Tafana riveti Simon, 1903

De Oliveira, Luiz Fernando M. & Brescovit, Antonio Domingos, 2021, Taxonomic revision and cladistic analysis of ghost spiders of the genus Tafana Simon, 1903 (Araneae: Dionycha, Anyphaenidae), with the descriptions of twelve new species, European Journal of Taxonomy 742, pp. 1-77 : 15-17

Tafana riveti Simon, 1903

Figs 6G , 9B, E–F , 10 , 32F , 34

Tafana riveti Simon, 1903: 124 .

Tafana riveti – Berland 1913: 105, figs 91–96 (descr. of male). — Roewer 1954: 546. — Bonnet 1959: 4232. — Brescovit 1997: 90, figs 225–238.

Diagnosis

Males of Tafana riveti resemble those of T. straminea , T. kunturmarqa sp. nov., T. humahuaca sp. nov. and T. nevada sp. nov., by the apophysis at the base of the embolus short, but differ by retrolateral tibial apophysis short and with large base in retrolateral view and visible subtegulum on palp when not expanded ( Fig. 10C–E ). Females differ from those of other species by the short atrium, triangular hood and lateral lobe curved in the anterior region and sclerotized ( Fig. 10F ).

Material examined

Holotype ECUADOR • ♀; Cerro Troya , 5 km S of Tulcan; [0.73333° N, 77.68333° W]; 1901; G. Riveti leg.; MNHN.

Other material

COLOMBIA – Valle del Cauca • 1 ♂, Lago Calima; [3°53′44″ N, 76°29′41″ W]; Jan. 1979; W. Eberhard leg.; MCZ . – Magdalena • 1 ♂; Sierra Nevada de Santa Marta ; [10°53′27.3″ N, 73°59′43.9″ W]; 28 Sep. 2017; L. Martínez leg.; IBSP 223307 .

ECUADOR – Quito • 1 ♀; Pichincha, Monte Pichincha ; [0°15′00″ S, 78°35′00″ W]; Oct.1944; G.W. Prescott leg.; MCZ • 1 ♂, 1 ♀; Pichincha; 15 km from Pifo; [0°15′00″ S, 78°35′00″ W]; 5 Oct. 1982; A. Roig leg.; MACN-Ar • 3 ♂♂, 4 ♀♀; same collection data as for preceding; MCN 24624 • 2 ♀♀; Pichincha, 10 km from Pifo; [0°15′00″ S, 78°35′00″ W]; 30 Sep. 1978; A. Roig leg.; MACN-Ar • 1 ♂; Pichincha, Eucalyptus forest; [0°15′00″ S, 78°35′00″ W]; 30 Sep. 1978; A. Roig leg.; MACN-Ar • 1 ♂; Cotopaxi ; [0°40′ S, 78°26′ W]; 18 Apr. 1982; A. Roig leg.; MACN-Ar • 1 ♀; Loja [3°59′ S, 79°12′ W]; 21–22 Aug. 1977; L.E. Peña leg.; AMNH • 1 ♀; Azuay; [2°53′57″ S, 79°00′55″ W]; 20 Aug. 1988; W. Maddison leg.; MCZ • 1 ♂; Chimborazo; [2°8′59″ S, 78°54′27″ W]; Danas; 1904; G. Riveti leg.; MNHN • 2 ♂♂; Casitagua; [3°20′00″ S, 78°29′00″ W]; 1903; G. Riveti leg.; MNHN .

PERU – Cuzco • 1 ♂; Machu Picchu ; [13°9′47″ S, 72°32′44″ W]; 19–23 Jun. 1964; B. Malkin leg.; AMNH .

Remark

The type locality of Cerro Troya, 5 km south of Tulcan was defined using a database of Wharton Lab, Department of Entomology, Texas A&M University from http://mx.speciesfile.org/projects/8/public/public_content/show/21273?content_template_id=88.

Description

Male and female described by Brescovit (1997: 90, figs 225–238).

Distribution

Colombia, Peru and Ecuador ( Fig. 34 ).

