Philanthinus, de Beaumont, 1949

Malash, Alyaa A., Edmardash, Yusuf A. & Gadallah, Neveen S., 2023, The genera Philanthus Fabricius, 1790 and Philanthinus de Beaumont, 1949 (Hymenoptera: Crabronidae: Philanthinae) in Egypt, with a new record and the description of a new species, Zootaxa 5249 (2), pp. 151-189 : 158-159

Key to species of Philanthinus and Philanthus recorded from Egypt.

1. Inner eye margins simple, slightly concave and greatly bowed away from each other ( Fig. 3C ); pronotal collar distinctly reduced, depressed and appressed to mesoscutum ( Fig. 3A ) (genus Philanthinus )....... Philanthinus integer de Beaumont

- Inner eye margins distinctly emarginate (as in Fig. 4B); pronotal collar well-developed (Fig. 4D) (genus Philanthus )...... 2

2. Dorsal surface of propodeum entirely, densely punctate ( Fig. 18D ); apical margin of clypeus with two median teeth in female, and three in male................................................. Philanthus triangulum abdelcader Lepeletier

- Dorsal surface of propodeum at least partly smooth and shiny; apical margin of clypeus without teeth.................. 3

3. Propodeal dorsum with broad median rugose sulcus, lateral side of sulcus shiny, with some points; head and thorax with long whitish pilosity masking some parts ( Fig. 14C ); mesopleuron less shiny ( Fig. 14C ); male clypeal moustache dense, nearly meeting medially ( Fig. 14B ); thorax entirely black ( Fig. 14C ).............................. Philanthus rutilus Spinola

- Propodeal dorsum smooth and shiny, with or without shagreened median sulcus; head and thorax without long pilosity; mesopleuron shiny; male clypeal moustache either meeting or not meeting medially; thorax with some yellow........... 4

4. Malar space conspicuously long ( Fig. 8D ), as long as 2/3 F 1 in male and half length of F 1 in female; body predominantly yellow ( Fig. 7A, B ); gaster with superficial, dense punctures ( Fig. 8E )........................ Philanthus genalis Kohl

- Malar space inconspicuous; gaster clearly punctate, punctures sometimes spaced; body colour variable................ 5

5. Upper mesopleuron densely punctate like lower one ( Fig. 19C ); male clypeal moustache not meeting medially ( Fig. 20C ).. 6

- Upper mesopleuron smooth and shiny, unlike punctate lower area ( Fig. 6B ); male clypeal moustache meeting medially ( Fig. 5D )................................................................................................ 7

6. Mesoscutum shiny ferruginous ( Fig. 6B ) (rest of thorax yellow except black median sulcus), nearly smooth (impunctate), densely punctate at junction with scutellum; gastral T 2 sparsely punctate ( Fig. 19E )......... Philanthus variegatus Spinola

- Mesoscutum shiny black, scutellum and postscutellum ferruginous, propodeum black with two lateral yellow patches ( Fig. 21B ); mesoscutum sparsely punctate, punctures dense posteriorly ( Fig. 21B ); gastral T 2 with more or less dense punctures ( Fig. 21C )............................................................... Philanthus variegatus ecoronatus Dufour

7. Gastral T 3 yellow or with a continuous or medially interrupted broad yellowish to whitish band like those of T 2 and T 4; T 1 or T 1-3 with ferruginous; T 3 finely punctate................................................................... 8

- Gastral T 3 black, or with very reduced yellow markings ( Fig. 5A, C ), unlike those of T 2 and T 4; T 1 sometimes with ferruginous; T 3 finely shagreened, with very fine widely spaced punctures.......................... Philanthus coarctatus Spinola

8. Pronotal collar distinctly emarginate antero-medially ( Fig. 9C ); mesopleuron and propodeum mostly black; propodeal dorsum with rugose middle sulcus extending to base as triangle ( Fig. 10A )............................ Philanthus minor Kohl

- Pronotal collar indistinctly emarginate antero-medially; yellow colour more developed; propodeal dorsum not as above... 9

9. Face between antennal bases and clypeus and along inner eye margin below emargination smooth and shiny, impunctate ( Fig. 22A ); gastral T 3 with broad whitish yellow band interrupted medially, densely coarsely punctate, each half of fascia containing round black spot ( Fig. 22D )...................................................... Philanthus pulawskii sp. nov.

