Stalisyne veromanduiorum, Szwedo, Jacek, Bourgoin, Thierry & Lefebvre, Fabrice, 2006

Szwedo, Jacek, Bourgoin, Thierry & Lefebvre, Fabrice, 2006, New Mnemosynini taxa (Hemiptera, Fulgoromorpha: Cixiidae) from the Palaeogene of France with notes on their early association with host plants, Zootaxa 1122, pp. 25-45 : 31-34

Stalisyne veromanduiorum sp. nov.

( Figs. 13–18 )

MATERIAL: Holotype, male. Specimen MNHN PA 2770; collection “Langlois­Meurinne / De Ploëg”, mounted in Canada balsam, deposited in Muséum national d’Histoire naturelle, Paris. State of preservation: right part of body partly destroyed, fore, mid and hind femora partly, preserved fore, mid and hind tibiae and tarsi missing, anterior part of body, face and part of tegmina covered with milky veil.

TYPE LOCALITY: Farm Le Quesnoy, Chevrière, region of Creil, Oise department (Northern France).

STRATIGRAPHIC POSITION: Lowermost Eocene, circa 53 Ma, Sparnacian, level MP7, of the mammal fauna of Dormaal ( Feugueur 1963; De Ploëg et al. 1998; Nel et al., 1999, Nel et al. 2004; Cavagnetto 2000).

ETYMOLOGY: Named after Gallo­Celtic tribe Veromandui, inhabiting area of river Oise valley.

DIAGNOSIS: Bigger than Stalisyne lutetiorum ; Vertex in mid line longer than wide at base (about as long as wide in S. lutetiorum ); Postocular carinae close to the compound eyes; RA with two terminalia (three in S. lutetiorum ), RP with five terminalia (four in S. lutetiorum ) M with six terminalia (five terminalia in S. lutetiorum ).

DESCRIPTION: General coloration in amber brownish, head, pronotum and mesonotum slightly lighter. Tegmina brownish, with darker spot in apical, external portion. Tegmina, both cells and veins covered with short chaetae.

Total length (male) 13.2 mm. Head with compound eyes (2.4 mm) distinctly wider than long; Anterior margin of head rounded in dorsal view. Vertex with lateral margins distinctly elevate, posterior margin angulately incised; vertex at base (0.6 mm) narrower than long in mid line (0.76 mm), narrowing apicad. Face very long (4.8 mm), with median carina and median ocellus. Two longitudinal carinae at the margin of frons and vertex, two transverse carinae, the upper delimiting vertex, and the lower uniting longitudinal carinae with lateral carinae present. Frons 1.52 mm long in mid line, with disc slightly concave; Frontoclypeal suture arcuate; Postclypeus 1.92 mm long in mid line, convex, with distinct lateral carinae; Anteclypeus 1.44 mm long in mid line, with median carina.

Pronotum about 3.44 mm wide, wider than head with compound eyes, 0.56 mm long in mid line, longer in lateral portion, with median carina and postocular carinae distinct, postocular carinae shifted anteriad close to the compound eyes; posterior margin incised at wide angle; Tegula with median carina, wider than long.

Mesonotum longer (3.12 mm) than wide (about 2.72 mm), with disc concave and five longitudinal carinae, scutellum flat.

Tegmen 10.5 mm long, 3.5 mm wide, clavus long reaching 2/3 of tegmen length, stigma distinct, elongately triangular, costal margin mildly curved at base, with distinct, short basicostal field, almost straight in median portion, rounded in apical portion; basal cell about twice as long as wide, ScR forked near base, ScRA1 reaching the stigma, RA forked apicad, with two terminalia, RP forked basad of nodal line, then forked apicad, with five terminalia; M simple at base, forked slightly apicad of ScRA1 forking, with six terminalia; CuA with two terminalia, CuA1 curved; CuA terminalia in left tegmen fused (aberrancy); first row of transverse veinlets uniting RP and M and M and CuA1 basad of nodal line, transverse veinlets of apical line not very regular, not forming distinct line, additional transverse veinlet apicad of third M forking on right tegmen; claval veins Pcu and A1 united apicad of half of length of clavus.

Hind wing with m­cu veinlet very short, and M branching slightly apicad of ir veinlet.

Male abdomen about 3.2 mm long; Pygofer 0.8 mm long in mid line, with median process, posterior margin incised lateral of median process, lobed of pygofer in lateral view without projections; Anal tube long (1.5 mm), wide and flattened, tapering apicad, with lateral carination, apical portion with distinct, wide ventrad lobes; Anal style long (0.48 mm); Genital styles quite short, without distinct widenings at apex, apical margin covered with chaetae. Aedeagus with a single long lateral spine (of periandrium?) visible.

Figures

FIGURES 13 – 16. Stalisyne veromanduiorum gen. et sp. nov. Fig. 13. Anterior part of body in dorsal view; Fig. 14. Left tegmen; Fig. 15. Part of right tegmen; Fig. 16. Part of hind wing. Scale bar: 1 mm.

FIGURES 17 – 18. Stalisyne veromanduiorum gen. et sp. nov. Fig. 17. Male pygofer in ventral view; Fig. 18. Male genital block in lateral view. Scale bars: 0.5 mm for 17, 1 mm for 18.