Figures

Fig. 6. A. Arachosia praesignis (Keyserling, 1891), ♂, cephalothorax, dorsal view (arrow shows row of the eyes) (IBSP 87975). B. Patrera fulvastra Simon, 1903, ♂, cephalothorax, dorsal view (MCN 24207). C. Arachosia praesignis, ♂, chelicera, ventral view (IBSP 87975). D. Aysha prospera Keyserling, 1891, ♀, palp femur, prolateral view (arrows show setal row) (IBSP 19780). E. Tafana silhavyi (Caporiacco, 1955), ♀, prolateral view (arrow indicating the absence of setae) (MNHN 11312). F. Arachosia praesignis, ♂, leg I, tibia, ventral view, scopulae (IBSP 87975). G. Tafana riveti Simon, 1903, ♂, leg I, tibia, ventral view, scopulae (MCZ). H. Arachosia praesignis, ♂, palp, retrolateral view (IBSP 87975). I. Aysha prospera, ♂, palp, retrolateral view (IBSP 38914). J. Arachosia praesignis, ♂, palp, ventral view (arrow shows the basal notch on the tegulum) (IBSP 87975). K. Xiruana gracilipes (Keyserling, 1891), ♂, palp, ventral view (IBSP 72660). L. Patrera fulvastra, ♂, palp, retrolateral view (MCN 24207). Abbreviations: AEP=apophysis of embolic process; C= cymbium; Co =conductor; Co2 =secondary conductor; E =embolus; MA =median apophysis; MTA=median tibial apophysis; PMA =paramedian apophysis; RTA= retrolateral tibial apophysis; SD =sperm duct; ST=subtegulum; T =tegulum; VTP=ventral tegular process. Scale bars: C=0.2 mm; D, K =0.4 mm; E= 0.3 mm; F, H–J =0.5 mm; G, L=1 mm.

Fig. 9. A, C–D. Tafana silhavyi (Caporiacco, 1955), ♂ (MNHN 11312). A. Left palp unexpanded, ventral view (arrow indicating character). C. Left palp expanded, prolateral view. D. Left palp expanded, retrolateral view (arrow indicating character). B. Tafana riveti (MCZ), ♂, left palp. E–F. Tafana riveti (MCZ), ♂. E. Left palp, retrolateral view. F. Left palp, prolateral view (arrow indicating character). Abbreviations: 1= primary loop of the sperm duct; 2= secondary loop of the sperm duct; ABE=apophysis of base of the embolus; BH =basal haematodocha; C=cymbium; Ch =character; E =embolus; EP=embolic process; F = fundus; MA=median apophysis; MH= median haematodocha; P=petiole; SD = sperm duct; ST =subtegulum; T=tegulum. Scale bars: A–B, E–F =0.25 mm; C–D=0.5 mm.

Fig. 10. Tafana riveti Simon, 1903 (A, C–E =♂, MCZ; B, F–G =♀, MCZ). A. Habitus, dorsal view. B. Habitus, dorsal view. C. Left palp, ventral view (arrow indicating character). D. Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations:A=atrium; Ch =character; CD=copulatory ducts; CO=copulatory opening; E= embolus; FD = fertilization ducts; H = hood; LL =lateral lobes; MA=median apophysis; PA= patellar apophysis; RP = retrolateral projection; RTA=retrolateral tibial apophysis; SD = sperm duct; ST= subtegulum; S1 =primary spermathecae; S2 =secondary spermathecae; T =tegulum. Scale bars: A–B =3.5 mm, C–E, G =0.25 mm, F =0.4 mm.

Fig. 32. Female genitalia, digested. A. Arachosia praesignis (Keyserling, 1891) (Amaurobioidinae) (IBSP 87975). B. Xiruana gracilipes (Keyserling, 1891) (Anyphaeninae) (IBSP 166120). C. Aysha prospera Keyserling, 1891 (IBSP 19783). D. Anyphaena accentuata (Walckenaer, 1802) (MCN). E. Patrera fulvastra Simon, 1903 (MCN 24208). F. Tafana riveti Simon, 1903 (MCZ). G. Tafana straminea (L. Koch, 1866) (AMNH). H. Tafana quelchi (Pocock, 1895) (IBSP 32131). I. Tafana kunturmarqa sp. nov. (ICN-Ar). J. Tafana humahuaca sp. nov. (MACN-Ar 41788). K. Tafana pastaza sp. nov. (AMNH). L. Tafana nevada sp. nov. (AMNH). Abbreviations: CD =copulatory ducts; CO= copulatory opening; FD = fertilization ducts; H =hood; S1= primary spermathecae; S2 =secondary spermathecae. Asterisks to globose membranous extension of proximal copulatory ducts. Scale bars: A–B, D–I, K–L=0.25 mm; C=0.3 mm; J = 0.5 mm.

Fig. 34. Distribution map of species of Tafana.