Krateriske Huber

Huber, John T., 2015, World reclassification of the Gonatocerus group of genera (Hymenoptera: Mymaridae), Zootaxa 3967 (1), pp. 1-184 : 38-39

Krateriske Huber , gen. n.

( Figs 146–174 )

Type species. Krateriske guianensis Huber , by present designation.

Diagnosis. Within Gonatocerini, the species of Krateriske are distinguished by the following combination of features: face ( Fig. 163 ) with subantennal sulci meeting above mouth margin and continuing to mouth as short, median sulci; fore wing ( Figs 157–159 ) with 2 or 3 hypochaetae on parastigma midway between proximal and distal macrochaetae; propodeum ( Figs 159 , 169, 171, 172 ) with median oval, crater-like elevation.

Description. FEMALE. Large specimens, 2125–2500 Μm in length. Colour. Body generally light brown except scape and clava dark; wings mostly with distinct brown suffusion. Head. Head ( Figs 146 , 164, 165 ) thick, about 1.6–1.8× as wide as long and about 1.1–1.2× as wide as high; in lateral view with strongly and evenly rounded anterior surface. Face ( Figs 147, 148 , 163, 167 ) almost 0.7× as wide as high; subantennal sulci present, strongly converging ventrally to meet just above mouth margin and continuing as short median sulcus to mouth; preorbital sulcus appressed against eye almost to its ventral margin, then strongly curved inward to dorsolateral corner of mouth. Toruli separated by their own width from transverse trabecula. Eye in lateral view almost as long as high, extending to back of head ( Figs 164–167 ). Malar space about 0.2× eye height; malar sulcus shallow, extending ventroanteriorly from posterior angle of eye almost to mouth and posterodorsally along posterior eye margin to level of foramen ( Figs 165–168 ). Gena in lateral view absent dorsally, wide ventrally and separated from occiput by blunt angle. Vertex in lateral view oblique, merging smoothly into face (separated by transverse trabecula), posteriorly merging smoothly into occiput. Ocelli with LOL about 0.3–0.7× POL and OOL 0.5–0.6× POL, with two setae between lateral ocelli. Occiput entire. Labrum with about 8 setae. Mandible with 3 teeth ( Fig. 168 ). Antenna. Scape about 7× as long as wide, with radicle distinct, narrow, about 0.3× scape length; pedicel about 0.11× scape length, as short and narrow as fl1; funicle 8-segmented, fl1 without mps, fl2–fl8 each with only 1 mps; clava almost 0.8× funicle length, with numerous (about 30) short, oblique mps ( Figs 146 , 151, 152 ). Mesosoma. About 1.9× as long as wide, 1.8× as long as high, and 0.8–0.9× as wide as high. Pronotum ( Figs 153 , 169 ) in dorsal view short but visible, longitudinally divided, the lobes abutting medially, with dorsal surface merging smoothly into lateral surface and lateral surface concave in posterior 0.6 (between posterior half of propleura and spiracle). Pronotal spiracle larger than propodeal spiracle, but its opening small ( Fig. 170 ). Propleura normal. Prosternum rhomboidal, posteriorly divided by longitudinal suture extending at least half its length, but in one species apparently not divided at all. Mesoscutum ( Figs 153 , 169 ) in dorsal view with rather wide but shallow, barely diverging notauli. Transscutal suture straight. Scutellum slightly longer than wide. Axilla normal. Prepectus narrow and strap-like, slightly wider medially than apically. Mesopleuron spindle-shaped and truncate at both ends, with sulcus separating mesepimeron from mesepisternum ( Figs 146 , 170 ). Metanotum with rectangular dorsellum, its posterior margin distinctly convex ( Figs 153 , 169, 171, 172 ). Metapleuron triangular, separated from propodeum by curved carina becoming thicker anteriorly ( Fig. 170 ). Propodeum in lateral view ( Fig. 170 ) sloping fairly strongly, in different plane from dorsellum; in dorsal view ( Figs 160 , 169, 171, 172 ) with posteromedian crater-like oval area bounded by thick, flat carinae with several short submedian carinae and a long, median carina extending anteriorly from it. Propodeal spiracle small, separated by less than its diameter from metanotum. Wings. Fore wing at least 4.0× as long as wide, and densely and uniformly covered with microtrichia to base of parastigma ( Figs 157–159 ). Venation about 0.4× wing length. Submarginal vein with the usual two basal setae (1 macrochaeta and 1 hypochaeta), and a hypochaeta apically, next to proximal macrochaeta of parastigma. Remaining venation (parastigma + stigmal vein) slightly longer than submarginal vein, with 2 or 3 hypochaetae midway between proximal and distal macrochaetae, and more than 10 shorter setae between the macrochaetae. Stigmal vein with apex oblique. Hind wing slightly wider towards apex than at level of hamuli. Venation about 0.45× wing length. Metasoma. Petiole short, slightly longer than wide ( Figs 153, 154 , 171 ). Gaster about 1.5× as long as high. Terga narrow and widely separated by white membrane between them ( Figs 146 , 154 , 160, 162 ). Ovipositor sheath shorter than gaster, shorter than metatibia and not exserted beyond apex of gaster ( Figs 146 , 161, 162 ), with 1 subapical seta.

