Anatoma tobeyoides, Geiger & Jansen, 2004

Geiger, Daniel L. & Jansen, Patty, 2004, Revision of the Australian species of Anatomidae (Mollusca: Gastropoda: Vetigastropoda), Zootaxa 415 (415), pp. 1-35 : 21-23

Anatoma tobeyoides new species: Figures 10–12 , 18

Anatoma “East coast”: Jansen, 1999: 49, figs. 13–15.

Type material. HOLOTYPE AMS C.431072. 1.47 mm.

PARTAYPES: AMS C.29025, 8. AMS C.402671, 1. AMS C.402672, 1. AMS C.402673, 7. AMS C.402674, 1. AMS C.406337, 5.

Type locality. West of D’Entrecasteaux Channel , SE Tasmania, Australia, 43.045°S 147.347°E .

Etymology. The sculpture of fine, irregularly intersecting lines is reminiscent of the paintings of Mark Tobey (1890–1976: cf. Anonymous, 1997): tobey­: referring to the painters surname; ­oides: Latin: like, similar to.

Description. Shell globular, medium size to large (4.2 mm). Protoconch 3/4 whorls, flocculant sculpture, no apertural varix. Teleoconch I fewer than 0.5 whorls, no spiral cord in position of selenizone, fine centrifugal markings, strong axials present. Teleoconch II up to 2 1/4 whorls (1.6 mm shell). Sculpture with axials predominant, approximately 85 on body whorl on fully grown shell, approximately 15 spiral threads on shoulder at apertural margin. Approximately 25 spiral threads on base. Umbilicus continuously sloping from base, narrow, deep. Selenizone at periphery, keels distinct, growth marks poorly coordinated with axials. Aperture rounded, somewhat flared. Animal unknown.

Differential diagnosis. Anatoma tobeyoides n. sp. is most similar to A. australis . However, A. tobeyoides n. sp. lacks the protoconch varix found in A. australis , has a teleoconch I of less than half a whorl, whereas in A. australis it has more than half a whorl, A. tobeyoides n. sp. lacks a spiral cord on teleoconch I whereas A. australis has a prominent one, and A. tobeyoides n. sp. has much weaker spirals than axials and centrifugal spirals on teleoconch I and early teleoconch II, whereas A. australis has spirals and axials of equal strength and lacks the centrifugal spirals.

Distribution. South Australia, Tasmania (43.2°S) through Queensland (21.7°S), 27– 1330 m shells only, 154 m live.

Specimen Records. West of D’Entrecasteaux Channel , SE Tasmania, Australia, 43.045°S, 147.347°E (holotype AMS C.431072, seven paratypes AMS C.402673) .

QLD. 75 m, off Moreton Bay , 27.517S, 153.667E ( AMS C.402674, 1: paratype) . 77 m, off Moreton Bay , 27.456S, 153.65E ( AMS C.402724, 18). Amity Point, Stradbroke Is, Moreton Bay., 27.4S, 153.433E ( AMS C.402726, 1) . 27 m, GBR, Swain Reefs , Bylund Gillett Cay, 21.717S, 152.417E ( AMS C.402671, 1: paratype) . 64 m, GBR, Swain Reefs , 3 km NE of W side of Bylund Gillett Cay, 21.7S, 152.433E ( AMS C.402725, 7) .

NSW. 219 m, E of Brush Is , 35.433S, 150.633E ( AMS C.402672, 1: paratype) . 40 m, N of Sydney , 33.757S, 151.36E ( AMS C.402675, 1) . 45.5 m, off Port Stephens , 32.708S, 152.25E ( AMS C.402727, 1) . 73 m, 9 ml NE of Coffs Harbour , 30.25S, 153.317E ( AMS C.402730, 1) . 13 m, E of Yamba , 29.5S, 153.367E ( AMS C.402731, 1) . 86 m, off Ballina , 29.17S, 153.728E ( AMS C.402728, 1) . 103 m, off Ballina , 29.167S, 153.783E ( AMS C.402729, 1) . 154 m, Taupo Guyot, off Newcastle, Tasman Sea , 33.103S, 156.155E ( AMS C.404497, 1: complete) .

VIC. 2000 m, Bass Strait, 30 mls S of Cape Nelson, 38.958S, 141.542E ( AMS C.402739, 1) . 60 m, Bass Strait, ca 40 km S of Lakes Entrance , 38.317S, 147.917E ( AMS C.402740, 1) . 165 m, Bass Strait, ca. 27 mls SE of Cape Everard, 38.25S, 149.2E ( AMS C.402738, 8) . 75 m, Between Cape Howe & Lakes Entrance , 37.917S, 149E ( AMS C.404965, 1).

SA. 1330 m, Galathea station 554, Australian Bight, 3728’S 138°55’E ( ZMUC, 1) .

TAS. 212 m, S of Storm Bay , 43.783S, 147.808E ( AMS C.402636, 6) . 82 m, W of Port Davey , 43.338S, 145.803E ( AMS C.402734, 1) . 183 m, off Cape Pillar , 43.217S, 148.083E ( AMS C.29025, 8: paratypes) . 95 m, N of Cape Pillar, 43.167S, 148.023E ( AMS C.402732, 1) . 82.5 m, Maria Is , 2.5 ml NE Beaching Bay, 42.458S, 148.2E ( AMS C.402733, 2) . 205 m, off Cape Forestier., 42.167S, 148.578E ( AMS C.402737, 6) . 113 m, off Long Point , N of Bicheno, 41.758S, 148.517E ( AMS C.402735, 1; AMS C.406337, 5) . 88 m, S of West Point , 41.153S, 144.403E ( AMS C.402736, 1) . 399 m, off Cape Naturaliste , 40.843S, 148.775E ( AMS C.402626, 11) .

Figures

FIGURE 10. Anatoma turbinata. Protoconchs of A–C eroded, scale bars = 200 µm. A. AMS C.402663. Arafura Sea, ca. 330 km NW of Wessel Island, NT, Australia. B. AMS C.402662. Arafura Sea, ca 210 miles NE Croker Island, NT, Australia. C. AMS C.402665. North West Shelf, ca 100 miles NW of Broome, WA, Australia. D. Protoconch profile of A. E–F. AMS C.402648. 260 m, SE of Tongatapu, Tonga. Scale bars D–F= 50 µm.

FIGURE 11. Anatoma tobeyoides n. sp. Holotype AMS C.431072. W of D’Entrecasteaux Channel, TAS, Australia. 1.47 mm. Scale bar shell = 200 µm. Scale bars spire and protoconch = 50 µm.

FIGURE 12. Anatoma tobeyoides Paratypes. A. AMS C.29025. 183 m, off Cape Pillar, TAS, Australia. B. AMS C.402671. 27 m, GBR, Swain Reefs, Bylund Gillett Cay, QLD, Australia. C. AMS C.402672. 219 m, E of Brush Island, NSW, Australia. Scale bars shell = 200 µm. Scale bars spire and protoconch = 50 µm.

FIGURE 18. Distribution maps of Australian species of Anatomidae. Circle with star: type locality. Solid dots: specimen records. Squares: literature records.