Ctenocheles madagascariensis Secrétan, 1964

Charbonnier, Sylvain, Garassino, Alessandro & Pasini, Giovanni, 2012, Revision of Mesozoic decapod crustaceans from Madagascar, Geodiversitas 34 (2), pp. 313-357 : 313-357

publication ID	https://doi.org/ 10.5252/g2012n2a5
persistent identifier	https://treatment.plazi.org/id/143B927A-FF90-FFC7-FF24-FF0106D212F7
treatment provided by	Marcus
scientific name	Ctenocheles madagascariensis Secrétan, 1964
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Treatment

Ctenocheles madagascariensis Secrétan, 1964 ( Fig. 18 B-D)

Ctenocheles madagascariensis Secrétan, 1964: 149-152 , pl. 19, figs 1-4, 11-14; pl. 20, figs 9-16. — Collignon 1970b: 35. — Förster & Mundlos 1982: 154. — Feldmann et al. 1995: 8. — Feldmann et al. 2010: 340. — Schweitzer et al. 2010: 40.

TYPE MATERIAL. — Albian-Cenomanian of Antsatramahavelona: 8 syntypes ( MNHN.F.A33425, R 03931, R 03932, R 03933, gisement 117, grès à Stoliczkaia ).

Middle Santonian of the “Coupe de Ampolypoly-Antsirasira-Behamotra”: 5 syntypes ( MNHN.F. A33427 View Materials , A33429 View Materials , gisement 280, Texanites hourcqi zone).

Lower Campanian of the “Coupe de Bevaho”: 11 syntypes ( MNHN.F. A33437 View Materials , R 03881, gisement 251, Anapachydiscus wittekindi and Eulopoceras jacobi zone).

Lower Campanian of the “Coupe de Berere”: 5 syntypes ( MNHN.F.A33421, gisement 163) ; 18 syntypes ( MNHN.F. A33359 View Materials , A33424, gisement 166) ; 4 syntypes ( MNHN.F.A33423, A33424, gisement 167) ; 4 syntypes ( MNHN.F. A33376 View Materials , R03880, gisement 171, Karapadites karapadensis zone) .

Lower Campanian of the “Coupe de Ampolypoly-Antsirasira-Behamotra”: 82 syntypes ( MNHN.F.A33398, A33399 View Materials , A33400, A33401, gisement 302, Menabites boulei and Anapachydiscus arrialoorensis zone); 25 syntypes ( MNHN.F.A33419, gisement 317).

Middle Campanian of the “Coupe de Ampolypoly- Antsirasira-Behamotra”: 40 syntypes ( MNHN.F. A33417 View Materials , A33418, gisement 317).

Middle Campanian of the “Coupe de Andimaka”: 4 syntypes ( MNHN.F. R 03882, A33110 View Materials , A33215, A33374, gisement 203, Pachydiscus grossouvrei zone).

Middle Campanian of the “Coupe de Ankilizato”: 43 syntypes ( MNHN.F.A33407, A33408, A33409, gisements 153-155, Pachydiscus grossouvrei zone); 1 syntype ( MNHN.F. A33433 View Materials , gisement 181, Pachydiscus grossouvrei zone); 85 syntypes ( MNHN.F.A33391, A33392, A33393, A33394, A33395, A33396, A33397, gisement 154, Pachydiscus grossouvrei zone); 32 syntypes ( MNHN.F.A33330, A33406, A33381, gisement 155, Pachydiscus grossouvrei zone); 25 syntypes ( MNHN.F.A33406, A33381, A33405, gisement 159, Delawarella subdelawarensis and Australiella australis zone); 25 syntypes ( MNHN.F.A33358, gisement 177, Pachydiscus grossouvrei zone); 29 syntypes ( MNHN.F.A33410, A33411 View Materials gisement 179, Pachydiscus grossouvrei zone); 6 syntypes ( MNHN.F.A 33344, R 03883, gisement 181, Pachydiscus grossouvrei zone).

