Trachysphaera schmidtii Heller, 1858
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https://doi.org/10.11646/zootaxa.5047.3.3 |
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Trachysphaera schmidtii Heller, 1858 |
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5. Trachysphaera schmidtii Heller, 1858
Figs 1F, G View FIGURE 1 , 7–10 View FIGURE 7 View FIGURE 8 View FIGURE 9 View FIGURE 10 , 12A View FIGURE 12 .
Trachysphaera Schmidtii (sic!) Heller, 1858: 317, figs 1–6.
Trachysphaera schmidtii — Sillaber (1987: 179–188, figs 1–3, 5, 7, 9, 11–18); Tabacaru (1988: 53; 1990: 26–29, fig. 1).
Trachysphaera schmidti (sic!) — Thaler & Neuherz (1978: 34); Kime & Enghoff (2011: 38, 134).
Gervaisia noduligera Verhoeff, 1906: 812 View in CoL , 815, 819, figs 14, 17, 18 [synonymized by Sillaber (1987: 181)].
Gervaisia noduligera View in CoL —Verhoeff (1980: 531, 533, fig. 3); Schubart (1934: 54, 55, figs 73, 74); Attems (1929: 313; 1943: 79, 82–84, figs 9–13); Strasser (1958: 164, fig. 26).
Gervaisia noduligera noduligera View in CoL — Jermy (1942: 59, 60, 68, fig. 202).
Trachysphaera noduligera auct.— Thaler & Neuherz (1978: 33, figs 1, 3, 4, 10, 11).
Trachysphaera noduligera noduligera — Strasser (1971: 13).
Trachysphaera noduligera — Tabacaru (1988: 55).
Gervaisia noduligera styrica Verhoeff, 1906: 812 View in CoL , 819, figs 1, 2 [appeared in the synonym list of T. n. noduligera View in CoL by Strasser (1971: 13)].
Gervaisia noduligera var. styrica View in CoL — Verhoeff (1908: 532); Attems (1929: 313).
Trachysphaera noduligera styrica — Strasser (1971: 13).
Gervaisia noduligera hungarica Jermy (1942: 60 View in CoL , 68, 73, fig. 201).
Trachysphaera noduligera hungarica — Tabacaru (1988: 55).
Gervaisia illyrica Verhoeff, 1908: 531 View in CoL , 534, figs 2, 12, 13 [synonymized with a question mark (?) under noduligera by Strasser & Minelli (1984: 198), later appeared in the synonym lists of noduligera by Foddai et al. (1995: 12)].
Gervaisia illyrica View in CoL — Strasser (1938: 402, 403; 1962: 467); Attems (1929: 313; 1943: 79, 86; 1959: 334, figs 65–69); Jermy (1942: 64, 65).
Gervaisia noduligera var. illyrica View in CoL — Strasser (1959: 6, figs 2, 5).
Trachysphaera illyrica — Tabacaru (1988: 55).
Trachysphaera noduligera illyrica — Strasser (1971: 13).
Gervaisia costata var. acutula Latzel, 1884: 89 View in CoL , figs 41, 42, syn. n. [since probably most of the numerous subsequent identifications of acutula fail to concern Latzel’s (1884) acutula , but belong to acutula in the sense of Verhoeff (1898, 1899, 1906), here we refer only to the original name)].
Gervaisia multiclavigera Verhoeff, 1898: 163–165 View in CoL , figs 5–7, 9, 10, syn. n.
Gervaisia multiclavigera View in CoL — Verhoeff (1906: 810, 811, 819; 1908: 531); Attems (1929: 313; 1943: 79, 84, 85, figs 15, 16).
Gervaisia costata multiclavigera View in CoL — Verhoeff (1899: 185, 186, fig. 3).
Gervaisia multiclavigera declivis Verhoeff, 1906: 811 View in CoL , 819, figs 12, 13 [synonymized under multiclavigera View in CoL by Strasser (1971: 13)].
Gervaisia multiclavigera declivis View in CoL — Attems (1929: 313).
Trachysphaera multiclavigera — Strasser (1971: 13); Tabacaru (1988: 54); Kime & Enghoff (2011: 37, 131).
