Eotrichocera (Archaeotrichocera) spatiosa, Liu, Luxi, Shih, Chungkun & Ren, Dong, 2012

Liu, Luxi, Shih, Chungkun & Ren, Dong, 2012, Two new species of Ptychopteridae and Trichoceridae from the Middle Jurassic of northeastern China (Insecta: Diptera: Nematocera), Zootaxa 3501, pp. 55-62 : 59

Eotrichocera (Archaeotrichocera) spatiosa sp. nov.

( Fig. 3 )

Diagnosis: Sc rather long, about 0.84 times of the wing length, and ending slightly beyond R2; crossvein sc-r at the two-thirds of Rs; r-m at about one-third of d cell; d cell small, 0.19 times length of wing; A2 short, about 0.21 times of the wing length.

Description: Body: A well-preserved female adult fossil. Head subcircular and narrower than thorax. The compound eyes with clear facets. One antenna with three segments visible. The thorax poorly preserved. Abdomen length 11 mm, width 1.5 mm, with 8 clear segments. Female with ovipositor preserved. The legs long and slender. Wing: Length 12 mm and width 4.5 mm, 2.7 times as long as wide. Sc rather long, about 0.84 times of the wing length and terminating slightly beyond R2; Crossvein sc-r at the two-thirds of Rs. Rs arising from less than basal one-third of length of wing, Rs at 0.63 times length of wing ramified into R2+3+4 and R5; R2+3+4 short, being 0.6 times as long as R2+3; R2 short, about one-seventh of the length of dR1, and nearly one-fourth length of R2+3; R3 about 1.3 times as long as R2+3; section bR5 rather short, 0.3 times as long as r-m. Stem vein M ramified nearly at the same level of sc-r; bM1+2 longer than r-m, bM1 +2 about three-fifths mM1+2, and bM1+2 and mM1+2 strongly bent at r-m, mM1+2 distinctly longer than dM1+2; cell m1 as long as d cell, about twice as long as dM1+2, M1 slightly longer than M2. Crossvein m-m shorter than bM3, M3 distinctly longer than M4, r-m at about one-third of d cell; d cell broader distally, small, 0.19 times length of wing, m-cu meeting M3+4 slightly before its fork. CuA long. A1 long and reaching wing margin. A2 short, about 0.21 times length of wing and curving toward to posterior margin.

Material: Holotype, female, CNU-DIP-D-NN2008130 p/c, a well-preserved body with wings.

Etymology: The species name refers to the large size of this specimen.

Remarks: The new species is similar to Eotrichocera (Archaeotrichocera) ephemera Zhang, 2006 , but the former can be easily distinguished by its larger size; Sc 0.84 times length of wing and terminating slightly beyond R2; cell m1 as long as d cell, twice as long as dM1+2; d cell 0.19 times length of wing; A2 short, about 0.21 times length of wing. The new species is similar to the Eotrichocera (Archaeotrichocera) conica Krzemińska, Krzemiński et Ren, 2009 , but can be easily separated from the latter by Sc longer; sc-r at two-thirds of Rs, Rs fork 0.63 times length of wing; r-m at about one-third of d cell; d cell slight smaller; A2 slight shorter. Compared with Eotrichocera (Archaeotrichocera) rara Krzemińska, Krzemiński et Ren, 2009 , the main differences are: Sc longer, sc-r at two-thirds of Rs; Rs fork in 0.63 times length of wing; R2+3 longer than R2+3+4; d cell smaller; dM1+2 longer (about half length of cell m1); m-cu meeting M3+4 slightly before its fork; A2 slightly longer.

Figures

FIGURE 3. Eotrichocera (Archaeotrichocera) spatiosa sp. nov., holotype, CNU-DIP-D-NN 2008130 p / c. A. Photograph of counterpart; B. Photograph of part; C. Line drawing of part. Scale bars = 2 mm.