
| STEPHENSEN K. 1913: 123 |
| STEPHENSEN K. 1912: 89 |
| STEPHENSEN K. 1912: 614 |
| TATTERSALL W. M. 1909: 210 |
| STEBBING T. R. R. 1906: 721 |
| TATTERSALL W. M. 1906: 29 |
| CHEVREUX E. 1905: 1 |
Euryporeia gryllus
| GRIEG J. A. 1925: 21 |
| BRUGGEN E. 1907: 215 |
| HOLMES S. J. 1903: 277 |
| CHEVREUX E. 1899: 147 |
| CHEVREUX E. 1895: 426 |
| DELLA VALLE A. 1893: 848 |
| SARS G. O. 1891: 86 |
Eurythenes gryllus
| PREMKE K. & MUYAKSHIN S. & KLAGES M. & WEGNER J. 2003: 283 |
| CHEREL Y. & BOCHER P. & DE BROYER C. & HOBSON K. A. 2002: 272 |
| THURSTON M. H. & PETRILLO M. & DELLA CROCE N. 2002: 205 |
| TREUDE T. & JANSSEN F. & QUEISSER W. & WITTE U. 2002: 1281 |
| BLUHM B. A. & BREY T. & KLAGES M. & ARNTZ W. E. 2001: 642 |
| BUHRING S. I. & CHRISTIANSEN B. 2001: 369 |
| DAUBY P. & SCAILTEUR Y. & DE BROYER C. 2001: 81 |
| JOHNSON W. S. & STEVENS M. & WATLING L. 2001: 198 |
| KLAGES M. & VOPEL K. & BLUHM H. & BREY T. & SOLTWEDEL T. & ARNTZ W. E. 2001: 293 |
| TAKEUCHI I. & WATANABE K. & TANIMURA A. & FUKUCHI M. 2001: 653 |
| CHRISTIANSEN B. & MARTIN B. 2000: 3027 |
| JANSSEN F. & TREUDE T. & WITTE U. 2000: 3011 |
| CREASEY S. S. & ROGERS A. D. 1999: 28 |
| WITTE U. 1999: 142 |
| JONES E. M. & COLLINS M. A. & BAGLEY P. M. & ADDISON S. & PRIEDE I. G. 1998: 1124 |
| MATSUMASA M. & KIKUCHI S. & TAKEUCHI I. 1998: 686 |
| TAKEUCHI I. & WATANABE K. 1998: 285 |
| GEBRUK A. V. & GALKIN S. V. & VERESCHAKA A. L. & MOSKALEV L. I. & SOUTHWARD A. J. 1997: 116 |
| WAKABARA Y. & TARARAM A. S. & MIYAGI V. K. 1996: 355 |
| CHRISTIANSEN B. 1996: 345 |
| FRANCE S. C. & KOCHER T. D. 1996: 633 |
| VINOGRADOV M. E. & VERESCHCHAKA A. L. & VINOGRADOV G. M. 1996: 8 |
| HARGRAVE B. T. & PHILLIPS G. A. & PROUSE N. J. & CRANFORD P. J. 1995: 1905 |
| THURSTON M. H. & BETT B. J. 1995: 201 |
| THURSTON M. H. 1994: 14 |
| HARGRAVE B. T. & PROUSE N. J. & PHILLIPS G. A. & CRANFORD P. J. 1994: 1489 |
| BRITTON J. C. & MORTON B. 1994: 391 |
| ISHIMARU S. 1994: 58 |
| KAUFMANN R. S. 1994: 54 |
| BRITTON J. C. & MORTON B. 1993: 369 |
| CHRISTIANSEN B. & THIEL H. 1993: 12 |
| DE BROYER C. & JAZDZEWSKI K. 1993: 67 |
| HEINRICH A. K. & PARIN N. V. & RUDYAKOV Y. A. & SAZHIN A. F. 1993: 6 |
| LOPES M. F. R. & MARQUES J. C. & SANTINI D. 1993: 209 |
| HARGRAVE B. T. & GERMAIN P. & PHILIPPOT J. C. & HEMON G. & SMITH J. N. 1992: 37 |
| HARGRAVE B. T. & HARDING G. C. & VASS W. P. & ERICKSON P. E. & FOWLER B. R. & SCOTT V. 1992: 41 |
| SMITH K. L. & KAUFMANN R. S. & EDELMAN J. L. & BALDWIN R. J. 1992: 669 |
| BARNARD J. L. & KARAMAN G. S. 1991: 486 |
| BOUDRIAS M. A. 1991: 13 |
| GAGE J. D. & TYLER P. A. 1991: 78 |
| GONZALEZ E. 1991: 59 |
| PALERUD R. & VADER W. 1991: 35 |
| RAUSCHERT M. 1991: 37 |
| STEELE D. H. & STEELE V. J. 1991: 1250 |
| THURSTON M. H. 1990: 262 |
| CHRISTIANSEN B. & PFANNKUCHE O. & THIEL H. 1990: 35 |
| RAUSCHERT M. 1990: 454 |
| VINOGRADOV G. M. 1990: 42 |
| WAKABARA Y. & TARARAM A. S. & BERARDO M. T. & OGIHARA R. M. 1990: 5 |
| CHARMASSON S. S. & CALMET D. P. 1989: 163 |
| COSTELLO M. J. & HOLMES J. M. C. & MCGRATH D. & MYERS A. A. 1989: 32 |
| ANDRES H. G. 1987: 96 |
| INGRAM C. L. & HESSLER R. R. 1987: 1889 |