MALE. Body length 2200–2460. Colour. Body generally darker than in female. Antenna. Scape less than 2× as long as wide, with radicle short and fused with scape; pedicel narrower and much shorter than fl1; flagellomeres each with numerous (30+) mps in three irregular whorls ( Fig. 149 ). Metasoma. Genitalia with wide, flat scoop-like lateral projections (volsellae?) at base of curved aedeagal apodemes that do not extend posteriorly as far as apex of apodeme of genital sternite ( Figs 155 , 156 ).

Etymology. From Greek “krater” meaning bowl or basin, and “-iske,” a suffix meaning little. The name refers to the small, crater-like elevation on the propodeum. The gender is feminine.

Distribution. Krateriske species occur in northern South America.

Hosts and habitat. Hosts are unknown. Specimens have been collected in forests.

Included species:

Krateriske ecuadorensis Huber ; holotype ♀ in CNC. TL: Ecuador, Napo, Yasuni Biological Research Station. Krateriske guianensis Huber ; holotype ♀ in CNC. TL: French Guiana, Saül, Mt. Fumée. Krateriske peruensis Huber ; holotype ♀ in CNC. TL: Peru, Huanuco. Rio Llullapichis.

Figures

FIGURES 146 – 148. Krateriske spp.: 146, K. guianensis, paratype, habitus; French Guiana, 41.5 km S. Matoury, 4 ° 37.37 ' N 52 ° 22.59 ' W, 50 m, 29. v – 9. vi. 1997, J. Ashe, R. Brooks; 147, K. guianensis, head anterior; 148, K. peruensis, head anterior. Scale bars: 146 = 1000 Μm, 147 and 148 = 200 Μm.

FIGURES 149 – 152. Krateriske spp., antenna: 149, K. guianensis, male (+ enlargement, showing sensilla); 150, K. ecuadorensis, female (fl 6 – fl 11 missing); 151, K. guianensis, female; 152, K. peruensis, female. Scale bars = 200 Μm.

FIGURES 153 – 155. Krateriske spp., wings: 153, K. ecuadorensis; 154, K. guianensis; 155, K. peruensis. Scale bars = 500 Μm.

FIGURES 156 – 158. Krateriske guianensis, female: 156, propodeum + metasoma, dorsal; 157, genitalia, ventral; 158, metasoma, dorsolateral. Scale bars = 500 Μm.

FIGURES 159 – 162. Krateriske guianensis, male: 159, mesosoma + base of metasoma, dorsal; 160, metasoma, dorsal; 161, genitalia, dorsal; 162, genitalia, ventral. Scale bars: 159, 160 = 500 Μm; 161, 162 = 100 Μm.

FIGURES 163 – 168. Krateriske guianensis, head: 163, anterior; 164, dorsal; 165, lateral (+ propleura); 166, dorsoposterior; 167, ventral; 168, mouthparts. Scale bars = 100 Μm.

FIGURES 169 – 174. Krateriske guianensis: 169, mesosoma, dorsal; 170, mesosoma, lateral; 171, posterior of frenum – anterior of gt 1, posterodorsal; 172, propodeum, laterodorsal; 173, gaster, dorsal; 174, gaster, lateral. Scale bars = 200 Μm.