Campsicnemus

Evenhuis, Neal L., 2009, Review of Campsicnemus (Diptera: Dolichopodidae) of the Marquesas, French Polynesia, with description of four new species groups, Zootaxa 2004, pp. 25-48 : 45

Campsicnemus ee Evenhuis, new species

( Figs. 22 , 24 )

Diagnosis. Distinguished from other members of the aa group by the strong apical seat on the hook-like process of the fore tibia and the strong long black setae on the swollen medial area of the ventral surface of the fore tibia.

Description. Male. Body length: 3.2 mm. Wing length: 2.6 mm.

Head. Black, face dark brown to black with reddish highlights, dark brown near clypeus; oc and vt black, about one-third length of antennal arista; front, occiput, and vertex black with blue highlights; face constricted at middle, almost holoptic, eyes separated below antennae by width of 1–2 ommatidia; palp small, dark brown; proboscis brown, extending slightly below eye in lateral view; scape and pedicel dark brown; postpedicel brown, short, conical, length subequal to width; arista slightly longer than head height.

Thorax: Mesoscutum, scutellum, and pleura dark brown to brown throughout, with a few metallic bluish highlights dorsolaterally; thoracic setae black: 1 + 3 dc; 2 np; 2 ph; 1 pa; 1 + 1 sc; ac absent; halter stem and knob white.

Legs: CI yellowish, CII and CIII brown; FI and FII yellowish brown, FIII yellowish brown basally, becoming darker brown on apical one-fourth; remainder of legs yellowish brown; FII with 2 strong black setae, one medially on ventral surface, one on apical one-third of ventral surface, remainder of femora unmodified, without MSSC; TiI ( Fig. 22 ) slightly thicker just beyond middle on ventral surface, with large hook-like process apically, length of hook-like process subequal to width of tibia, swollen area with 4 long, strong, stiff setae, apical hook-like process ( Fig. 24 ) with dense tibial comb (pecten) and bearing strong stiff black seta at tip (MSSC), remainder of foreleg unmodified; TiII slightly bowed and swollen, with long seta medially on dorsal surface and one subapically (MSSC). IIt1 about one-half length of Iit2, with 2 long black curved setae laterally, their length subequal to IIt2, and with dense wavy setae on mesal surface; Iit2 with row of long wavy setae on mesal surface, length subequal to segment. Remainder of leg segments without MSSC.

Wing: pale smoky throughout; posterior crossvein perpendicular to vein M.

Abdomen. Brown, dark brown along posterior margin, with short black hairs dorsally on each tergite, a few longer hairs laterally; sternites yellowish brown. Hypopygium dark brown with brown cerci, not dissected.

Female. As in male except for lack of MSSC; postpedicel length about 1.5 x width; legs normal, without modifications.

Types. Holotype ɗ (BPBM Type 16,629) [BPBM1013140] and 2Ψ paratypes [BPBM1013141] from FRENCH POLYNESIA: Marquesas: UA POU: Anakooma River, 1300 ft [396 m], cascade, sweep wall, 19 Aug 2001, R.A. Englund. Other paratype: FRENCH POLYNESIA: Marquesas: U A P OU: 1ɗ [BPBM1013142], Paeoa River, 1150–1500 ft [350–457 m], 17 Aug 2001, pools/cascades, R.A. Englund. Holotype and paratypes in BPBM.

Etymology. The specific epithet derives from the Marquesan ‘e‘e = “disappeared”; referring to the relative cryptic nature of the species, only being found in samples after collection of a number of flies from the same locality. The name is treated as a noun in apposition.

Figures

FIGURE 21 – 22. Campsicnemus aa group fore tibiae, mesal view. 21. C. aa, n. sp. 22. C. ee, n. sp.

FIGURES 23 – 27. Campsicnemus apices of fore tibiae showing hook-like process and setal arrangement. 23. C. aa, n. sp. 24. C. ee, n. sp. 25. C. ii, n. sp. 26. C. oo, n. sp. 27. C. uu, n. sp.