Cothornobata

Li, Xuankun, Marshall, Stephen A. & Yang, Ding, 2015, A review of the Oriental species of Cothornobata Czerny (Diptera, Micropezidae, Eurybatinae), Zootaxa 4006 (2), pp. 201-246 : 205-207

publication ID	https://doi.org/ 10.11646/zootaxa.4006.2.1
publication LSID	lsid:zoobank.org:pub:B154E526-0C08-4A92-868E-A36E45F2F1A1
DOI	https://doi.org/10.5281/zenodo.6094482
persistent identifier	https://treatment.plazi.org/id/03ED4575-FFCB-FF98-759D-FA58DEE673AF
treatment provided by	Plazi
scientific name	Cothornobata
status

Treatment

Key to the Oriental species (males) of Cothornobata View in CoL

NOTE: Several species appear to be practically identical except for characters of the male terminalia. This key will therefore not work for females of those species.

Females remain unknown for 12 of the 19 Oriental species in the genus.

1 Body pale reddish-brown; mid and hind femora with yellow apex ( Fig. 24 View FIGURE 24 )........................ C. viriata Enderlein View in CoL

- Body mostly black or brown; mid and hind femora with brown or black apex ( Figs. 2 View FIGURE 2 A–23A, 25A–B)................. 2

2 Orbital seta absent (1 fronto-orbital seta) ( Fig. 14 View FIGURE 14 B). Genital fork with medial tubercle often large and strong, broadly convex ( Figs. 13 View FIGURE 13 D–16D, 18D, 19D, 21D). Distiphallus ususually distinctly longer than phallapodeme, apical filaments straight ( Figs. 13 View FIGURE 13 G–16G, 19G, 21G) (except in C. mentogensis View in CoL sp. nov. ( Figs. 18 View FIGURE 18 G)). Pregonite not lobed apically. Postgonite without apical seta ( Figs. 13 View FIGURE 13 G–16G, 18G, 19G, 21G)..................................................... C. uniseta View in CoL group …3

- Orbital seta usually present (thus 2 strong fronto-orbital setae) ( Fig. 4 View FIGURE 4 B) (except in C. paieroi View in CoL sp. nov. ( Fig. 9 View FIGURE 9 A)). Genital fork either with medial tubercle vestigial ( Figs. 2 View FIGURE 2 D–4D, 6D, 9B, 10D, 11D, 22D, 23D), or genital fork elongated, extending to sternite 7, and medial tubercle small and finger-like ( Figs. 7 View FIGURE 7 D–8D). Distiphallus relatively short, at most slightly longer than phallapodeme, apical filaments curved. Pregonite lobed apically. Postgonite with 1 apical seta ( Figs. 2 View FIGURE 2 G–4G, 6G–8G, 9B, 10G, 11G,

22G, 23G).......................................................................................... 9 3 Genital fork with basal tubercle ( Figs. 13 View FIGURE 13 D–15D, 18D, 21D). Distiphallus usually inflated apically, glans present ( Figs. 13 View FIGURE 13 GH– 15GH, 21GH) (except in C. mentogensis View in CoL sp. nov. ( Figs. 18 View FIGURE 18 GH))............................................... 4

- Genital fork without basal tubercle ( Figs. 16 View FIGURE 16 D, 17D, 20C). Distiphallus inflated apically, glans absent ( Figs. 16 View FIGURE 16 GH, 19GH, 20D).............................................................................................. 8

4 Genital fork with basal tubercles low and indistinctly convex ( Fig. 14 View FIGURE 14 D). Distiphallus curved ventrally ( Fig. 14 View FIGURE 14 G)............................................................................................... C. curva View in CoL sp. nov.

- Genital fork with basal tubercles minute and narrow, finger-like ( Figs. 13 View FIGURE 13 D, 15D, 18D). Distiphallus straight ( Figs. 13 View FIGURE 13 G, 15G, 18G)............................................................................................... 5

5 Palpus yellow. Scutellum flat, directed posteriorly ( Fig. 18 View FIGURE 18 I). Genital fork with basal tubercles separate (not contiguous) ( Fig. 18 View FIGURE 18 D). Distiphallus inflated apically, glans absent, slightly longer than phallapodeme ( Fig. 18 View FIGURE 18 G).... C. mentogensis View in CoL sp. nov.

