Pseudopterogramma fenestrata ( Richards, 1973 ) Kuwahara & Marshall, 2022

Kuwahara, Gregory K. & Marshall, Stephen A., 2022, A revision of the Australian species of Howickia Richards (Diptera: Sphaeroceridae: Limosininae), Zootaxa 5192 (1), pp. 1-152 : 9

Pseudopterogramma fenestrata ( Richards, 1973) NEW COMBINATION

Leptocera (Limosina) fenestrata Richards, 1973: 379 .

[As an unplaced species of Limosininae ] Marshall, 1989: 607.

Biroina fenestrata (Richards) , Roháček et al., 2001: 125.

Comments. When Richards (1973) originally described this species in the (then) subgenus Limosina Macquart, 1835 , he noted that P. fenestrata resembled Pterogramma Spuler, 1924 , as it has patterned wings with a short R 2+3. In the catalog of Australian/Oceanic Sphaeroceridae, Marshall (1989) disagreed with the placement of P. fenestrata as a species of Limosina , but was unable to place the species in any of the known genera at that time. In the world catalog of Sphaeroceridae, Roháček et al. (2001) moved P. fenestrata into Biroina , though they noted that this generic placement was tentative and in need of confirmation.

This species is here moved to Pseudopterogramma on the basis of several characters, most notably the two pairs of large dorsocentral bristles, with the anterior pair located on the transverse suture and almost as long as the posterior pair. Other characters shared between Pseudopterogramma fenestrata and other Pseudopterogramma include patterned wings, a short sinuate R 2+3, costa ending at R 4+5, small (1.2–1.4 mm) body size, mid tibia with two pairs of dorsal setae (at the basal third and apical quarter), banded legs, and a simple hypandrium (lacking the ventral lobe and posterodorsal apodeme found in Howickia ). In Howickia , the dorsocentral bristles are all postsutural, with the anterior pair always at most half the size of the posterior pair. R 2+3 can be short in Howickia but in these species the wing is always infuscate but not distinctly patterned.