Physodera Eschscholtz

Ma, Yunlong, Shi, Hongliang & Liang, Hongbin, 2017, Revision of the Oriental Genus Physodera Eschscholtz, 1829 (Coleoptera, Carabidae, Lebiini, Physoderina), with the descriptions of two new species, Zootaxa 4243 (2), pp. 297-328 : 298

Key to species of Physodera Eschscholtz

1. Pronotum bicolor, dark with distinct light pattern; sternum and tergum VII blackish, both with striking yellowish pattern ( Figs. 45–47)..............................................................................................2

- Pronotum unicolor , sometimes lateral expansion a little lighter, but never forming distinct pattern; sternum and tergum VII usually uniformly blackish, sometimes only tergum VII with vague yellowish pattern around spiracles (Fig. 48)...........6

2. Pronotum light pattern plain, not forming callosity; elytra dark with light pattern on disc or apex ( amplicollis -group).......3

- Pronotum each side with an ivory callosity ( Figs. 29, 30 ); elytra unicolor ( dejeani -group)............................5

3. Elytra with large orange-yellow zigzag pattern on disc ( Figs. 27, 28 ); interval punctures dense and same size as stria punctures, strial puncture rows indistinct........................................................ P. bousqueti Mateu, 1990

- Elytra with narrow yellowish band at apex ( Figs. 13 , 25, 26 ); interval punctures much smaller than stria punctures, strial punc- ture rows distinct......................................................................................4

4. Pronotal hind angles fully round; pronotal lateral expansions yellow, without dark spot ( Fig. 13 ).. P. diglena Andrewes, 1930

- Pronotal hind angles more or less distinct; pronotal lateral expansions yellow to orange yellow, each side with a dark spot in ( Figs. 25, 26 )................................................................ P. amplicollis Van de Poll, 1889

5 Pronotal callosities large, distinctly larger than eyes; pronotum base shortly lobed, lateral notch (between posterior angle and basal lobe) width more than two times as length ( Fig. 52); elytral interval finely punctate; larger species, 9.5–10.5mm; widely distributed in Oriental Realm, except for Malay Peninsula and Borneo..................... P. dejeani Eschscholtz, 1829

- Pronotal callosities small, about same size as eyes; pronotum base strongly lobed, lateral notch width about same as length ( Fig. 53); elytral interval impunctate; smaller species, 8.5–9.0mm; Malay Peninsula and Borneo...................................................................................................... P. chalceres Andrewes, 1930

6. Elytra dark, disc with a pair of ivory callosities; antennomeres 5–11 distinctly widened (Fig. 59) ( eburata -group).........7

- Elytra without callosity, unicolor or bicolor; apical antennomeres not widened (Fig. 58)..............................9

7 Elytral callosity ovate, larger, length more than 0.4 times as elytral length ( Fig. 18 ); Borneo..... P. bifenestrata Heller, 1923

- Elytral callosity smaller, length about 0.2–0.25 times as elytral length............................................8

8. Elytral callosity nearly round, without distinct notch ( Fig. 19 ); third interval with five or six setigerous pores; fifth interval with three or four pores ( Fig. 62); Sulawesi..................................................... P. sciakyi sp. n.

- Elytral callosity distinctly notched posteriorly ( Fig. 17 ); third interval with two setigerous pores; fifth interval with one pore near base; the Philippines............................................................. P. eburata Heller, 1923

9. Elytra dark metallic, with two reddish spots near apex........................................................ 10

- Elytra uniformly metallic, without spot................................................................... 11

10. Head and pronotum reddish brown, elytra metallic blue; elytral red spots occupied intervals 1 to 6 ( Fig. 20 ); the Philippines............................................................................... P. andrewesi ( Jedlička, 1934)

- Head and pronotum dark brown, elytra metallic dark green; elytral red spots occupied intervals 2 to 4 ( Fig. 21 ); New Guinea, China (Xizang) . ................................................................ P. bacchusi Darlington, 1971

11. Pronotal lateral margins more or less angulate near middle ( Fig. 49); the fifth elytral interval with one large setigerous pore near base only................................................................... P. eschscholtzii Parry, 1849

- Pronotal lateral margins about evenly round near middle; the fifth elytral interval with three or more setigerous pores..... 12

12. Body form relative flat; pronotal hind angles blunt, not projected; elytral intervals with fine punctures; SE. China................................................................................................. P. unicolor sp. n.

- Body form strongly convex; pronotal hind angles about acute, a little projected laterally; elytral intervals almost impunctate; Sulawesi ................................................................... P. cyanipennis Van de Poll, 1889

Figures

FIGURES 25 – 30. Habitus, scale bars = 2 mm: Fig. 25, P. amplicollis van de Poll, Taiwan; Fig. 26, P. amplicollis van de Poll, Thailand; Fig. 27, P. bousqueti Mateu, Hainan; Fig. 28, P. bousqueti Mateu, Malaya; Fig. 29, P. dejeani Eschscholtz, Yunnan; Fig. 30, P. chalceres Andrewe s, Sabah

FIGURES 13 – 18. Habitus of type materials, scale bars = 2 mm: Fig. 13, holotype of P. diglena Andrewes; Fig. 14, syntype of P. dejeani Eschscholtz; Fig. 15, syntype of P. noctiluca Mohnike; Fig. 16, holotype of P. chalceres Andrewes; Fig. 17, lectotype of P. eburata Heller; Fig. 18, lectotype of P. bifnestrata Heller.

FIGURES 19 – 24. Habitus of type materials, scale bars = 2 mm: Fig. 19, holotype of P. sciakyi sp. n.; Fig. 20, holotype of Lachnoderma andrewesi Jedlička; Fig. 21, holotype of P. bacchusi Darlington; Fig. 22, holotype of P. unicolor, sp. n.; Fig. 23, syntype of P. eschscholtzii Dejean; Fig. 24, lectotype of P. davidis Fairmaire.