Gervaisia cultrifera Verhoeff, 1906: 810 View in CoL , 815, 816, 820, 821, syn. n.
Gervaisia cultrifera View in CoL — Verhoeff (1908: 530, fig. 5); Verhoeff (1910: 162); Attems (1943: 79, 86, fig. 20); Strasser (1958: 162, figs 23, 24).
Trachysphaera cultrifera — Strasser & Minelli (1984: 198); Tabacaru (1988: 54); Kime & Enghoff (2011: 36, 126).
Material examined. Serbia ( IZB): Mountain Fruška gora : 4 ♂♂, 6 ♀♀, 2 juveniles, Glavica , 25.03.2000, I. Kara- man leg .; 4 ♂♂, 1 ♀, 4 juveniles, Savina voda, Glavica , 05.03.2001, I. Karaman leg. ; 1 ♀, Stražilovo , 15.11.2011, D. Savić leg. ; 1 juvenile, Rakovački potok, 02.06.2018, D. Antić, S. Makarov leg. ; 2 ♂♂, 1 juvenile, Vrdnik , 02.06.2018, D. Antić, S. Makarov leg. ; 2 ♂♂, 3 ♀♀, same locality, but 01.03.2019, D. Antić, D. Stojanović, M. Šević, R. Beretić leg. ; 1 ♀, same locality, but 28.11.2019, D. Antić, D. Stojanović, S. Aničić leg. ; Zasavica : 1 ♂, Vrbovac, 2013, M. Stanković leg.; 1 ♂, 1 ♀, 1 juvenile, Trebljevine, Banovo Polje , 27.11.2019, D. Antić, D. Stojanović, S. Aničić, M. Šević leg. ; 2 ♀♀, same locality, but 13.03.2020, M. Stanković, M. Šević leg. ; Belgrade: 5 ♂♂, 7 ♀♀, 5 juveniles, Topčider, Milošev konak, 06.10.1992 ; 200 ex. (♂♂, ♀♀, juveniles), same locality, but 29.05.1993 ; 3 ♂♂, 4 ♀♀, Avala Mt., Čarapićev brest, 04.03.2021, D. Antić leg. ; Valjevo : 3 ♂♂, 3 ♀♀, 4 juveniles, Petnička Cave, village Petnica , 22.06.2001, S. Ćurčić leg. ; 5 ♂♂, 5 ♀♀, 1 juvenile, same locality, but 16.06., M. Petković leg. ; Kosjerić: 3 ♂♂, 1 juvenile, Filipići , 28.04.2000, I. Karaman leg. ; Miroč Mt.: 1 ♂, 2 ♀♀, Buronov Ponor Cave , 14.07.2002, S. Ognjenović leg. ; 1 juvenile, same locality, but 17.03.2015, D. Antić leg. ; Despotovac: 1 ♀, 4 juveniles, in front of the cave Bela Sala, Suvaja Canyon , 19.04.1997, S. Pešić leg. ; 1 ♂, 2 ♀♀, 2 juveniles, same locality, but 24.07.1997, I. Karaman leg. Bosnia and Herzegovina ( IZB) : 1 ♂, Snježnica Cave, near Jajce , 18.11.2012, T. Rađa leg. Slovenia ( IZB): 2 ♂♂, 4 ♀♀, 6 juveniles, Jama pri Metliki Pit , Metlika, 21.07.2015, Ž. Fišer, A. Croxeil, E. Premate, D. Škufca leg. ; 2 ♀♀, 1 juvenile, Hajdučka jama Pit, Bedenj , Črnomelj, 23.07.2015, Ž. Fišer, D. Škufca, E. Premate, A. Croxeil leg. ; 1 ♂, 1 juvenile, Džud Pit, Mali Nerajec , Črnomelj, 28.07.2015, Ž. Fišer, D. Škufca, E. Premate, A. Croxeil leg.
Attems, C. (1929) Die Myriopodenfauna von Albanien und Jugoslavien. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 56, 269 - 356.
Attems, C. (1943) Von der Gattung Gervaisia. Zoologischer Anzeiger, 143 (3 - 4), 76 - 89.