| BALDWIN R. J. & SMITH K. L. 1987: 425 |
| BUCKLIN A. & WILSON R. R. & SMITH L. S. 1987: 1795 |
| CHARMASSON S. S. & CALMET D. P. 1987: 1509 |
| MARIE B. & HARGRAVE B. T. 1987: 436 |
| AINLEY D. G. & FRASER W. R. & SULLIVAN C. W. & TORRES J. J. & HOPKINS T. L. & SMITH W. O. 1986: 848 |
| HASEGAWA M. & KUROHIJI Y. & TAKAYANAGI S. & SAWADAISHI S. & YAO M. 1986: 70 |
| REINHARDT S. B. & VAN VLEET E. S. 1986: 151 |
| DESBRUYERES D. & GEISTDORFER P. & INGRAM C. L. & KHRIPOUDOFF A. & LAGARDERE J. P. 1985: 233 |
| HARGRAVE B. T. 1985: 443 |
| HOPKINS T. L. 1985: 202 |
| LAVER M. B. & OLSSON M. S. & EDELMAN J. L. & SMITH K. L. 1985: 1136 |
| RAUSCHERT M. 1985: 319 |
| WILSON R. R. & SMITH K. L. & ROSENBLATT R. H. 1985: 1248 |
| UMEZU T. 1984: 128 |
| SMITH K. L. & BALDWIN R. J. 1984: 1179 |
| ANDRES H. G. 1983: 186 |
| INGRAM C. L. & HESSLER R. R. 1983: 683 |
| LAMPITT R. S. & MERRETT N. R. & THURSTON M. H. 1983: 73 |
| PETTER A. J. 1983: 177 |
| SMITH C. R. & PRESENT T. M. C. 1983: 183 |
| WICKINS J. F. 1983: 83 |
| UMEZU T. 1982: 2 |
| HESSLER R. R. 1981: 397 |
| KAMENSKAYA O. E. 1981: 95 |
| HALLBERG E. & NILSSON H. L. & ELOFSSON R. 1980: 280 |
| JUST J. 1980: 164 |
| PRINCE P. A. 1980: 63 |
| ANDRES H. G. 1979: 96 |
| DAHL E. 1979: 168 |
| GEORGE R. Y. 1979: 283 |
| GEORGE R. Y. 1979: 63 |
| HESSLER R. R. & INGRAM C. L. & SMITH C. R. 1979: 704 |
| ORTIZ M. 1979: 19 |
| SMITH K. L. & WHITE G. A. & LAVER M. B. & MCCONNAUGHEY R. R. & MEADOR J. P. 1979: 57 |
| THURSTON M. H. 1979: 56 |
| INTES A. 1978: 4 |
| GRIFFITHS C. L. 1977: 97 |
| DAHL E. & LAUBIER L. & SIBUET M. & STROMBERG J. - O. 1976: 75 |
| LOWRY J. K. & BULLOCK S. 1976: 90 |
| SHULENBERGER E. & BARNARD J. L. 1976: 241 |
| RANNOU M. & NOUGUIER J. 1974: 142 |
| SHULENBERGER E. & HESSLER R. R. 1974: 185 |
| PAUL A. Z. 1973: 289 |
| BOWMAN T. E. & MANNING R. B. 1972: 193 |
| BIRSTEIN J. A. & VINOGRADOV M. E. 1970: 420 |
| BECK J. R. 1969: 34 |
| TEMPLEMAN W. 1967: 215 |
| BIRSTEIN J. A. & VINOGRADOV M. E. 1962: 36 |
| BARNARD J. L. 1961: 35 |
| BIRSTEIN J. A. & VINOGRADOV M. E. 1960: 183 |
| OLDEVIG H. 1959: 19 |
| BIRSTEIN J. A. & VINOGRADOV M. E. 1958: 228 |
| BARNARD J. L. 1958: 92 |
| HURLEY D. E. 1957: 2 |
| BIRSTEIN J. A. & VINOGRADOV M. E. 1955: 225 |
| DAHL E. 1954: 3 |
| GURJANOVA E. F. 1951: 265 |
| GORBUNOV G. 1946: 43 |
| SHOEMAKER C. R. 1945: 186 |
| BARNARD K. H. 1937: 144 |
| STEPHENSEN K. 1932: 356 |
| SCHELLENBERG A. 1927: 679 |
| STEPHENSEN K. 1925: 110 |
| CHILTON C. 1911: 563 |
| STEBBING T. R. R. 1906: 73 |
| STEBBING T. R. R. 1893: 80 |
| CHEVREUX E. 1889: 298 |
| HANSEN H. J. 1888: 67 |
| SMITH S. I. 1884: 54 |
Eurysthenes gryllus
| GILCHRIST I. & MACDONALD A. G. 1980: 35 |
| SMITH S. I. 1884: 181 |
Eurytenes gryllus
| STEPHENSEN K. 1912: 528 |
| STUXBERG A. 1880: 62 |
| BOECK A. 1872: 144 |
| BOECK A. 1871: 105 |
Lysianassa gryllus
| BATE C. S. 1867: 229 |
| GOES A. 1866: 517 |
Eurytenes magellanicus
| LILLJEBORG W. 1865: 11 |
Lysianassa magellanica Milne Edwards 1848: 398
| BETHUNE A. 1869: 431 |
| SARS M. 1869: 260 |
| LILLJEBORG W. 1865: 3 |
| LILLJEBORG W. 1865: 2 |
| BATE C. S. 1862: 66 |
| LUCAS H. 1857: 13 |
| MILNE EDWARDS H. 1848: 398 |