- Lower face punctate, either sparsely or densely; T 3 differs in colour and sculpture................................. 10

10. Lower face densely punctate (Fig. 4B), interspaces equal to puncture diameter........ Philanthus ammochrysus W. Schulz

- Lower face sparsely punctate, interspaces distinctly larger than puncture diameter................................. 11

11. Body entirely yellow ( Fig. 11A,C ), with thin black streak along upper part of inner eye margin that does not reach emargination ( Fig. 11B, D, E ); hind femoral venter with short setae; propodeal enclosure smooth, with inconspicuously rugose median sulcus ( Fig. 11A, C )..................................................................... Philanthus pallidus Klug

- Black more extensive than above, especially on head and thorax ( Fig. 17A ); hind femoral venter with long, fine setae; propodeal enclosure with coarsely rugose broad median sulcus, that extends to propodeal foremargin as inverted triangle (similar to that of P. minor ) ( Fig. 16B ).......................................................... Philanthus schulthessi Maidl

Figures

FIGURE 3. Philanthinus integer (de Beaumont, 1949) (♀): A, dorsal habitus; B, lateral habitus; C, frontal view of head; D, dorsal view of thorax and gaster.

FIGURE 5. Philanthus coarctatus Spinola, 1839: A, dorsal habitus (♀); B, frontal view of head (female); C, dorsal habitus (male); D, frontal view of head (♁).

FIGURE 6. Philanthus coarctatus Spinola, 1838:A, dorsal view of head and thorax (♀); B, lateral view of head and thorax (♁).

FIGURE 7. Philanthus genalis Kohl, 1891: A, dorsal habitus (♀); B, dorsal habitus (♁).

FIGURE 8. Philanthus genalis Kohl, 1891: A, frontal view of head (♀); B, frontal view of head (♁); C, lateral view of head and thorax (♀); D, lateral view of head and thorax (♁); E, dorsal view of propodeum and gaster (♀).

FIGURE 9. Philanthus minor Kohl, 1891: A, dorsal habitus (♀); B, lateral habitus (♀); C, frontal view of head (♀); D, frontal view of head (♁).

FIGURE 10. Philanthus minor Kohl, 1891: A, dorsal view of head and thorax (♀); B, dorsal view of head and thorax (♁); C, D, lateral view of head and thorax (♁); E, dorsal view of propodeum and gaster (♁).

FIGURE 11. Philanthus pallidus Klug, 1845: A, dorsal habitus (♀); B, frontal view of head (♀); C, dorsal habitus (♁); D, E, frontal view of head (♁).

FIGURE 14. Philanthus rutilus Spinola, 1839:A, frontal view of head (♀); B, frontal view of head (♁); C, lateral view of head and thorax (♁).

FIGURE 16. Philanthus schulthessi Maidl, 1924 (♀): A, lateral view of thorax; B, dorsal view of thorax; C, dorsal view of gaster (OÖLM).

FIGURE 17. Philanthus schulthessi Maidl, 1924 (♀): A, dorsal habitus; B, frontal view of head; C, lateral view of head and thorax; D, dorsal view of thorax (part) and gaster.

FIGURE 18. Philanthus triangulum abdelcader Lepeletier, 1845: A, frontal view of head (♀); B, frontal view of head (♁); C, lateral view of head and thorax (♀); D, dorsal view of head and thorax (♀); E, dorsal view of gaster.

FIGURE 19. Philanthus variegatus Spinola, 1839: A, frontal view of head (♀); B, frontal view of head (♁); C, lateral view of head and thorax (♀); D, dorsal view of head and thorax (part); E, dorsal view of thorax (part) and gaster.

FIGURE 20. Philanthus variegatus ecoronatus Dufour, 1854: A, lateral habitus (♁); B, frontal view of head (♀); C, frontal view of head (♁).

FIGURE 21. Philanthus variegatus ecoronatus Dufour, 1854 (♀): A, frontolateral view of head and thorax; B, dorsal view of head and thorax; C, dorsal view of gaster.

FIGURE 22. Philanthus pulawskii Malash, Edmardash & Gadallah, sp. nov. (holotype, ♀): A, frontal view of head; B, dorsal view of thorax; C, lateral view of thorax; D, dorsal view of gaster.