Macohasa Dubatolov, Volynkin & Kishida, 2018

Dubatolov, Vladimir V., Volynkin, Anton V. & Kishida, Yasunori, 2018, Review of the Prabhasa Moore, 1878 genus-group (Lepidoptera, Erebidae, Arctiinae), Zootaxa 4407 (3), pp. 383-400 : 385-388

Macohasa Dubatolov, Volynkin & Kishida , gen. nov.

Type species: Macohasa cernyi Dubatolov, Volynkin & Kishida , sp. nov. Macotasa: HolloWay 2001: 305 .

Prabhasa: Černý & Pinratana 2009: 139 .

Description. Forewing shape resembles that of Eilema , but dorsum noticeably conVeX in its basal half; its outer edge folded inwards, stronger in males. Costal Vein with androconial scales in basal half. Male genitalia: uncus long, narrow, with terminal denticle; scaphium heaVily sclerotized; apeX of ValVa broad, without distal process, costal margin straight; medial area of ValVe with a harpe-like fold; sacculus short, with bented apeX; aedeagus short, Vesica with seVeral diVerticula. Female genitalia: oVipositor short and broad; apophyses long and thin; ductus bursae, narrow, short, membranous, with a large, heaVily sclerotized plate Ventrally; corpus bursae large, membranous, with one signum; appendiX bursae short, bulbous, situated latero-posteriorly.

Remarks. 1. The new genus differs significantly either from species of Prabhasa , or Zadadra (type species Zadadra distorta ( Moore, 1872) ( Fig. 30 )); broad and weakly sclerotized apeX of cucullus is not characteristic for both genera (only “ Zadadra ” cucullata Joshi, Kirti & Singh, 2015 has broad distal part of ValVe, but it is heaVily sclerotized (see: Joshi et al. 2015: figs. 2–4), and the generic placement of this species will be clarified in the further reVision of the genus Zadadra ); harpe-like fold is also an autapomorphic character of the genus. 2. Macohasa orientalis ( Hampson, 1905) , comb. nov. , Macohasa tetraspila (Černý, 2009) , comb. nov. and Macohasa dimorpha ( Hampson, 1918) , comb. nov. were included formerly in Macotasa ( Holloway 1982; 2001; Fang 2000; Černý & Pinratana 2009; Bucsek 2012), but they are clearly not congeneric to the type species of Macotasa , M. biplagella (Butler, 1877) ( Figs. 31 , 41 ): in the male genitalia, M. biplagella has the bifid uncus, the furcated juXta, the large basal saccular process, and the small sclerotized harpe in the medial section of ValVe, whereas species of Macohasa haVe a simple uncus, a plate-like juXta, no basal saccular process, and no harpe (but a membranous fold in the medial section of ValVe is presented); in the female genitalia, M. biplagella has no sclerotized Ventral plate in ductus bursae which is characteristic for species of Macohasa . Another character of Macotasa , “a fold in the cell on male forewing ridges the dorsal surface, the basal part of this ridge coVered by an area of coarse scales to just beyond the ridge” ( Holloway 1982; 2001), is not autapomorphic, but common in Macohasa , Prabhasa, Chinasa , some Teulisna Walker, 1862 , genera with quite different male genitalia.

Etymology. The genus name is a combination of the generic names Macotasa and Prabhasa . Gender is feminine.

Figures

FIGURES 30–31. Lithosiina spp.: male genitalia (genital capsules in ventral vieW). 30, Zadadra distorta, Nepal, slide MWM 31500 Volynkin; 31, Macotasa biplagella, holotype, slide BMNH(E) Arct-2840 (©NHMUK).

FIGURES 36–41. Lithosiina spp.: female genitalia (ventral vieW). 36, Prabhasa venosa, [India, Meghalaya] Khasis, slide BMNH(E) Arct-4557f HolloWay (©NHMUK); 37, Macohasa cernyi, paratype, N Vietnam, slide MWM 31774 Volynkin; 38, Macohasa orientalis, Malaysia, slide MWM 31776 Volynkin; 39, Macohasa tetraspila, paratype, Thailand, Chantaburi prov., slide AV4186m Volynkin; 40, Macohasa dimorpha, Philippines, PalaWan, slide MWM 31778 Volynkin; 41, Macotasa biplagella, slide BMNH(E) Arct-2849 (©NHMUK).