- Palpus dark brown. Scutellum slightly convex, angled dorsally ( Fig. 21 View FIGURE 21 I). Genital fork with basal tubercles contiguous ( Fig. 13 View FIGURE 13 D, 15D, 21D). Distiphallus inflated apically, glans present, distinctly longer than phallapodeme ( Fig. 13 View FIGURE 13 GH, 15GH, 21GH).................................................................................................... 6

6 Wing lightly infuscated ( Fig. 21 View FIGURE 21 C). Genital fork with arms strongly divergent basally (strongly convergent on distal 1/3); width at widest point almost twice as wide as base at midpoint ( Fig. 21 View FIGURE 21 D)............................... C. uniseta View in CoL sp. nov.

- Wing relatively darkly infuscated ( Figs. 13 View FIGURE 13 C). Genital fork with arms not strongly divergent; widest point across arms only slightly wider than 1.5X width of base at midpoint ( Figs. 13 View FIGURE 13 D, 15D)............................................ 7

7 Postocellar seta absent ( Fig. 13 View FIGURE 13 I). Bm-cu in line with CuA2 ( Fig. 13 View FIGURE 13 C). Postgonite (viewed ventrally) slightly angled laterally ( Fig. 13 View FIGURE 13 GH)............................................................................. C. atra View in CoL sp. nov.

- Postocellar seta present ( Fig. 15 View FIGURE 15 I). Bm-cu basal to CuA2 ( Fig. 15 View FIGURE 15 C). Postgonite (viewed ventrally) strongly strongly bent, creating a “V-shape” ( Fig. 15 View FIGURE 15 GH)......................................................... C. elegantula View in CoL sp. nov.

8 Wing discal band relatively large, anterior to R4+5 ( Fig. 16 View FIGURE 16 C). Pregonite slender, needle-like; postgonite cuspate ( Fig. 16 View FIGURE 16 GH).................................................................................. C. longigonitea View in CoL sp. nov.

- Wing discal band relatively small, posterior to R4+5 ( Fig. 19 View FIGURE 19 C). Pregonite broader, crescent-like; postgonite twisted, corkscrewlike ( Fig. 19 View FIGURE 19 GH)................................................................... C. nigrigenu Enderlein View in CoL

9 Fore tibia black with yellow apex ( Fig. 6 View FIGURE 6 A). Genital fork large, broadly arched, wider than abdomen ( Fig. 6 View FIGURE 6 D)............................................................... Cothornobata ingensfurca View in CoL group … C. ingensfurca View in CoL sp. nov.

- Fore tibia black, yellow, or yellow to brown with darker base and apex. Genital fork narrower than abdomen, not conspicuously enlarged ( Figs. 2 View FIGURE 2 D–4D, 7D, 8D, 9B, 10D, 11D, 22D, 23D).............................................. 10

10 Genital fork elongate, extending to sternite 7 ( Figs 4 View FIGURE 4 A, 7A); arm twice as long as base ( Figs. 2 View FIGURE 2 D–4D, 7D, 8D)......... 11

- Genital fork short, not exceeding sternite 6; arm less than 1.5X as long as base of genital fork ( Figs. 9 View FIGURE 9 B, 10D, 11D, 22D, 23D)................................................................................................... 15

11 Arm of genital fork subparallel or only slightly incurved on distal 1/3 with short setulae at apex; medial tubercle indistinctly convex or absent ( Figs. 2 View FIGURE 2 D–4D)..................................................... C. breviseta View in CoL group … 12

- Arm of genital fork strongly incurved on distal 1/3 with long setulae on incurved section; medial tubercle small but distinctive ( Figs. 7 View FIGURE 7 D, 8D)................................................................... C. longifurca View in CoL group … 14

12 Wing discal band as dark as distal band, distal band without pale strip between R2+3 and R4+5 ( Fig. 2 View FIGURE 2 C). Arm of genital fork incurved apically; basal tubercle papillary ( Fig. 2 View FIGURE 2 D).......................................... C. breviseta View in CoL sp. nov.

- Wing discal band much paler than distal band, distal band with minor pale strip between R2+3 and R4+5 ( Figs. 3 View FIGURE 3 C, 4C). Arm of genital fork bowed at base and subparallel on distal 2/3; basal tubercle broadly convex ( Figs. 3 View FIGURE 3 D, 4D)................ 13

13 Body brownish-yellow ( Fig. 3 View FIGURE 3 A). Wing discal band rather small, reaching middle of penultimate section of M1, basal to dm-cu; cell cup short, A1+CuA2 more than 3X as long as CuA2 ( Fig. 3 View FIGURE 3 C). Postgonite (viewed ventrally) strongly bent, creating a “Vshape” ( Fig. 3 View FIGURE 3 G)......................................................................... C. fusca View in CoL sp. nov.