LOCALITIES. — Antsatramahavelona (northern Ananalava, Peninsula of Radama, northern Mahajanga Basin, Mahajanga Province); Coupes de Bevaho, Ampolypoly- Antsirasira-Behamotra, Berere Andimaka, Ankilizato (Belo-sur-Thsiribihina, Menabe region, central Morondava Basin, Tulear Province).

AGES. — Albian-Cenomanian to Middle Campanian.

ADDITIONAL MATERIAL EXAMINED. — Menabe region, central Morondava Basin. From the middle Campanian of the “Coupe de Ankilizato”: 11 specimens [MNHN.F.A33353, A33354, A33388 View Materials , A33436, gisements 176, 180 ( Pachydiscus grossouvrei zone), 189].

“Coupe de Berere”: 156 specimens (MNHN.F.A33345, A33346, A33350, A33356, A33367 View Materials , A33402 View Materials , A33403, A33404, A33420, A33430, A33435 View Materials , gisements 161, 162 [basal Lower Campanian, Anapachydiscus wittekindi and Eulophoceras jacobi zone], 184 [Lower Campanian, Menabites boulei and Anapachydiscus arrialoorensis zone], 200, 208 [Middle-Upper Santonian, Texanites hourcqi zone]).

From the Upper Turonian of the “Coupe de Berere”: 9 specimens (MNHN.F.A33428, gisement 246, [Masiaposa], Coilopoceras requieni and Romaniceras deveriai zone). From the Middle Campanian of the “Coupe de Andimaka”: 4 specimens (MNHN.F.A33175, A33351, A33380, gisements 201, 202, Delawarella subdelawarella and Australiella australis zone).

Lower Campanian of the “Coupe de Ampolypoly-Antsirasira-Behamotra”: 21 specimens (MNHN.F. A33217 View Materials , A33422, A33426, A33431 View Materials , A33438, gisements 283, 285, 287 [ Anapachydiscus wittekindi and Eulophoceras jacobi zone], 319, 322 [ Anapachydiscus wittekindi and Eulophoceras jacobi zone]).

Middle Campanian of the “Coupe de Ampolypoly- Antsirasira-Behamotra”: 5 specimens (MNHN.F.A33434, gisement 330).

Lower Campanian of the “Coupe de Bevaho”: 1 specimen (MNHN.F. A33377 View Materials , gisement 261, Karapadites karapadensis zone).

From the Middle Coniacian of Analabe (Belo-sur-Tsiribihina): 17 specimens (MNHN.F.A33432, gisement 344, Kossmaticeras theobaldi and Barroisiceras onilayense zone).

Sikily region, Tulear Province, southern Morondava Basin. Lower Maastrichtian of Soatana (Manera): 1 specimen (MNHN.F. A33225 View Materials ).

Northern Manambolo River, southern Maintirano region, northern Morondava Basin. Lower Campanian of Trangahy: 92 specimens (MNHN.F.A33218, A33338, Antsalova, gisement 240, Hoplitoplacenticeras marroti zone).

Uncertain localities and stratigraphic levels. 20 specimens (MNHN.F.A33221).

STRATIGRAPHIC RANGE. — Albian, Cenomanian, Late Turonian, Middle Coniacian, Middle Santonian, Early to Middle Campanian and Early Maastrichtian.

DISCUSSION

Upon re-examination of the Malagasy material studied by Secrétan (1964), we agree with her generic attribution. As noted by Secrétan (1964), the preservational state of the specimens studied allows confirmation of the presence of two different propodus types: one globose with a very elongate index and the other flat with an unknown index. Dactyli invariably are always fragmentary. However, because two kinds of propodus are typical of Ctenocheles , the lack of diagnostic characters, such as the number of teeth along the occlusal margins of specimens studied precludes a detailed diagnosis of the Malagasy species and a comparison with other Cretaceous forms. Although C. madagascariensis is still considered to be the oldest fossil species known to date ( Schweitzer & Feldmann 2002), only the discovery of more complete specimens can refine its morphological features.