Amplirhagada Iredale, 1933

Köhler, Frank, 2010, Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae), Records of the Australian Museum 62 (2), pp. 217-284 : 220

Amplirhagada Iredale, 1933

Amplirhagada Iredale, 1933: 52 ; Solem, 1981a: 147–320; Solem, 1988: 28–32; Solem, 1991: 187–202. Type species Helix (Hadra) sykesi Smith, 1894 by original designation.

Tenuigada Iredale, 1939: 68 . Type species Tenuigada percita Iredale, 1939 by original designation.

Diagnosis

Rock or free sealer with medium sized (15–30 mm in diameter), thin and translucent to moderately thick, broadly conical to dome-shaped shell with moderately to strongly elevated spire. Umbilicus concealed by columellar reflection or narrowly open. Protoconch with weak to strongly developed radially elongated pustulations; transition to teleoconch inconspicuous. Teleoconch with very weak axial growth lines that may become almost invisible on last whorl or sculptured by well-developed, regular axial lirae. Last whorl moderate to wide in cross-section; periphery well rounded to slightly angulate (often transitions are found within single populations). Shell colour usually variable even within populations, background often yellowish brown to horn, frequently with darker, brown bands located below suture and on periphery of whorl; some species are uniform in colour. Genitalia typically; development depends on seasonal activity and maturity. Penis with well-developed sheath, extending entire length, thin proximally, thick distally. No well-developed epiphallus present. Penial retractor muscle attached at apex of penial complex. Vas deferens entering sheath from halfway up to almost apically; entering penial chamber through verge. Inner penial wall supports characteristic pustulation and basal pilasters; a main stimulatory pilaster varies in development from undifferentiated to very large, cone-shaped. Spermatheca relatively simple, short reaching base of spermoviduct; duct and head usually well differentiated; inner spermathecal and vaginal wall with longitudinal pilasters that vary in development and finer structure. Albumen gland elongate. Hermaphroditic duct tightly undulating. Radula rectangular in shape, usually between 3.5 and 5.5 mm long with 120–170 rows of teeth. Tooth formula variable, C+12–20+3–4+15–22.