- Body blackish-brown ( Fig. 4 View FIGURE 4 A). Wing discal band relatively large, basal to middle of penultimate section of M1, distal to dmcu, cell cup long, A1+CuA2 twice as long as CuA2 ( Fig. 4 View FIGURE 4 C). Postgonite (viewed ventrally) slightly angled laterally ( Fig. 4 View FIGURE 4 G)................................................................................. C. shuimanensis View in CoL sp. nov.

14 Fore tibia entirely black ( Fig. 7 View FIGURE 7 A). Bm-cu in line with CuA2 ( Fig. 7 View FIGURE 7 C). Genital fork with basal tubercle small and medial tubercle finger-like, medial tubercle at midpoint of arm; inner surface of arms convergent basally, apex slightly inflated ( Fig. 7 View FIGURE 7 D)..................................................................................... C. cyanea (Hendel)

- Fore tibia yellow, with brown rings at base and apex ( Fig. 8 View FIGURE 8 A). Bm-cu proximal to CuA2 ( Fig. 8 View FIGURE 8 C). Genital fork with basal and medial tubercles subquadrate, medial tubercle on basal 1/3 of arm; inner surface of arms parallel basally, apex not inflated ( Fig. 8 View FIGURE 8 D).............................................................................. C. longifurca View in CoL sp. nov.

15 Bm-cu proximal to CuA2 ( Figs. 10 View FIGURE 10 C, 11C). Apex of arm distinctly inflated, fist-like ( Figs. 9 View FIGURE 9 B, 10D, 11D). Distiphallus elongate, exceeding base of expanded apex of phallapodeme ( Figs. 9 View FIGURE 9 C, 10G, 11G).................. C. pugnoa View in CoL group … 16

- Bm-cu distal to CuA2 ( Figs. 22 View FIGURE 22 C, 23C). Apex of arm not inflated ( Figs. 22 View FIGURE 22 D, 23D). Distiphallus distinctly shorter than phallapodeme, not reaching the expanded apex of phallapodeme ( Figs. 22 View FIGURE 22 G, 23G).............. C. vietnamensis View in CoL group … 18

16 Arm of genital fork with basal tubercle broad and shallow ( Fig. 10 View FIGURE 10 D). Distiphallus shorter than phallapodeme ( Fig. 10 View FIGURE 10 G)......................................................................................... C. pugnoa View in CoL sp. nov.

- Arm of genital fork with basal tubercle narrow and pointed ( Fig. 9 View FIGURE 9 B, 11D). Distiphallus slightly longer than phallapodeme ( Fig. 9 View FIGURE 9 C, 11D)...................................................................................... 17

17 Fore tibia entirely black ( Fig. 9 View FIGURE 9 A). Median occipital sclerite uniformly dark reddish-brown. Arm 1.7X as long as base, slender basally ( Fig. 9 View FIGURE 9 B)....................................................................... C. paieroi View in CoL sp. nov.

- Fore tibia dark yellow to brown with darker base and apex ( Fig. 11 View FIGURE 11 A). Median occipital sclerite with a triangular dark yellow spot at middle ( Fig. 11 View FIGURE 11 B). Arm about 1.2X as long as base, thick basally and slender before apex ( Fig. 11 View FIGURE 11 D).................................................................................................. C. zhangae View in CoL sp. nov.

18 Fore tibia entirely black; tarsi blackish-brown, except mid basitarsus dark yellow with blackish-brown apex ( Fig. 22 View FIGURE 22 A). Wing discal band darker than distal band ( Fig. 22 View FIGURE 22 C). Arm of genital fork subparallel basally and incurved apically at right angle, medial tubercle indistinctly convex ( Fig. 22 View FIGURE 22 D)........................................... C. bubengensis View in CoL sp. nov.

- Fore tibia yellow with brown base; fore tarsus pale yellow, mid and hind tarsi dark yellow, except mid basitarsus pale yellow ( Fig. 23 View FIGURE 23 A). Wing discal band lighter than distal band ( Fig. 23 View FIGURE 23 C). Arm of genital fork arcuate, medial tubercle absent ( Fig. 23 View FIGURE 23 D)................................................................................. C. vietnamensis View in CoL sp. nov.