Foddai, D., Minelli, A., Scheller, U. & Zapparoli, M. (1995) Chilopoda, Diplopoda, Pauropoda, Symphyla. In: Minelli, A., Ruffo, S. & La Posta, S. (Eds.), Checklist delle Specie della Fauna Italiana, 32 - 33, pp. 1 - 35.
Heller, C. (1858) Beitrage zur osterreichischen Grotten-Fauna. Sitzungsberichte der Mathematisch-Naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften, Wien, 26 (1), 313 - 326.
Jermy, T. (1942) [Taxonomy of the Hungarian Plesiocerata (Diplopoda)]. Matematikai es termeszettudomanyi kozlemenyek, 39 (4), 1 - 82. [in Hungarian]
Kime, R. D. & Enghoff, H. (2011) Atlas of European millipedes (Class Diplopoda). Vol. 1. Pensoft Publ., Sofia, 282 pp.
Latzel, R. (1884) Die Myriopoden der Osterreichisch-ungarischen Monarchie. Zweite Halfte. Die Symphylen, Pauropoden und Diplopoden. Alfred Holder, Wien, 414 pp.
Schubart, O. (1934) Tausendfussler oder Myriapoda. Die Tierwelt Deutschlands und der angrenzenden Meeresteile nach ihren Merkmalen und nach ihrer Lebensweise. Teil 28. Gustav Fischer Verlag, Jena, 318 pp.
Sillaber, H. (1987) Zur Trachysphaera schmidtii in Karnten (Myriapoda, Diplopoda). Carinthia II, 177 (97), 179 - 188.
Strasser, K. (1938) Neue Diplopoden der Insel Cherso. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 71 (4 - 6), 389 - 407.
Strasser, K. (1959) Una Gervaisia troglobia del Veneto. Bollettino della Societa adriatica di scienze naturali in Trieste, 50, 1 - 11.
Strasser, K. (1971) Catalogus Faunae Jugoslaviae. III / 5. Diplopoda. Academia Scientarum et Artium Slovenica, Ljubljana, 48 pp.
Strasser, K. & Minelli, A. (1984) Elenco dei diplopodi d'Italia - A checklist of Italian diplopods. Societa Veneziana di Scienze Naturali Lavori, 9 (2), 193 - 212. [Stronczynski, K.] (1855) In: [Stronczynski, K., Taczanowski, W. & Waga, A.], Sprawozdanie z podrozy naturalistow odbytcj w r. 1854 do Ojcowa. Biblioteka Warszawska, 2, pp. 142 - 172.
Tabacaru, I. (1988) Sur le genre Trachysphaera Heller, 1858 (Diplopoda, Glomerida, Trachysphaeridae). 3 e partie. Travaux de l'Institut de Speologie Emile Racovitza , 27, 51 - 66.
Tabacaru, I. (1990) Le genre Trachysphaera Heller, 1858 (Diplopoda, Glomerida, Trachysphaeridae) dans les grottes de Roumanie. 2. Trachysphaera schmidtii Heller. Travaux de l'Institut de Speologie Emile Racovitza , 29, 25 - 31.
Thaler, K. & Neuherz, H. (1978) Uber zwei Hohlen-Diplopoden der Steiermark (Diplopoda, Chilognatha: Gervaisiidae und Attemsiidae). Mitteilungen der Abteilung fur Zoologie am Landesmuseum Joanneum in Graz, 7 (1), 31 - 38.
Verhoeff, K. W. (1898) Uber Diplopoden aus Bosnien, Herzegowina und Dalmatien. V. Theil: Glomeridae und Polyzoniidae (Schluss). Archiv fur Naturgeschichte, 64 (1), 161 - 176. https: // doi. org / 10.5962 / bhl. part. 6891
Verhoeff, K. W. (1899) Beitrage zur Kenntniss palaarktischer Myriopoden. IX. Aufsatz: Zur Systematik, Phylogenie und vergleichenden Morphologie der Juliden und uber einige andere Diplopoden. Archiv fur Naturgeschichte, 65 (1), 183 - 220.
