Calliax de Saint Laurent, 1973
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https://doi.org/ 10.5281/zenodo.5402949 |
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https://treatment.plazi.org/id/03FF2C63-FFAD-FFAB-2EF2-7A34FD87751B |
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Marcus |
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Calliax de Saint Laurent, 1973 |
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Genus Calliax de Saint Laurent, 1973
TYPE SPECIES. — Callianassa (Callichirus) lobata de Gaillande & Lagardère, 1966 , by original designation.
DIAGNOSIS. — Carapace lacking dorsal oval; rostrum short, with blunt tip, rostral spine absent. Second abdominal segment 2 longest, longer than sixth, no lateral tufts of setae on segments 3-5. Telson about 1.2 times as wide as long, lateral border curved, posterior border straight or slightly convex.
Eyestalk about twice as long as wide, slightly flattened dorso-ventrally; cornea small, weakly pigmented. A1 peduncle shorter than that of A2. Mxp1 epipod large, tapering anteriorly. Mxp2 with small, leaf-like epipod. Mxp3 subpediform, propodus and dactylus rounded, exopod absent.
P1 unequal, dissimilar; fixed finger as long as dactylus and terminal in major P1, shorter than dactylus and subterminal in minor P1, with wide proximal gap and large triangular proximal tooth on cutting edge. P3 with small proximal heel on propodus, P5 subchelate. Paired arthrobranch on Mxp3 and P1-4.
Male and female Plp1 uniramous, male and female Plp2 biramous, all lacking appendix interna, male Plp2 with appendix masculina overreaching endopod. Plp3- 5 biramous, foliaceous, appendix interna finger-like in both sexes. Uropodal endopod and exopod slightly longer than telson, with rounded posterior border; exopod with dorsal plate terminating in short distal setal row.
REMARKS
The genus Calliax was considered by Manning & Felder (1991: 781) as distinct from their newly established Eucalliax , but Sakai (1999a: 110) synonymised the two taxa. A number of Calliax and Eucalliax species have been described recently. Their examination confirms the validity of the latter genus and permits the former to be better defined.
Both Calliax and Eucalliax comprise species of Eucalliacinae having no exopod on the Mxp3. It is proposed that two of these are assigned to Calliax : Calliax lobata (de Gaillande & Lagardère, 1966) (type species) and Calliax doerjesti Sakai, 1999 .
A short diagnosis for the American species of Eucalliax was given by Manning & Felder (1991: 781). It is here amended after examination of new materials.
Rostrum triangular, rostral spine small or absent; cornea flatttened, almost terminal. A1 peduncle shorter than that of A2. Mxp3 without exopod, operculiform. Chelipeds equal and similar, major without meral hook. Male and female Plp1 slender and uniramous, lacking appendix interna; male and female Plp2 slender and biramous, male Plp2 with appendix interna and appendix masculina, female Plp2 often with appendix interna (absent in E. quadracuta ). Plp3- 5 foliaceous and biramous in both sexes, with finger-like appendix interna. Telson over 1.5 times as wide as long, posterior border straight or slightly concave, often with dorsal transverse carina (absent in E. bulimba ). Uropods longer than telson, endopod overreaching telson by at least half its length.
It can be noted that the Mxp3 figured by Manning & Felder (1991: fig. 3) and labeled as Eucalliax actually belongs to Calliax lobata as indicated in the legend.
The following species are placed in Eucalliax : Eucalliax quadracuta (Biffar, 1970) (type species), Eucalliax aequimana (Baker, 1907) n. comb., Eucalliax bulimba ( Poore & Griffin, 1979) n. comb., Eucalliax cearaensis Rodriguez & Manning, 1992 , Eucalliax jonesi ( Heard, 1989) , Eucalliax mcilhennyi Felder & Manning, 1994 .
Differences between Calliax and Eucalliax (figures from Felder & Manning 1994 for Eucalliax mcilhennyi ) species are: 1) A1 peduncle is distinctly shorter than that of A2, not overreaching the proximal border of the last article in Calliax (reaching mid-length of last article in Eucalliax ); 2) Mxp3 is subpediform in Calliax ( Fig. 17K, L View FIG ) (operculiform in Eucalliax [ Fig. 18P View FIG ]); 3) P1 is unequal; the minor with the fixed finger shorter than and separated from the dactylus by a wide gap, bearing a large triangular proximal tooth in Calliax ( Fig. 17D, E View FIG ) (P1 is subequal and similar in Eucalliax , Fig. 18M, N View FIG ); 4) male Plp2 has an appendix masculina ( Fig. 18D View FIG ), female Plp2 lacking an appendix interna in Calliax ( Fig. 18B View FIG ) (male Plp2 has both appendix interna and appendix masculina [ Fig. 18L View FIG ] female Plp2 often with appendix interna in Eucalliax , Fig. 18J View FIG ); 5) the telson is less than 1.5 times as wide as long with the posterior border straight or slightly convex and no dorsal transverse carina in Calliax ( Fig. 17J View FIG ) (the telson is over 1.5 times as wide as long with the posterior border straight or slightly concave with the dorsal transverse carina often present in Eucalliax [ Fig. 18O View FIG ]); and 6) the uropods are about as long or slightly longer than the telson in Calliax ( Fig. 17J View FIG ) (uropods are long- er with the endopod overreaching the telson by at least half of its length in Eucalliax [ Fig. 18O View FIG ]).
