Amplirhagada kessneri, Köhler, 2010

Köhler, Frank, 2010, Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae), Records of the Australian Museum 62 (2), pp. 217-284 : 242-243

publication ID	https://doi.org/ 10.3853/j.0067-1975.62.2010.1554
publication LSID	lsid:zoobank.org:pub:749A271A-6942-4D4C-B0DF-B968462BFF0C
persistent identifier	https://treatment.plazi.org/id/03F987C5-FF88-F746-01D7-16024ED1C9AE
treatment provided by	Felipe
scientific name	Amplirhagada kessneri
status	sp. nov.

Treatment

Amplirhagada kessneri View in CoL n.sp.

Type locality ( Fig. 1 View Figure 1 ). Western Australia, northwestern Kimberley, Bonaparte Archipelago, Bigge Island , 4.3 km north of Savage Hill , 14°35'15"S 125°11'05"E; KIS 1-41. Vine thicket on mid slopes north of small stream, dolerite, on trees and bushes in drier sections (leg. V. Kessner, 17 February 2008) GoogleMaps .

Type material. Holotype WAM S34610 View Materials (Pl. 1.16) . Paratypes AMS C463701 (4 preserved specimens, as holotype), WAM S36875 View Materials (10 preserved specimens, as holotype) , WAM S36477 View Materials (5 shells, 14°35'16"S 125°11'02"E) GoogleMaps , AMS C463702 (9 preserved specimens, 14°35'16"S 125°11'02"E), AMS C C.463703 (2 shells, 14°35'16"S 125°11'02"E), WAM S36658 View Materials (10 preserved specimens, 14°35'16"S 125°11'02"E) GoogleMaps , WAM S36876 View Materials (16 preserved specimens, 14°35'18.5"S 125°10'54.7"E) GoogleMaps , WAM S36475 View Materials (3 shells, 14°35'16"S 125°11'06"E) GoogleMaps , WAM S36476 View Materials (6 shells, 14°35'14"S 125°11'08"E) GoogleMaps .

Etymology. Named in honour of Vince Kessner in recognition of his lasting achievements as one of Australia’s finest experts in the field.

Shell ( Fig. 36A–C View Figure 36 , Pl. 1.16). Semi-globose to conical with low spire; thin to solid (translucent). Periphery evenly rounded to slightly angulate; upper sector of whorls rounded, basal sector somewhat flattened. Umbilicus open, forming a chink or narrowly winding opening, 30–90 percent concealed by columellar reflection. Background colour horn to light brown, exceptionally dark brown; sub-sutural and mid-whorl bands absent or diffuse to well marked, brown, thin to moderately thick, most conspicuous on last whorls; ventral colour horn to light brown; outer lip colour lighter than shell, horn; inner lip translucent. Protoconch c. 2 mm in diameter, comprising 2 whorls, almost smooth. Teleoconch with regularly spaced axial lirae becoming somewhat oblique on last whorl.Angle of aperture 45°; outer lip rounded, sharp to moderately thick, slightly expanded, not or slightly reflected, basal node absent or very weak, palatal node absent. Parietal wall of inner lip absent.

Pallial morphology. Pallial cavity deep, extending one whorl. Mottled mantle pigmentation dark grey to black. Kidney extending about half of pallial cavity.

Genital morphology ( Figs. 37–38 View Figure 37 View Figure 38 ). Penis straight, more or less of same length as anterior part of oviduct. Vas deferens forms simple loop before entering penis. Penial retractor muscle longer than penis. Penial sheath evenly thin. Penial verge short (<1 ⁄ 8 penial chamber), slender to spatulate, with truncated tip. Penial wall pustules small, densely arranged in rows over entire length of inner penial wall. Main stimulatory pilaster well-differentiated, forming a large, elongated cone, sculptured by smooth horizontal ridges, comprising apical to median portion of penial chamber. Vas deferens entering penial sheath in upper third.Vagina rather long, tubular; inner vaginal wall densely covered by tiny pustules. Spermathecal duct moderately thick, internally with longitudinal rows of densely packed ciliae. Spermathecal head globular to elongately inflated, connected with oviduct by connective tissue, internally smooth, densely ciliated, with thin wall. Free oviduct comprising±half of anterior part of oviduct, straight. Spermoviduct longer to much longer than anterior part of oviduct. Talon embedded in albumen gland close to anterior end.

Radular morphology ( Fig. 36E–G View Figure 36 ). Rectangular. Tooth formula C+14–17+0–4+18–21. In average with 117±2.9 rows of teeth, 27.6±0.7 rows per mm (n = 3). Central teeth with sharply pointed, triangular mesocones, shorter than base of tooth; ectocones tiny. Lateral teeth with sharply pointed, triangular mesocones, length equal to base of tooth; ectocones tiny, endocones vestigial. Marginal teeth with broad, flattened mesocones; ectocones shorter and narrower than mesocones; endocones reduced in size.

Comparative remarks. Differs from A. decora , its congener from Bigge Island, by flatter, uniformly coloured shell, much smaller main pilaster and finer pustulation of inner penial wall.