Physodera dejeani

Ma, Yunlong, Shi, Hongliang & Liang, Hongbin, 2017, Revision of the Oriental Genus Physodera Eschscholtz, 1829 (Coleoptera, Carabidae, Lebiini, Physoderina), with the descriptions of two new species, Zootaxa 4243 (2), pp. 297-328 : 301-302

dejeani -group

The dejeani -group contains two species: P. dejeani Eschscholtz and P. chalceres Andrewes.

The dejeani -group distributes in southeast Asia, including south Chinese continent, Taiwan Island, Indo-China Peninsula, northeastern India, Malay Peninsula, the Philippines, Borneo, Java, and Sumatra ( Fig. 64 ).

The diagnostic characters of this species group are: Pronotum with a pair of ivory callosities on each side; elytral dark purplish, without pattern; tergum and sternum VII dark, with yellowish patch on each side ( Fig. 47); elytral apex strongly curved; third interval with only two setigerous pores; terminal labial palpomeres tubiform in both sexes (Fig. 60); male mesotarsus with adhesive hairs rudimentary on the first two tarsomeres; males with one pair of setae on sternum VII. Aedeagus stout and straight, apical lamella very small; internal sac with main flagellum thick, reaching apical orifice; trumpet-form expansion large, length 0.4–0.45 times of the main flagellum; secondary flagellum and apical bursa absent ( Figs. 37 , 38 ).

These two species are doubtless closest in this genus. They seems to be relatively distant from other species groups for these following unique characters: (1) Male terminal labial palpomeres tubiform (securiform in other species groups); (2) elytral apex strongly curved (truncated in other species groups); (3) males with only one pair of setae on sternum VII (with two pairs in other species groups); (4) aedeagal secondary flagellum absent (present in other species groups); (5) apical segment of ovipositor slenderer than other species groups, length about 4 times as basal width (membranous extension excluded); length 2.5–3.4 times as basal width in other species groups ( Fig. 54–57).

Figures

FIGURE 37. Median lobe of male genitalia, right-lateral, ventral, left-lateral, and dorsal view of P. dejeani Eschscholtz, Yunnan, scale bar = 0.5 mm.

FIGURE 38. Median lobe of male genitalia, right-lateral, ventral, left-lateral, and dorsal view of P. chalceres Andrewes, a paratype from Perak, scale bar = 0.5 mm.

FIGURE 64. Known distribution of Physodera dejeani group: ● P. dejeani Eschscholtz; Ɨ P. chalceres Andrewes, type localities for valid names in red.