Verhoeff, K. W. (1906) Uber Diplopoden. 5. (25.) Aufsatz. Zur Kenntnis der Gattung Gervaisia (Opisthandria). Zoologischer Anzeiger, 30 (24), 790 - 822.
Verhoeff, K. W. (1908) Uber Diplopoden. 9. (29.) Aufsatz: Gervaisia und Polyzonium. Zoologischer Anzeiger, 32 (18), 521 - 536.
Verhoeff, K. W. (1910) Aufsatz: Beitrage zur Kenntnis der Glomeriden, Juliden, Ascospermorpha (!) und Lysiopetaliden, sowie zur Fauna Siziliens, Untersuchungen uber Art- und Gruppensystematik; Morphologie, nachembryonale Entwicklung, Biologie und Geographie. Nova Acta. Abhandlungen der Kasiserl. Leop. - Carol. Deutschen Akademie der Naturforscher, 92 (2), 139 - 448.
FIGURE 1. Trachysphaera species in situ, Serbia (IZB). A–C. T. cristangula (Attems, 1943) from Mucina voda, near Ivanjica (A, B—♂, C—♀). D, E. T. lobotarsus (Attems, 1943) from Medvednik Mt. (D—♂, E—♂ and ♀). F, G. T. schmidtii Heller, 1858 (F—specimens from Avala Mt., G—♀ from Stražilovo, Fruška gora Mt.). H, I. T. similicostata (Radu & Ceuca, 1951) from Stara Planina Mt. (H—specimen from near Ponor, Dojkinci village, I—specimens from no-name cave I, Temska village). All pictures by Dragan Antić, except for G by Dragiša Savić.
FIGURE 7. Trachysphaera schmidtii Heller, 1858 from Serbia (IZB). A, C, E. Specimens from Petnička Cave. B, D. ♂ from Vrdnik, Fruška gora Mt. F–I. ♂ from Trebljevine, Zasavica. A. ♂, habitus, dorsal view. B. Habitus, dorsal view. C. ♂, habitus, lateral view. D. Habitus, lateral view. E. Juvenile, habitus, lateral view. F. Leg-pair 17, anterior view. G. Leg-pair 18, anterior view. H. Telopods, anterior view. I. Left telopod, posterior view. Scale bars: 1 mm (A–E) and 0.2 mm (F–I).
FIGURE 8. Trachysphaera schmidtii Heller, 1858. A, B, H. ♂ syntype, ex. Gervaisia costata var. acutula Latzel, 1884, syn. n., from the Eastern Alps or Carpathians (NHMW MY1822). C, D, E, I. ♂♂, ex. Gervaisia cultrifera Verhoeff, 1906, syn. n. (NHMW MY1838 and NHMW MY1841), F, G, J. ♂ syntype, ex. Gervaisia multiclavigera Verhoeff, 1898, syn. n., from Bosnia and Herzegovina (NHMW MY1865). A, B. Habitus, lateral and dorsal views, respectively. C, D. ♂ syntype from Sabine Mountains (Sabinergebirge), Italy, habitus, lateral and dorsal views, respectively (NHMW MY1838). E. ♂ from Camerata Vec- chia, Italy, habitus, lateral view (NHMW MY1841). F, G. Habitus, lateral and dorsal views, respectively. H. Gervaisia costata var. acutula Latzel, 1884, syn. n., telopods, anterior view (NHMW MY1822). I. Gervaisia cultrifera Verhoeff, 1906, syn. n., telopods, anterior view (NHMW MY1841). J. Gervaisia multiclavigera Verhoeff, 1898, syn. n., telopods, posterior view (NHMW MY1865). Scale bars: 1 mm (A–G) and 0.2 mm (H–J).
FIGURE 9. Trachysphaera schmidtii Heller, 1858, telopods, anterior views (IZB). A. ♂ from Trebljevine, Zasavica, Serbia. B. ♂ from Topčider, Belgrade, Serbia. C. ♂ from Petnička Cave, Serbia. D. ♂ from Buronov Ponor Cave, Serbia. E. ♂ from Džud Pit, Črnomelj, Slovenia. F. ♂ from Jama pri Metliki Pit, Metlika, Slovenia. Scale bars: 0.2 mm.
