Paramacroxiphus securiformis, Ingrisch, 2008

Ingrisch, Sigfrid, 2008, Revision of the genera Paramacroxiphus C. Willemse 1961 and Pseudomacroxiphus C. Willemse 1961 (Orthoptera: Tettigoniidae: Conocephalinae: Agraeciini), Zootaxa 1755 (1), pp. 1-34 : 22-27

Paramacroxiphus securiformis sp. n.

Figs. 10 , 26–28 , 38 , 60–62 , 74–75 , 95

Holotype (male): Indonesia: Papua, Dutch New Guinea, Maffin Bay , 12.vi.1944, E.S. Ross . Holotype in Collection of Fer Willemse , Eygelshoven, Netherlands [later to be deposited in RMNH] .

Specimen examined (Allotype): 1 female, same data as holotype, 1.ix.1944 (CW).

Description. Fastigium verticis: eye diameter 1.4: 2.1 mm; projecting 0.8 mm in front of eyes. Femora with the following number of spines on ventral margins: profemur 7–9 external, 6–7 internal; mesofemur 7 external, 3–4 internal; postfemur 13–15 external, 15–18 internal.

Male. Stridulatory file 3.11 mm long, slightly sinuate and with large teeth in basal half, curved in about middle and with narrow teeth in apical half ( Fig. 10 ); with about 117 teeth or 37.6 teeth per mm; in middle 35.9 teeth per mm, in basal half 20.4 teeth per mm. Mirror quadrate, apex substraight, 1.74 mm long and wide. Tenth abdominal tergite globular and with central area prolonged behind; prolongation furrowed in middle, setose and with sloping lateral areas; apex faintly sinuate ( Fig. 28 ). Epiproct triangular. Cerci slightly curved; with a large dorso-internal tooth behind middle, which has a very broad base and acute apex; behind that tooth, cercus dorso-ventrally compressed and curved mediad; apex broad with an acute spine at posterior angle and a rounded lobe at anterior angle ( Figs. 26–27 ). Subgenital plate greatly narrowed behind basal area into a long medial projection; projection with almost parallel margins, little converging in apical area, and with broad obtuse lateral carinae, divided from apex for slightly less than half of its length; styli small ( Fig. 38 ).

Titillators separate, elongate, basal parts simple; apical parts subhyaline with broad brown band of stronger sclerotisation in midline; two-branched: lateral branch with narrow stiffened rim becoming conical at apex; apical branch with broad stiffened rim with little undulating margin and a short subapical cone ( Figs. 60–62 ). Apico-lateral sclerites small, rounded. Baso-lateral sclerites large, elongate with undulating margin.

Female. Subgenital plate transverse, narrowly divided in midline; with a membranous apical lobe that terminates into two long, acute, stiffened spines; lateral areas strongly curved dorsal ( Figs. 74–75 ). Very base of ventral valves of ovipositor above lateral apices of subgenital plate with a rather large pit ( Fig. 75 ).

Coloration. Uniformly ochre (formerly conserved in alcohol). Mandibles medium to dark brown; clypeus yellow. Female subgenital plate dark brown, apical prolongation light ochre. Ovipositor brown.

Measurements: body male 29, female 38; pronotum male 10.3, female 10.7; tegmen male 38, female 47; postfemur male 23, female 26; ovipositor 52; -height 3.5 mm.

Diagnosis. The new species is most closely related to P.rufus . It differs by the male tenth abdominal tergite having shorter apical projections that are not deviating nor widening towards apex; the male cerci have shorter apical teeth; the male subgenital plate has the long projection with parallel margins; the titillators differ in details especially that the main branch of the apical parts is weak but distinctly sclerotised and the apical cap has an obtuse projection; the female subgenital plate has the apical projections straight and approaching each other and the baso-lateral grooves of different shapes.

Etymology. The new species is named for the hatchet-shaped male cerci.

Figures

FIGURES 7–10. Stridulatory files on underside of left tegmen of Paramacroxiphus males: 7, P.aberrans paratype; 8, P. rufus holotype; 9, P. maculatus holotype; 10, P. securiformis holotype.

FIGURES 16–28. Male cercus and tenth abdominal tergite of Paramacroxiphus species: 16, 19, 22, 26, cercus in dorsal view; 17, 20, 23, do. ventral view; 24, 27, do. internal view; 18, 21, 25, 28, male tenth abdominal tergite in dorso-apical (18) or dorsal view (21, 25, 28).—16–18, P.aberrans holotype; 19–21, P.rufus holotype; 22–25, P. maculatus holotype; 26–28, P. securiformis holotype.

FIGURES 35–44. Male subgenital plate (35–43) and stridulatory area of right tegmen (44) of Paramacroxiphus species: 35, P. elongatus holotype; 36, P. irregularius holotype; 37, P.rufus holotype; 38, P. securiformis holotype; 39, P.armatus holotype; 40, P. bifasciatus holotype; 41, P. maculatus holotype; 42, P. aberrans holotype; 43, P. brunneus holotype; 44, P.aberransparatype. Abbreviations: m mirror, s scrapper, sp field of spinules.

FIGURES 57–66. Male titillators of Paramacroxiphus species in ventro-distal view (57, 60, 63, 65), dorso-proximal view (58, 61, 64, 66), lateral view (59), and view on apex (62): 57–59, P. aberrans holotype; 60–62, P. securiformis holotype; 63–64, P. maculatus holotype; 65–66, P. rufus holotype. Abbreviations: ap apical part of titillator; as apico-lateral sclerite; bp basal part of titillator; bs baso-lateral sclerite. Scale bars = 1 mm.

FIGURES 67–79. Female subgenital plate of Paramacroxiphus species in ventral view (67, 69, 70, 72, 74, 76, 78), lateral view (68, 71, 73, 75, 77) or recurved (79): 67–68, P. aberrans allotype (67) and paratype (68); 69, P. tessellatus holotype [the white arrow points at the apical cones]; 70–71, P. maculatus allotype; 72–73, P. uniformis holotype; 74–75, P. securiformis allotype; 76–77, P.rufus allotype (76) and paratype (77); 78, P.armatus paratype; 79, P. brunneus allotype. Abbreviations: ac apical cone; m membranous area; po pits at very base of ovipositor; ps pits at lateral area of subgenital plate; sd inner [= dorsal] surface of recurved subgenital plate; vv base of ventral valves of ovipositor usually hidden under subgenital plate.