Calliax lobata (de Gaillande & Lagardère, 1966) ( Figs 17 View FIG ; 18 View FIG )
Callianassa (Callichirus) lobata de Gaillande &
Lagardère, 1966: 259, pls 1-4.
Calliax lobata – de Saint Laurent 1973: 514. — de
Saint Laurent & Božić 1976: 28, figs 7, 15, 23, 27,
34. — Thessalou-Legaki & Zenetos 1985: 311. —
°Thessalou-Legaki 1986: 183. — Manning 1987:
397. — Števčić 1985: 313; 1990: 218. — Manning &
Felder 1991: 783, figs 15f-j. — ° Froglia 1995: 7. —
Falciai & Minervini 1996: 145, 4 figs. — Sakai 1999a:
110, fig. 27a-c. — d’Udekem d’Acoz 1999: 155. —
Tudge et al. 2000: 145, figs 3, 4 (cladogram). —
° Türkay 2001: 289.
TYPE MATERIAL. — Holotype: #, Port-Miou, near
Toulon, France, de Gaillande coll., X.1964 ( MNHN
Th 1296); paratypes: # 1 (MNHN Th 1298), 1 ,
1 ( MNHN Th 84), 1, 1 ( RMNH D 23011 View Materials ) (# previously deposited in the Station marine d’Endoume, France).
MATERIAL EXAMINED. — France. Port-Miou , near Toulon, 5-10 m, de Gaillande coll., X.1964, holotype cl. 8mm (figured) (dissected by the authors) ( MNHN Th 1296) ; 1 paratype cl. 6 mm, tl. 21.5 mm, eyes and both Plp 1, Plp 2 lost ( MNHN Th 1298), 1 paratype cl. 5 mm, tl. 20 mm, 1 broken paratype (figured) ( MNHN Th 84), 1, 1 paratypes ( RMNH D 23011 View Materials ) ; same data, 1 cl. 7 mm, poor condition (figured) (# MNHN Th 1297) ; same locality, de Gaillande coll., 1972, 3 tl. 21- 25 mm, 1 tl. 25 mm ( MNHN Th 674) .
Croatia. Rovinj, Števčić coll., 1983, 1 cl. 4.5 mm, tl. 18 mm approx., poor condition ( MNHN Th 675). OTHER MATERIAL EXAMINED. — Eucalliax quadracuta (Biffar, 1970) , Venezuela, paratypes: 1 ovig.
( MCZ 12872a), 1, 1 ( MCZ 732). — Eucalliax bulimba ( Poore & Griffin, 1979) , Queensland, Australia, 1 ( MV J12184 View Materials ). — Eucalliax aequimana (Baker, 1907) , King George Sound, 1 ( MNHN Th 579).
DISTRIBUTION. — Mediterranean: Port-Miou (between Toulon and Marseille, France) (de Gaillande & Lagardère 1966; de Saint Laurent & Božić 1979); near Rovinj, Adriatic ( Števčić 1990), Aegean Sea (Thessalou-Legaki 1986).
DIAGNOSIS
As for the genus with details added: eyestalk ( Fig. 17C View FIG ) with cornea small, weakly pigmented, sometimes recognisable dorsodistally, according to de Saint Laurent & Božić (1976), indistinct at present in most material examined. Md ( Fig. 18E View FIG ) incisor process with triangular proximal and distal tooth separated by pectinate border. Mx1 ( Fig. 18F View FIG ) and Mx2 ( Fig. 18G View FIG ) as figured. Mxp3 ( Fig. 17K, L View FIG ) subpediform, ischium-merus length about 2.5 merus width, mesial ridge of ischium with eight or nine spinules; both propodus and dactylus rounded, propodus nearly as long as wide, dactylus more slender, about 1.3 as long as wide.
P1 unequal ( Fig. 17D, E View FIG ), minor subchelate; both major and minor P1 with spines on lower border of ischium; lower border of merus with spines in major P1, unarmed in minor P1, carpus and propodus unarmed in both; fixed finger of major P1 with large median tooth on cutting edge, fixed finger of minor P1 shorter than dactylus with large triangular proximal tooth. P2-P5 ( Fig. 17 View FIG F-I) as figured. Telson ( Fig. 17J View FIG ) with posterior border weakly convex. Male and female Plp1, Plp2 ( Fig. 18A, B and 18C, D View FIG ) as figured. Plp3-5 ( Fig. 18H View FIG ) with finger-like appendix interna, sitting side by side with mesial border of endopod. Uropodal exopod ( Fig. 17J View FIG ) with lateral notch near upper third of posterior border.
Colour
Whitish (de Gaillande & Lagardère 1966).
Size
Small: cl. 4.5- 8 mm, tl. 18-25 mm approximately.
ECOLOGY
Living in mud or silty mud, between 2 and 21 m (de Gaillande & Lagardère 1966; Števčić 1990).
REMARKS
There is probably an error in the shape of the eyestalks presented by de Saint Laurent & Božić (1976) for this species and reproduced here ( Fig. 17A View FIG ). The female figured, from the type locality near Toulon, is at present in the collection of the MNHN (MNHN Th 674). The eyestalks are not as wide as depicted (about 0.7 time as wide as long) nor are they as wide in the holotype ( Fig. 17C View FIG ) or paratypes (see Sakai 1999a: fig. 27a) (about 0.45-0.5 time as wide as long).
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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Calliax de Saint Laurent, 1973
| Ngoc-Ho, Nguyen 2003 |
Calliax lobata
| SAINT LAURENT M. & DE 1973: 514 |