FIGURE 10. Trachysphaera schmidtii Heller, 1858, juveniles from Topčider, Belgrade, Serbia (IZB). A. II stadium. B. III stadium. C. IV stadium. D. V stadium. Scale bars: 0.5 mm.
R |
Departamento de Geologia, Universidad de Chile |
T |
Tavera, Department of Geology and Geophysics |
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Trachysphaera schmidtii Heller, 1858
Antić, Dragan, Šević, Mirko, Macek, Oliver & Akkari, Nesrine 2021 |
Trachysphaera noduligera
Tabacaru, I. 1988: 55 |
Trachysphaera noduligera hungarica
Tabacaru, I. 1988: 55 |
Trachysphaera illyrica
Tabacaru, I. 1988: 55 |
Trachysphaera schmidtii
Tabacaru, I. 1990: 26 |
Tabacaru, I. 1988: 53 |
Sillaber, H. 1987: 179 |
Trachysphaera cultrifera
Kime, R. D. & Enghoff, H. 2011: 36 |
Tabacaru, I. 1988: 54 |
Strasser, K. & Minelli, A. 1984: 198 |
Trachysphaera schmidti
Kime, R. D. & Enghoff, H. 2011: 38 |
Thaler, K. & Neuherz, H. 1978: 34 |
Trachysphaera noduligera
Thaler, K. & Neuherz, H. 1978: 33 |
Trachysphaera noduligera noduligera
Strasser, K. 1971: 13 |
Trachysphaera noduligera styrica
Strasser, K. 1971: 13 |
Trachysphaera noduligera illyrica
Strasser, K. 1971: 13 |
Trachysphaera multiclavigera
Kime, R. D. & Enghoff, H. 2011: 37 |
Tabacaru, I. 1988: 54 |
Strasser, K. 1971: 13 |
Gervaisia noduligera var. illyrica
Strasser, K. 1959: 6 |
Gervaisia noduligera noduligera
Jermy, T. 1942: 59 |
Gervaisia noduligera hungarica
Jermy, T. 1942: 60 |
Gervaisia illyrica
Attems, C. 1943: 79 |
Jermy, T. 1942: 64 |
Strasser, K. 1938: 402 |
Attems, C. 1929: 313 |
Gervaisia noduligera
Attems, C. 1943: 79 |
Schubart, O. 1934: 54 |
Attems, C. 1929: 313 |
Gervaisia multiclavigera declivis
Attems, C. 1929: 313 |
Gervaisia noduligera var. styrica
Attems, C. 1929: 313 |
Verhoeff, K. W. 1908: 532 |
Gervaisia illyrica
Foddai, D. & Minelli, A. & Scheller, U. & Zapparoli, M. 1995: 12 |
Strasser, K. & Minelli, A. 1984: 198 |
Verhoeff, K. W. 1908: 531 |
Gervaisia cultrifera
Attems, C. 1943: 79 |
Verhoeff, K. W. 1910: 162 |
Verhoeff, K. W. 1908: 530 |
Gervaisia noduligera
Sillaber, H. 1987: 181 |
Verhoeff, K. W. 1906: 812 |
Gervaisia noduligera styrica
Strasser, K. 1971: 13 |
Verhoeff, K. W. 1906: 812 |
Gervaisia multiclavigera
Attems, C. 1943: 79 |
Attems, C. 1929: 313 |
Verhoeff, K. W. 1906: 810 |
Gervaisia multiclavigera declivis
Strasser, K. 1971: 13 |
Verhoeff, K. W. 1906: 811 |
Gervaisia cultrifera Verhoeff, 1906: 810
Verhoeff, K. W. 1906: 810 |
Gervaisia costata multiclavigera
Verhoeff, K. W. 1899: 185 |
Gervaisia multiclavigera
Verhoeff, K. W. 1898: 165 |
Gervaisia costata var. acutula
Latzel, R. 1884: 89 |
Trachysphaera Schmidtii
Heller, C. 1858: 317 |