Prismognathus sukkitorum Nagai, 2005

Huang, Hao & Chen, Chang-Chin, 2012, A review of the genera Prismognathus Motschulsky and Cladophyllus Houlbert (Coleoptera: Scarabaeoidea: Lucanidae) from China, with the description of two new species, Zootaxa 3255, pp. 1-36 : 15

Prismognathus sukkitorum Nagai, 2005

( Figs. 67 , 97 , 129 , 155 , 191 , 215 , 235 )

Prismognathus sukkitorum Nagai, 2005: 37 [type locality: Chudo Razi, northeastern Kachin, northern Myanmar], Figs. 21–23 , males and female; Fujita 2010: 168, plate 101, partim [Figs. 505 (1–5), males from northern Myanmar].

Material examined. CHINA: Yunnan: 1 3 ( IEAS), Baoshan City, Tengchong County, Gaoligongshan Nature Reserve, Da-hao-ping station, on the old road from Tengchong to Baoshan, 17.IX.1991, Z.-Y. Liu, T.-Q. Wang & H.-S. Yin leg.; 1 Ƥ ( IEAS), same data; 1 3 ( CLYB), Nujiang Autonomous Prefecture, Gongshan County, VIII.2011, Y.- B. L i leg..

Identification. This species was originally described from northeastern Myanmar and compared with P. kanghianus . It differs from P. kanghianus in males “by having the body black, almost without luster, and the angulated frontal corners of pronotum” ( Nagai 2005). Fujita (2010) illustrated some reddish brown specimens alongside the black specimens from the same area; therefore the black color of the body is not a constant character. The male specimen from western Yunnan examined by us matches the original description of P. s u kk i to r um except for the redder color of the body. On the underside of the male specimen, each of the abdominal ventrites bears a transversal band of yellow setae near the posterior margin; this character is not found in all other species. The female was illustrated in the original description and very briefly described. The following peculiar characters have been noticed by us: 1) anterior margin of the pronotum markedly wider than head, with anterolateral corners more or less produced; 2) lateral angels of the pronotum a little protruding. The female from western Yunnan examined by us matches the above-mentioned characters. However, the female illustrated by Fujita (2010: plate 101, Fig. 505-6) does not match these characters and probably does not belong to P. sukkitorum .

Taxonomic notes. P. sukkitorum is sympatric with P. kanghianus kittii in northeastern Myanmar.

Distribution. Western Yunnan (Tengchong); northeastern Myanmar.

Figures

FIGURE 235. Distribution of the other Chinese Prismognathus.

FIGURES 57 – 72. Head, pronotum and proleg of male in dorsal view. 57 – 60 — P. alessandrae; 57 — Weixi, Yunnan; 58 — Muli, Sichuan; 59 — Zhongdian, Yunnan, 60 — Pantiange, Weixi, Yunnan; 61 – 64 — P. shani; 61, 63 — paratypes, Siping, Yingjing, Sichuan; 62 — paratype, Erlangshan, Sichuan; 64 — holotype, Erlangshan; 65 — P. arcuatus, Siping, Sichuan; 66 — P. haojiani, holotype, Fanjingshan, Guizhou; 67 — P. sukkitorum, Tengchong, Yunnan; 68 — P. prossi, Siping, Sichuan; 69 — P. kanghianus kanghianus, Hekou, Yunnan; 70 – 71 — P. triapicalis ssp.; 70 — Dayaoshan, Guangxi; 71 — Fengyangshan, Zhejiang; 72 — P. prossi, Maoershan, Guangxi. Scale bar 10 mm.

FIGURES 95 – 110. Head, pronotum and proleg of female in dorsal view. 95 — P. arcuatus, Siping, Sichuan; 96 — P. haojiani, paratype, Fanjingshan, Guizhou; 97 — P. sukkitorum, Tengchong, Yunnan; 98 — P. dauricus, Liaoning; 99 – 100 — P. p ro s s i; 99 — Fanjingshan, Guizhou; 100 — Maoershan, Guangxi; 101 — P. kanghianus kanghianus, Hekou, Yunnan; 102 — P. formosanus, Taiwan; 103 – 105 — P. triapicalis subspecies; 103 – 104 — Dabashan, Chongqing; 105 — Fengyangshan, Zhejiang; 106 — P. triapicalis triapicalis, Erlangshan, Sichuan; 107 — P. s i n e n s i s, holotype, Kuatun; 108 – 110 — P. davidis davidis; 108 — Dabashan, Chongqing; 109 – 110 — Siping, Sichuan. Scale bar 10 mm.

FIGURES 111 – 131. Female canthus (not under the same scale). 111 — P. s h a n i, paratype, Labahe, Sichuan; 112 — P. nigricolor, paratype; 113 — P. arcuatus, Siping, Sichuan; 114 – 116 — P. p ro s s i; 114 — Siping, Sichuan; 115 — Fanjingshan, Guizhou; 116 — Maoershan, Guangxi; 117 – 119 — P. davidis davidis; 117 — Siping, Sichuan; 118 — Tianmushan, Zhejiang; 119 — Dabashan, Chongqing; 120 – 123 — P. triapicalis; 120 — Sichuan; 121 — Dabashan, Chongqing; 122 — Fengyangshan, Zhejiang; 123 — Dabashan, Chongqing; 124 — P. haojiani, paratype, Fanjingshan; 125 — P. dauricus, Liaoning; 126 — P. kanghianus kanghianus, Hekou, Yunnan; 127 — P. alessandrae, Zhongdian, Yunnan; 128 — P. sinen sis, holotype, Kuatun, Fujian; 129 — P. sukk ito rum, Tengchong, Yunnan; 130 — P. formosanus, Taiwan; 131 — P. siniaevi, Gongshan, Yunnan.

