Amplirhagada tricenaria, Köhler, 2010

Köhler, Frank, 2010, Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae), Records of the Australian Museum 62 (2), pp. 217-284 : 249-251

publication ID	https://doi.org/ 10.3853/j.0067-1975.62.2010.1554
publication LSID	lsid:zoobank.org:pub:749A271A-6942-4D4C-B0DF-B968462BFF0C
DOI	https://doi.org/10.5281/zenodo.5239039
persistent identifier	https://treatment.plazi.org/id/33C642CD-AF40-4B3C-B77F-58B905C2563D
taxon LSID	lsid:zoobank.org:act:33C642CD-AF40-4B3C-B77F-58B905C2563D
treatment provided by	Felipe
scientific name	Amplirhagada tricenaria
status	sp. nov.

Treatment

Amplirhagada tricenaria View in CoL n.sp.

Type locality ( Fig. 1 View Figure 1 ). Western Australia, Kimberley, Bonaparte Archipelago, Prince Frederick Harbour , north of Hunter River mouth, 15°01'54"S 125°23'13"E; KC-080 (leg. V. Kessner & A. Longbottom, 22 July 1988) GoogleMaps .

Type material. Holotype WAM S34614 View Materials (Pl. 1.20) . Paratypes AMS C463757 (2 preserved specimens), FMNH 219322 About FMNH (6 preserved specimens), WAM S41476 View Materials (4 preserved specimens) .

Etymology. From tricenaria (Latin = thirty), in reference to manuscript name assigned to this species by Solem (“ Amplirhagada n.sp. 30”).

Shell ( Fig. 49A–D View Figure 49 , Pl. 1.20). Semi-globose, with medium high spire; thin to solid (translucent). Periphery slightly angulate; upper and basal sectors of whorls rounded. Umbilicus forming a chink, 90–100 percent concealed by columellar reflection. Background colour dark horn to ochre; sub-sutural and mid-whorl bands diffuse to well defined, yellowish brown, thin to moderately broad, most conspicuous on last whorls; ventral colour horn; outer lip colour same as shell; inner lip yellowish white. Protoconch c. 2.6 mm in diameter, comprising 2 whorls, almost smooth. Teleoconch with pronounced axial growth lines, last whorl almost smooth. Angle of aperture 30°, outer lip rounded, sharp to moderately thick, slightly expanded, slightly reflected, basal node of lip absent or weak, palatal node absent. Parietal wall of inner lip inconspicuous.

Pallial morphology. Pallial cavity deep, extending one whorl. Pigmentation on mantle mottled, greyish or brownish (probably leached in actual specimens). Kidney extending about half of pallial cavity.

Genital morphology ( Figs. 48 View Figure 48 , 50 View Figure 50 ). Penis straight, more or less of same length as anterior part of oviduct. Vas deferens forms simple loop before entering penis. Penial retractor muscle shorter than penis. Penial verge very short (barely visible), slender to spatulate with pointed tip. Penial wall pustules small, elongated, arranged in rows on apical to median portion of penial chamber. Base of inner penial walls with many smooth longitudinal pilasters. Main stimulatory pilaster well-differentiated, elongate to cone-shaped, sculptured by smooth ridges, comprising apical to median portion of penial chamber. Vas deferens entering penial sheath in upper third.Vagina of medium length and thickness, tubular, posteriorly slightly inflated. Inner vaginal wall with smooth longitudinal pilasters. Spermathecal duct moderately thick, internally with smooth longitudinal pilasters. Spermathecal head globular to elongately inflated, connected with oviduct by connective tissue, internally smooth, with thin wall. Free oviduct comprising less than half of anterior part of oviduct, more or less straight. Spermoviduct longer than anterior part of oviduct. Talon embedded in albumen gland at junction with spermoviduct.

Radular morphology ( Fig. 49F–G View Figure 49 ). Rectangular. Tooth formula C+15–17+3+18–23. In average with 152±3.0 rows of teeth, 28.1±0.8 rows per mm (n = 2). Central teeth with sharply pointed, ovate mesocones, shorter than base of tooth; ectocones vestigial. Lateral teeth with bluntly pointed, ovate mesocones, length equal to base of tooth; ectocones small, endocones vestigial. Marginal teeth with triangular mesocones; ectocones shorter and narrower than mesocones; endocones reduced in size.

Comparative remarks. This species has a very distinctive colour, similar only to A. anderdonensis , which differs by smaller shell size, less globose shell shape, higher position of peripheral angulation. Rather flattened diameter of whorls and very broadly conical shape of shell are diagnostic. Material of the present species has been labelled as “ Amplirhagada sp. 30” by Solem.