Alexandre P. Marceniuk & Naércio A. Menezes, 2007, Systematics of the family Ariidae (Ostariophysi, Siluriformes), with a redefinition of the genera., Zootaxa 1416, pp. 1-126: 7

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Family Ariidae  Bleeker, 1862

Arii Bleeker, 1862: 7, 25. Type genus: Arius  ZBK  Valenciennes in Cuvier & Valenciennes, 1840b.

Hemipimelodinae  Gill, 1861: 46. Type genus: Hemipimelodus  ZBK  Bleeker, 1858.

Batrachocephalinae  Gill, 1893: 132. Type genus: Batrachocephalus  ZBK  Bleeker, 1846.

Doiichthyidae  Weber, 1913: 532. Type genus: Doiichthys  ZBK  Weber, 1913.

Bagreidae  Schultz, 1944: 182. Type genus: Bagre  Oken, 1817.

Osteogeneiosinae  Fowler, 1951: 3. Type genus: Osteogeneiosus  ZBK  Bleeker, 1846.

Vorhisiidae  † Frizzell, 1965: 179. Type genus: Vorhisia  † Frizzell, 1965.

Diagnosis. The family Ariidae  can be distinguished from all other Siluriformes by the following characters: external posterior branch of lateral ethmoid columnar; a bony blade anteriorly connecting the nasal tubules; lateral ethmoid and frontal bones connected mesially and laterally delimiting a fontanel (fig. 1) (shared with Pangasiidae  and Schilbeidae  ); presence of three infraorbitals (fig. 2) (shared with Doradidae  , Mochokidae  and Pimelodidae  ); lachrymal well developed (fig. 3); space between transcapular process and otic capsule small; otic capsules enlarged; wing process of parasphenoid present (shared with Bagridae  and Schilbeidae  ); subvertebral process well developed (fig. 4); anterior portion of second basibranchial expanded and very conspicuous (fig. 5); third pharyngobranchial boomerang shaped (fig. 6); anterior portion of proximal cartilage of fourth ceratobranchial narrow about one-half as wide as posterior portion; dorsal processes of pharyngeal tooth plates long (fig. 7); anterior and posterior nostrils close together (shared with Diplomystidae  ).

Remarks. To be consistent with the results of the phylogenetic analysis reported by Marceniuk (2003) in order to accommodate all the names available without creating unnecessary nomenclatural conflicts, the following genera are recognized: Amissidens  ZBK  , Arius  ZBK  , Aspistor  ZBK  , Bagre  , Batrachocephalus  ZBK  , Brustiarius  ZBK  , Cathorops  , Cephalocassis  ZBK  , Cinetodus  ZBK  , Cochlefelis  ZBK  , Cryptarius  ZBK  , Doiichthys  ZBK  , Galeichthys  ZBK  , Genidens  ZBK  , Hemiarius  ZBK  , Ketengus  ZBK  , Nedystoma  ZBK  , Nemapteryx  ZBK  , Neoarius  ZBK  , Netuma  , Notarius  ZBK  , Osteogeneiosus  ZBK  , Pachyula  ZBK  , Plicofollis  ZBK  , Potamarius  ZBK  and Sciades  . Three new genera have to be erected: Amphiarius  , Carlarius  and Potamosilurus  . The nominal genera Bagre  Oken, 1817, Glanis  ZBK  , Stearopterus  ZBK  , Breviceps  ZBK  non Merrem, 1820, Felichthys  ZBK  , Ailurichthys  ZBK  , Mystus  ZBK  non Scopoli, 1777 and Anemanotus  ZBK  are junior synonyms of Bagre  Cloquet, 1816. Hexanematichthys  ZBK  , Sciadeichthys  ZBK  , Selenaspis  ZBK  , Ariopsis  ZBK  and Leptarius  ZBK  are synonymous with Sciades  . The nominal genus Pseudarius  ZBK  is an objective synonym and Arioides  is subjective synonym of Arius  ZBK  . Sciadeops  ZBK  is synonymous with Notarius  ZBK  . Catastoma  ZBK  and Sarcogenys  ZBK  are synonymous with Netuma  . Tetranesodon  ZBK  is tentatively recognized as synonym of Pachyula  ZBK  . Hemipimelodus  ZBK  is synonymous with Cephalocassis  ZBK  . Septobranchus  ZBK  is a junior synonym of Cinetodus  ZBK  . Guiritinga  ZBK  is a new synonym of Genidens  ZBK  .

The genus Doiichthys  ZBK  formerly considered the single representative of the family Doiichthyidae  by Weber & de Beaufort (1913) and Berg (1940) and not examined either by Mo (1991) or de Pinna (1993), shares a series of synapomorphies with the other members of the Ariidae  (Kailola, 1990a, 2004; Marceniuk, 2003) and is included in this family. The nominal genus Tachysurus  ZBK  ( Lacépède, 1803) as already noted by Regan (1907) cannot be recognized in the Ariidae  contrary to the conclusion of Eigenmann & Eigenmann (1888), Miranda-Ribeiro (1918), Fowler (1928) and Chandy (1953) as emphasized by Wheeler & Baddokwaya (1981). Following Mo (1991) and de Pinna (1993), Ancharius  ZBK  is not considered a member of the Ariidae  . It belongs in its own family Anchariidae  (Ng & Sparks, 2005).













Alexandre P. Marceniuk & Naércio A. Menezes 2007


Frizzell 1965