Amplirhagada buffonensis, Köhler, 2010

Köhler, Frank, 2010, Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae), Records of the Australian Museum 62 (2), pp. 217-284 : 259-261

publication ID	https://doi.org/ 10.3853/j.0067-1975.62.2010.1554
publication LSID	lsid:zoobank.org:pub:749A271A-6942-4D4C-B0DF-B968462BFF0C
persistent identifier	https://treatment.plazi.org/id/03F987C5-FFBB-F770-01CF-10DE4EA1CF30
treatment provided by	Felipe
scientific name	Amplirhagada buffonensis
status	sp. nov.

Treatment

Amplirhagada buffonensis View in CoL n.sp.

Type locality ( Fig. 1 View Figure 1 ). Western Australia, Kimberley, Bonaparte Archipelago, Buffon Island , 14°54'40"S 124°44'13"S; KC-086 (leg. V. Kessner & A. Longbottom, 23 July 1988) .

Type material. Holotype WAM S34620 View Materials (Pl. 2.6) . Paratypes AMS C463758 (2 preserved specimens, as holotype), FMNH 219337 About FMNH (8 preserved specimens, same as holotype) , WAM S41482 View Materials (5 preserved specimens, as holotype) , FMNH 219365 About FMNH (8 preserved specimens, unnamed islet W of Buffon Island , 14°54'29"S 124°43'28"E) GoogleMaps , AMS C463759 (2 preserved specimens, as FMNH 219365 About FMNH ), WAM S41483 View Materials (5 preserved specimens, same as FMNH 219365 About FMNH ) .

Additional, non-type material. FMNH 219374 About FMNH , WAM S41484 (preserved specimens, unnamed island S of Buffon Island , 14°57'07"S 124°44'50"E) GoogleMaps .

Etymology. In reference to Buffon Island, where this species occurs.

Shell ( Fig. 65A–C View Figure 65 , Pl. 2.6). Broadly conical to almost flat with low spire; thin (translucent). Periphery evenly rounded to slightly angulate; upper and basal sectors of whorls rounded. Umbilicus open, forming narrowly winding opening, 10–20 percent concealed by columellar reflection. Background and ventral colour greyish horn to slightly beige, whitish cracks and growth lines visible; sub-sutural band absent; mid-whorl band absent or diffuse, yellowish brown, thin, only visible on last whorl; outer and inner lip whitish. Protoconch c. 2.7 mm in diameter, comprising 1.7 whorls, smooth. Teleoconch with faint axial growth lines only. Angle of aperture 45°, outer lip rounded, sharp to moderately thick, expanded, slightly reflected, basal node of lip absent to weak, palatal node absent. Parietal wall of inner lip inconspicuous.

Pallial morphology. Pallial cavity moderately deep, extending 3 ⁄ 4 whorl; mantle pigmentation comprises densely packed black spots or patches. Kidney extending about half of pallial cavity.

Genital morphology ( Figs. 64 View Figure 64 , 66 View Figure 66 ). Penis straight to slightly curved, more or less of same length as anterior part of oviduct. Vas deferens forms simple loop or coils before entering penis. Penial retractor muscle shorter than penis. Penial verge very short (barely visible), slender to spatulate, with pointed to rounded tip. Penial wall pustules very small, densely arranged in rows on apical to median portion of penial chamber, becoming oblique towards base of penial chamber. Main stimulatory pilaster well-differentiated, large, cone-shaped, sculptured by horizontal ridges that support little hooks, comprising almost entire length of penial chamber. Vas deferens entering penial sheath in upper third of penis. Vagina moderately wide, posteriorly inflated; inner vaginal wall with smooth longitudinal pilasters. Spermatheca of long, clearly extending over base of spermoviduct. Spermathecal duct rather wide, internally with smooth longitudinal pilasters. Spermathecal head elongately inflated with extended tubular or pyriform tip, connected with oviduct by connective tissue, internally smooth, with delicate wall. Free oviduct comprising less than half of anterior part of oviduct, more or less straight. Spermoviduct longer than anterior part of oviduct. Talon embedded in albumen gland close to anterior end of albumen gland.

Radular morphology ( Fig. 65E–F View Figure 65 ). Rectangular. Tooth formula C+14–15+3–4+18–22. In average 153±17.5 rows of teeth, 27.4±0.1 rows per mm (n = 2). Central teeth with bluntly pointed, triangular mesocones, shorter than base of tooth; ectocones vestigial. Lateral teeth with sharply pointed, triangular to ovate mesocones, length equal to base of tooth; ectocones well developed, endocones vestigial. Marginal teeth with triangular mesocones; ectocones shorter and narrower than mesocones; endocones reduced in size.

Comparative remarks. It has a low spired shell with reduced banding pattern. Differs from its sister species in the phylogeny, A. yorkensis , by angulate periphery, slightly larger shell size ( Table 1), and smaller verge. Material of this species has been labelled as “ Amplirhagada sp. 67” by Solem.