Jethsura rubricauda, Claridge, 2021

Claridge, Brandon, 2021, Revision of the genus Jethsura Cameron, 1902 (Hymenoptera: Ichneumonidae: Ichneumoninae: Phaeogenini), Zootaxa 5071 (2), pp. 223-241 : 234-235

Jethsura rubricauda sp. nov.

Figs 7–8 , 11

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Diagnosis. Easily differentiated from other known Jethsura species by the black head and mesosoma and brownish-red metasoma.

Description. Female ( Fig. 7A–H ). Body length: 8.0– 8.1 mm; fore wing length: 4.8–4.9 mm.

Color. Black head and mesosoma with limited yellowish-white to yellowish marks and brownish-red metasoma. Head black, except for mandible dark brownish-red with darker base and paraocular area yellowish-white medially from antenatal base to median ocellus and posterodorsal corner, in one specimen the left side with paraocular area venteromedially ventral to antennal base yellowish and venterolateral corner with faint impression of yellowish mark. Antenna black. Mesosoma black, except for the following areas yellowish-white: medial 0.3 of ventral margin of pronotum, posterior 0.8 of dorsal margin of pronotum, tegula, and small mark on subalar ridge. Legs: coxae, trochanters, and trochantelli dark brown to black; fore femur anteriorly with basal 0.3 dark brown and apical 0.7 brownish-yellow, posteriorly dark brown; mid and hind femur dark brown except, for brown apex; fore and mid tibiae anteriorly brownish-yellow, posteriorly yellowish-brown to brown; hind tibia reddish-brown at base becoming dark brown apically; tarsi with tarsomeres 1–4 reddish-brown and tarsomere 5 dark reddish-brown, tarsomeres 1–4 of hind tarsus lighter basally. Metasoma reddish-brown, except for holotype with large mottled brownish mark on T2. Wing: membrane clear; basal 0.1–0.3 of wing with veins white, remaining sections brown.

Head. Slightly tapering ventrally. Mandible moderately wide (MI: ~3.1×). Clypeus smooth and coarsely punctate with punctures separated by 0.5–1.5× their diameter. Supraclypeal area densely punctate with punctures adjacent forming transverse rugulae dorsally, punctures becoming coarser and sparser laterally. Gena smooth and finely punctate with punctures separated by 2.0–4.0× their diameter, denser posteriorly. Supra-antennal are smooth and moderately punctate with punctures separated by 0.5–1.5× their diameter. Vertex smooth and finely punctate with punctures separated by 1.0–3.0× their diameter, denser and coarser medially. Hypostomal and occipital carinae meeting immediately before mandibular base. Antenna with 26 flagellomeres.

Mesosoma . Of normal width, not laterally flattened. Pronotum smooth and moderately punctate with punctures sparser anterodorsally, coarser ventrally. Propleuron smooth and finely, densely punctate with punctures separated by 0.5–1.0× their diameter. Mesonotum smooth and sparsely punctate with punctures separated by 1.0–3.0× their diameter, anterior 0.1 with punctation denser. Scutellum smooth and sparely punctate with punctures separated by 2.0–4.0× their diameter. Mesopleuron smooth and densely punctate, dorsal 0.3 with punctures separated by 0.5– 1.0× their diameter. Speculum finely and sparely punctate, ventral 0.6 denser with punctures adjacent to subadjacent and some confluent forming longitudinal rugulae. Ventral division of metapleuron smooth coarsely, densely punctate with punctures subadjacent, posteriorly some punctures confluent and forming rugulae. Propodeum: of normal width, nearly equal in length and width; dorsal face moderately convex, surface sculpture smooth and coarsely, densely punctate laterally with punctures adjacent and some confluent forming rugulae, especially posteriorly, punctation sparser medially; posterior face punctate-rugose; carinae obsolete, except for pleural carina posterior transverse carina well-developed, median longitudinal carina posterior to posterior transverse carina subobsolete. Legs of normal length. Hind femur normal, not anteroposteriorly flattened.

Metasoma. MS1 smooth, postpetiole moderately, coarsely punctate laterally and impunctate to sparsely punctate medially. T2 smooth and finely to moderately punctate with punctures separated by 1.0–4.0× their diameter, sparser posteriorly. Gastrocoelus shallow, subobsolete. Thyridium essentially obsolete, only indicated by faint impression; narrow, 0.2–0.3× as wide as interthyridial width; distant from T2 anterior margin by 0.2× tergite length. T3 smooth and finely, sparsely punctate. T4–7 smooth and nearly impunctate except for very fine, shallow, sparse punctures.

Male ( Fig. 8A–C ). Body length: 8.0 mm; fore wing length: 5.5 mm. As in female, except for: yellowish-white or yellowish areas on head and mesosoma of female white in male; white marks on paraocular areas posteroventrally and posterodorsally; MS1 smooth and finely, densely punctate with punctation on postpetiole sparser medially; T2–7 finely, densely punctate with punctures separated by 0.5–1.0× their diameter. Antenna with 29 flagellomeres.

Material examined. Holotype: Mexico • 1♀; Durango, 10 mi W El Salto ; 9000 ft.; 12.iv.1964; W. R. M. Mason; CNCI.

Paratypes: Mexico • 1♀; same collection data as holotype; 23.vi.1964 • 1♂; same collection data as holotype; 07.vi.1965 .

Distribution. Known only from type locality ( Fig. 11 ).

Etymology. Formed by rubri- (Latin) meaning red and cauda (Latin) meaning tail and refers to the distinctive red metasoma.

Figures

FIGURE 7. Jethsura rubricauda sp. nov. female holotype. A. Habitus. B. Mesosoma, lateral view. C. Head, frontal view. D. Mandible, ventral view. E. Propodeum, dorsolateral view. F. MS1–2, dorsolateral view. G. Metasomal apex, lateral view. H. Metasoma, dorsal view. Scale bars: 5.0 mm (A); 1.0 mm (B–H).

FIGURE 8. Jethsura rubricauda sp. nov. male paratype. A. Habitus. B. Head, frontal view. C. Mandible, ventral view. Scale bars: 5.0 mm (A); 1.0 mm (B–C).

FIGURE 11. Distribution of Jethsura pyriformis (Provancher) and Jethsura rubricauda sp. nov.