Heteromeringia
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publication ID |
https://doi.org/ 10.3853/j.0067-1975.61.2009.1531 |
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persistent identifier |
https://treatment.plazi.org/id/03C887E9-2949-FFFE-1B12-FF44027EF9F6 |
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treatment provided by |
Felipe |
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scientific name |
Heteromeringia |
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Key to the Australian Heteromeringia View in CoL View at ENA
1 Thorax yellow with brown stripes. Wing lightly clouded ............................................................. 2
—— Thorax dark brown, but, if dark with lighter stripes ( H. australiae View in CoL ), wing banded ( Figs 7, 8 View Figures 1–8 ) ................................................................................................................. 5
2 Scutum with single central floating stripe. Fore tibia yellow. Frons evenly orange medially. Bristles black ................................................................ H. limacens n.sp.
—— Scutum with one pair of stripes that often join anteriorly and continue onto sides of scutellum. Fore tibia brown, at least laterally. Frons with black medial tint (darkest laterally). Bristles brown .................................................... 3
3 Fore tarsi brown with distal two or three segments white. Anterior and posterior margins of fore tibia yellow to light brown. Pleuron yellow with thin, oblique subnotal stripe or thick black stripe touching notal margin. Tergites 1 and 2 brown. Surstylus pointed ( Fig. 44 View Figures 38–46 ) ............................................................................... H. hypoleuca D.K. McAlpine View in CoL
—— Fore tarsi entirely brown. Fore tibia entirely dark brown. Pleuron with faint white or orange subnotal mottling. Tergites 1 and 2 yellow. Surstylus rounded to truncated ( Figs 38, 41 View Figures 38–46 ) .................................................................... 4
4 Notal stripes broad, connecting anteriorly and attaining anterior margin of scutum. First flagellomere with inner-basal spot. Stripes on back of head reaching foramen. Surstylus widest at base ( Fig. 38 View Figures 38–46 ) ......................................................................................................... H. hypobrunnea n.sp.
—— Notal stripes very thin, separate and not extending past transverse suture. First flagellomere entirely pale. Stripes on back of head restricted to outer margins. Surstylus widest at apex ( Fig. 41 View Figures 38–46 ) ................................................................................................................. H. digitula n.sp.
5 Hairs of arista much longer than width of central filament. Katepisternum yellow below anepisternal suture. Surstylus wrinkled ( Fig. 69 View Figures 69–72 ). Wing clear ............................................................................... H. ptenopa n.sp.
—— Hairs of arista approximately as long as width of central filament at base. Katepisternum entirely brown, sometimes excluding ventral margin ( H. asteia ). Surstylus smooth. Wing at least infuscated anterodistally ................................................................................................................ 6
6 Vibrissae short and straight—barely overlapping if folded towards each other. Length of gena no more than twice height ( Figs 3, 4 View Figures 1–8 ). First flagellomere lobate ventrally and recessed dorsally (sometimes indistinct), with arista relatively close to pedicel. Wing with anterodistal infuscation and sometimes with cloud around dm-cu, but never banded .................................................................................. 7
—— Vibrissae long and usually curved upwards—widely overlapping if folded inwards (relatively short in H. norrisi View in CoL ). Length of gena at least three to four times height ( Figs 1, 2 View Figures 1–8 ). First flagellomere bilaterally symmetrical dorsoventrally; arista not near pedicel. Wing variable, sometimes with pronounced spots or bands .................................................................... 9
7 Mid and hind femora yellow, sometimes with brownish spot on apex of hind femur, or pale brownish band medially on male mid femur. Male: frons yellowish around bases of reclinate orbital bristles; fore tibia brown; fore tarsi yellow with distal two tarsomeres brownish; epandrium not wider than pregenitalic segments and mid and hind femora yellow; sides of epandrium excavated to reveal prominent hypandrium+pregonite ( Fig. 32 View Figures 28–37 ); apex of phallus terminating in two long filaments on one side ( Fig. 34 View Figures 28–37 ). Female: fore tarsi relatively high and flat (height strongly tapering past second segment); fore femur yellow with dark inner-distal spot; tergite 8 not produced posterolaterally; sternite 8 only slightly tapering apically; spermathecae with minute medial papillae (Fig. 93) ............................................................. H. hardyi D.K. McAlpine View in CoL —— Hind and (usually) mid femora with wide dark brown medial bands. Male: frons dark brown around bases of reclinate orbital bristles; fore tibia partly brown; if tarsi yellow with distal segments somewhat darker, epandrium very large and bulbous; terminalia not as above. Female (unknown for H. magnicauda ): fore tarsi not modified as above; fore femur dark brown on distal ½–¾; tergite 8 wrapping around segment posterolaterally and sternite 8 strongly tapered apically; spermathecae smooth excluding shallow basal wrinkles (Fig. 99) ................................................................................................................. 8
8 Fore tarsi dark brown. Fore tibia yellow with inner-distal margin brownish. Epandrium much narrower than pregenitalic segments ( Fig. 47 View Figures 47–57 ). Surstylus small and rounded. Distiphallus with complex distal sclerite. Hypandrium+pregonite as in Fig. 49 View Figures 47–57 .......................... H. laticornis D.K. McAlpine View in CoL
—— Fore tarsi yellow with distal segments brownish or entirely brown. Fore tibia brown, sometimes with apex lighter. Epandrium very large and bulbous ( Fig. 54 View Figures 47–57 ). Surstylus longer than wide and triangular. Distiphallus bifid apically, without apical sclerites, and with medial “wings” ( Fig. 56 View Figures 47–57 ). Hypandrium+pregonite as in Fig. 57 View Figures 47–57 ..................................................................................................... H. magnicauda n.sp.
