Theognete bothynometopon Anderson, 2010

Anderson, Robert S., 2010, A taxonomic monograph of the Middle American leaf-litter inhabiting weevil genus Theognete Champion (Coleoptera: Curculionidae; Molytinae; Lymantini) 2458, Zootaxa 2458 (1), pp. 1-127 : 32-33

21. Theognete bothynometopon Anderson , new species ( Figures 1 , 7 , 90 , 100 , 109 , 316–319 )

Diagnosis. Size. Length, male, 4.10–4.95mm; female, 3.93–5.23mm. Width, male, 2.42–2.75mm; female, 1.97–2.81mm. Head with deep excavation behind eyes extended onto vertex, excavation separated into two halves by median raised carina or ridge; head not noticeably constricted behind eyes in lateral view. Pronotum elongate, with raised, variously developed median carina extended throughout middle of length but not extended to apex or base, carina sometimes low and irregular or absent; with prominent anterodorsal tubercles, each tufted with recurved suberect to erect scales; surface with coarse deep punctures. Front coxae widely separated by about the width of a coxa. Elytra with numerous low tubercles, each tufted with long erect recurved scales. Aedeagus as in Fig. 100 .

Geographical distribution. México (Oaxaca).

Natural history. Collected from berlese extraction of oak forest and oak-pine forest leaf litter at elevations ranging from 2200–2925m.

Derivation of specific name. Noun in apposition; named from the Greek bothynos, meaning hole, and metopon, meaning brow or forehead, in reference to the large pits on the posterior part of the head.

Material examined. 32♂, 22♀♀ ( BMNH, CMNC, CWOB, ECOS, MZLU) . Holotype ♂ ( CMNC) : MÉXICO: Oaxaca. 89.5km S. W. Valle Nacional, KM 142.5, 2430m, 28.VII.1992 -040, R.S. Anderson, wet alder/oak/pine for. leaf litt. berlese / HOLOTYPE ♂, Theognete bothynometopon sp. nov., R.S. Anderson. Aedeagus extracted. Paratypes: MÉXICO. Oaxaca. Llano de las Flores , 2713m, 17.VII.1987, R. Anderson, wet oak pine forest, 87–25,29 (8♂, 4♀♀ BMNH, CMNC, ECOS) . Oaxaca (17km. N.), 2200m, 14.IX.1986, R. Baranowski, sifting litter mixed pine forest (1♂ MZLU) . Oaxaca (20 km N.) 9500', 7.VI.1971, S. Peck, oak litter, BER210 (3♀♀ CMNC) . Oaxaca (78 km. N.), 2800m, 21.XI.1989, R. Baranowski, sifting litter in wet pine-oak forest (1♂ MZLU) . Oaxaca (79 km. N., Hwy. 175), 2900m, 2.X.1990, R. Baranowski, sifting litter wet mixed pine forest (1♂, 1♀ MZLU) . Valle Nacional (37 mi. S.), 8500', 24.V.1971, S. Peck, oak litter, BER206 (1♀ CMNC) . Valle Nacional (37 mi. S. Hwy 175), 9600', 3.VI.1983, C. W. & L. O'Brien & G.B. Marshall, berlese sifted oak cloud forest litter (9♂, 4♀♀ CWOB) . Valle Nacional (64.5 km. S. W.) , 2600m, 28.VII.1992, R. Anderson, oak forest leaf litter berlese (2♂ CMNC) . Valle Nacional (89.5 km. S. W.) , 2430m, 28.VII.1992, R. Anderson, wet alder/oak/pine forest leaf litter berlese (3♂, 2♀♀ CMNC) . Valle Nacional (66 km. S.), 2800m, 2.X.1990, R. Baranowski, sifting litter in pine-oak forest (1♂, 4♀♀ MZLU) . Valle Nacional (68 km. S.), 2800m, 17.IX.1990, R. Baranowski, sifting litter in mixed pine forest (1♂ MZLU) . Valle Nacional (70 km. S.), 2600m, 3.X.1986, R. Baranowski, sifting litter pine-oak forest (1♂ MZLU) . Valle Nacional (70 km. S.), 2500m, 10.IX.1986, R. Baranowski, sifting litter pine-oak forest (1♀ MZLU) . Valle Nacional (70 km. S.), 2700m, 29.IX.1990, R. Baranowski, sifting litter in mixed pine forest (1♂, 2♀♀ MZLU) . Valle Nacional (74 km. S.), 2900m, 21.XI.1989, R. Baranowski, sifting litter in pine-oak forest (1♀ MZLU) . Valle Nacional (84 km. S.), 2850m, 4.IX.1990, R. Baranowski, sifting litter pine-oak boreal forest (1♂ MZLU) . Oaxaca (80 km. N), 2900m, 8.IX.1994, R. Baranowski, sifting litter mixed pine forest (1♂ CMNC) .

Chorological relationships. Sympatric with T. giselae , T. birdi , T. weiri , T. baranowskii , T. semiopaca and T. lalibertei .

Figures

FIGURE 1. Habitus, Theognete bothynometopon Anderson, new species (México, Oaxaca).

FIGURES 2–7. Thoracic sterna, ventral view, Theognete species. 2. T. gracilis. 3. T. striatifemur. 4. T. fulgida. 5. T. semistriata. 6. T. caviventris. 7. T. bothynometopon.

FIGURES 89–94. Habitus, dorsal view, Theognete bothynometopon species group. 89. T. jeanae. 90. T. bothynometopon. 91. T. pragudemi. 92. T. giselae. 93. T. ashei. 94. T. birdi.

FIGURES 99–101. Aedeagus, dorsal and lateral view, Theognete bothynometopon species group. 99. T. jeaneae. 100. T. bothynometopon. 101. T. pragudemi.

FIGURES 108–116. Abdominal ventrite 5, male, ventral view, Theognete bothynometopon species group. 108. T. jeaneae. 109. T. bothynometopon. 110. T. pragudemi. 111. T. giselae. 112. T. ashei. 113. T. birdi. 114. T. genieri. 115. T. carinata. 116. T. chiapaneca.

FIGURES 316–331. Female genitalia (top to bottom; spermatheca, tergite VIII, sternite VIII, tergite VII), Theognete species, various species groups, representatives. 316–319. T. bothynometopon. 320–323. T. tzotzil. 324–327. T. terrestris. 328–331. T. hesmos.