FIGURES 150 – 167. Aedeagus in ventral view. 150 – 151 — Prismognathus arcuatus, Sichuan; 152 – 153 — P. haojiani; 152 — holotype, Fanjingshan; 153 — paratype, Leigongshan, Guizhou; 154 — P. kanghianus kanghianus, Hekou, Yunnan; 155 — P. sukkitorum, Tengchong, Yunnan; 156 – 157 — P. triapicalis triapicalis, Sichuan; 158 – 161 — P. triapicalis subspecies; 158 – 159 — Dabashan, Chongqing; 160 — Fengyangshan, Zhejiang; 161 — Dayaoshan, Guangxi; 162 – 163 — P. prossi; 162 — Siping, Sichuan; 163 — Fanjingshan, Guizhou; 164 – 165 — P. siniaevi; 164 — Jinping, Yunnan; 165 — Gongshan, Yunnan; 166 — Tetrarthrius sukkiti, Gongshan, Yunnan; 167 — Cyclorasis fukinukii, Motuo, Tibet. Scale bars 1 mm for all.

FIGURES 168 – 199. Flagellum of the median lobe. 168 – 171 — P. davidis davidis; 168 — Qinling, Shaanxi; 169 — Erlangshan, Sichuan; 170 — Tianmushan, Zhejiang; 171 — Dabieshan, Anhui; 172 — P. davidis cheni, Taiwan; 173 — P. nigricolor, Gongshan, Yunnan, same specimen as in Fig. 226; 174 – 178 — P. klapperichi; 174 — Simianshan, Chongqing; 175 — Wuyishan, Fujian; 176 — Maoershan, Guangxi; 177 — Nanling, Guangdong; 178 — Baishanzu, Zhejiang; 179 — P. dauricus, Liaoning; 180 — P. s h a n i, paratype, Siping; 181 — P. alessandrae, Zhongdian, Yunnan; 182 – 184 — P. p ro s s i; 182 — Siping, Sichuan; 183 — Fanjingshan, Guizhou; 184 — Maoershan, Guangxi; 185 — P. siniaevi, Gongshan; 186 – 187 — P. arcuatus, Sichuan; 188 – 189 — P. haojiani; 188 — holotype, Fanjingshan; 189 — paratype, Leigongshan, Guizhou; 190 — P. kanghianus kanghianus, Hekou, Yunnan; 191 — P. sukkitorum, Tengchong, Yunnan; 192 – 194 — P. triapicalis triapicalis, Sichuan; 195 – 196 — P. triapicalis subspecies; 195 — Dabashan, Chongqing; 196 — Fengyangshan, Zhejiang; 197 — P. formosanus, Taiwan; 198 — Tetrarthrius sukkiti, Gongshan, Yunnan; 199 — Cyclorasis fukinukii, Motuo, Tibet. Figs. 168 – 198 under the same scale. Scale bars 1 mm for all.

FIGURES 212 – 223. Female genitalia and the central protrusion of the 9 th tergites (at left bottom or right bottom). 212 — Prismognathus arcuatus, Sichuan; 213 — P. haojiani, paratype, Fanjingshan; 214 — P. kanghianus kanghianus, Hekou, Yunnan; 215 — P. sukkitorum, Tengchong, Yunnan; 216 – 217 — P. triapicalis triapicalis, Sichuan; 218 – 219 — P. triapicalis subspecies; 218 — Dabashan, Chongqing; 219 — Fengyangshan, Zhejiang; 220 — P. davidis davidis, Siping, Sichuan; 221 — P. siniaevi, Gongshan; 222 — P. prossi, Maoershan, Guangxi; 223 — P. formosanus, Taiwan. Scale bars 1 mm for all.

FIGURES 21 – 34. Male mandible. 21 – 30 — dorsal view; 21 — Prismognathus dauricus (Liaoning); 22 — P. davidis davidis (Beijing); 23 — P. davidis davidis (Zhejiang); 24 — P. klapperichi (Fujian); 25 — P. s i n i a e v i (Gongshan, Yunnan); 26 — P. siniaevi (Jinping, Yunnan); 27 — P. prossi (Sichuan); 28 — P. nigricolor (holotype); 29 — P. s h a n i (paratype, Sichuan); 30 — P. nigricolor (Gongshan); 31 – 34 — ventral view; 31 — P. davidis davidis (Zhejiang); 32 — P. klapperichi (Chongqing); 33 — P. klapperichi (Fujian); 34 — P. siniaevi (Gongshan, Yunnan). Arrow mark directed to dorsal subbasal tooth or gap on ventral teeth of the mandible.