Rugosothynnus

Brown, Graham, 2015, Rugosothynnus gen. nov. (Hymenoptera: Tiphiidae: Thynninae: Rhagigasterini), a newly recognised Australian genus, Zootaxa 3925 (3), pp. 361-386 : 383-384

Key to the species of Rugosothynnus

Males

1. Clypeus flat or convex over at least half length in profile and with or without small central polished area ( Fig. 2 ), not strongly depressed below branches of carina....................................................................... 2

- Clypeus distinctly concave either in profile and usually over about half of length but only near apex in R. monteithae sp. nov., or abruptly depressed below branches of carina when well defined, or both, much of depression polished ( Fig. 44 )......... 4

2(1). S2 distinctly subtuberculate anteromedially (as Figs 47–48 ); S1 sagittally carinate; pronotum rugose [SWA].................................................................................................. R. houstoni sp. nov.

- S2 not subtuberculate; S1 not sagittally carinate (beyond basal blade-like projection); pronotum closely punctate to transversely rugose-punctate................................................................................. 3

3(2). Clypeus with central inverted cordate area; parameres weakly lobed near apex of basiparameres; S1 strongly medially raised [SEQ]........................................................................... R. neocorrugatus sp. nov.

- Clypeus without a cordate area; parameres strongly medially lobed near apex of aedeagus; S1 broadly medially raised [SEA]................................................................................... R. corrugatus (Turner)

4(1). Legs ferruginous; vertex and pronotum shallowly and punctate to sparsely punctate ( Figs 1–3 ) [SEQ, NNSW]................................................................................................... R. burnsi (Given)

- Legs black; vertex and pronotum closely punctate to rugose.................................................... 5

5(4). Clypeus with carina well developed, branches broadly diverging, long and reaching clypeal margin, apical margin broadly emarginate ( Fig. 44 ); S8 with preapical spine reduced, weakly toothed, not spinose [SWA].......... R. depressus sp. nov.

- Clypeus not both deeply and broadly emarginate; S8 with well developed preapical spine Fig. 47 )...................... 6

6(5). Branches of clypeal carina strongly curved, well defined, strongly depressed between branches [SWA].. R. clypeatus sp. nov.

- Branches of clypeal carina, when present, not curved......................................................... 7

7(6). S1 sagittally carinate with second raised blade-like projection posteriorly ( Fig. 49 ) [SWA]............. R. tristis (Smith)

- S1 not raised posteriorly into second blade-like projection..................................................... 8

8(7). S2 weakly raised anteromedially but not tuberculate; vertex and dorsum of mesosoma rugosely punctate [SEQ].............................................................................................. R. monteithae sp. nov.

- S2 distinctly subtuberculate anteromedially; vertex and dorsum of mesosoma rugose [SWA].......................... 9

9(8). Cuspis bifid in profile with distinct lobes ( Figs 11 , 29 )....................................................... 10

- Cuspis truncate in profile without lobes ( Figs 20 , 39 )........................................................ 11

10(9). Lower lobe of cuspis projecting beyond upper lobe and about three times longer than upper lobe; hypopygial spine evenly curved in profile ( Figs 29–30 )............................................................ R. fulvescens sp. nov.

- Lower lobe of cuspis downturned and not projecting beyond apex of upper lobe, about two times longer than upper lobe; hypopygial spine angulate in profile ( Figs 11–12 )............................................ R. brunneus sp. nov.

11(9). Parameres lobed distal to apex of aedeagus and basiparameres ( Figs 20–21 )........................ R. confusus sp. nov.

- Parameres lobed near apex of basiparameres and proximal to apex of aedeagus ( Figs 39–40 )........... R. schichai sp. nov.

Figures

FIGURES 1 – 4. Rugosothynnus burnsi, male. 1. habitus; 2. face; 3. head and pronotum, dorsal view; 4. apex of metasoma, dorsal view.

FIGURES 44 – 45. Rugosothynnus spp. 44. R. depressus sp. nov., face, male; 45. Ru fulvescens sp. nov., abdomen, female, dorsal.

FIGURES 46 – 49. Rugosothynnus spp. 46. R. houstoni sp. nov., apex of metasoma, male, lateral, open with parameres partially exposed; 47 – 48 R. depressus sp. nov., S 1 – 2, male: 47 ventral, 48 lateral; 49. R. tristis, S 1 – 2, male, lateral, scale line = 0.2 mm.

FIGURES 11 – 13. Rugosothynnus brunneus sp. nov. 11. genitalia, male, right lateral view; 12. genitalia, male, dorsal view; 13. pygidium, female, dorsal view. Scale lines = 0.5 mm.

FIGURES 29 – 31. Rugosothynnus fulvescens sp. nov. 29. genitalia, male, right lateral view; 30. genitalia, male, dorsal view; 31. pygidium, female, dorsal view. Scale lines = 0.5 mm.

FIGURES 18 – 21. Rugosothynnus spp. 18. R. clypeatus sp. nov., genitalia, male, right lateral view; 19. R. clypeatus sp. nov., genitalia, male, dorsal view; 20. R. confusus sp. nov., genitalia, male, right lateral view; 21. R. confusus sp. nov., genitalia, male, dorsal view. Scale lines = 0.5 mm.

FIGURES 39 – 42. Rugosothynnus spp. 39. R. schichai sp. nov., genitalia, male, right lateral view; 40. R. schichai sp. nov., genitalia, male, dorsal view; 41. R. tristis, genitalia, male, right lateral view; 42. R. tristis, genitalia, male, dorsal view. Scale lines = 0.5 mm.