9 Two fronto-orbital bristles. Wing dark anterodistally, as well as around and between R 4+5 and M 1 to r-m; dusky around dm-cu. Antenna sometimes entirely dark brown ............................................................... H. bisetosa n.sp.
—— Three fronto-orbital bristles. Wing various, but not as above. Antenna always yellow, at least in part ............................................................................................ 10
10 Three dorsocentral bristles. Legs predominantly brown with tibiae paler (hind tibia sometimes with two dark bands) ................................................ H. trisetosa n.sp.
—— Two dorsocentral bristles, sometimes with one relatively well developed setula in front of anterior bristle. Mid and hind tibiae and femora usually entirely or predominantly yellow ( H. imitans View in CoL sometimes with dark legs) .................................................................................................................... 11
11 Wing with dark, distinct, anterodistal bar along distal half of costa connecting (or nearly connecting) distal and medial bands (Figs 7, 8); additional transverse band usually present at wing base ( Fig. 5 View Figures 1–8 ) ......................................................................................................................................... 12
—— Wing clear, dusky, or infuscated anterodistally; if cross veins surrounded by spot, then widely separated from distal spot ( Fig. 6 View Figures 1–8 ) ................................................ 13
12 Body length 4.1–4.5 mm. Wing with two bands (medial band sometimes appearing to fade into dusky basal region). Male sometimes with grey or beige colouration on scutum ( Fig. 22 View Figures 22–27 ). Palpus usually brown at base or apex. Male anepisternal disc at least seven times width of anepisternal bristle. Fore tibia entirely dark brown. Hind femur brown apically. Surstylus relatively broad— length not more than three times width at midpoint ( Fig. 24 View Figures 22–27 ). Phallus relatively straight (single lateral lobe present at midpoint), smooth, and with small, thin reticulated membrane at apex. Subterminal flagellum of ventral receptacle straight ......................................................... H. australiae Malloch View in CoL
—— Body length 2.8–3.5 mm. Wing with three distinct bands (fourth basal band near alar base sometimes indistinct). Male scutum entirely brown excluding variable yellow posterolateral tint. Palpus entirely yellow. Male anepisternal disc not more than four to five times width of anepisternal bristle. Fore tibia dark brown (some females) or pale with apex brown (all males and some females). Hind femur entirely yellow. Surstylus relatively thin—length at least four times width at midpoint ( Fig. 73 View Figures 73–77 , 78 View Figures 78–87 ). Phallus strikingly branched at midpoint, covered with minute spicules from midpoint to base and without thin reticulated membrane at apex ( Fig. 75 View Figures 73–77 ); sometimes with “phallic blade” ( Fig. 77 View Figures 73–77 ). Subterminal flagellum slightly to strongly coiled (Fig. 106) ................................................. H. spinulosa D.K. McAlpine View in CoL 13 Face, parafacial and gena entirely black. Wing dark with posterior margin clearer ................................................................................................... H. imitans Malloch View in CoL
—— Face, parafacial and gena yellow to orange (posterior half of gena sometimes dark brown). Wing dusky or with anterodistal infuscation; sometimes with dark spot around cross veins .......................................................... 14
14 Wing with dark distal band and medial cloud around cells dm and bm ( Fig. 17 View Figures 17–21 ). Notum dark brown with yellowish band extending from sides of scutellum to posterior margin of notum or anterior dorsocentral ............................................................................................................... H. asteia n.sp.
—— Wing dusky or with single anterodistal infuscation. Notum dark brown, sometimes with postpronotum and notopleuron slightly paler ............................................................................................................................................. 15
15 Tergite 1 yellow. Fore tibia yellow with brown distolateral mottling. First flagellomere brown ventrally (excluding inner base). Hypandrium+pregonite extending laterally as long, dark bars ( Fig. 67 View Figures 62–68 ) ................................................................................................................... H. patula n.sp.
—— Tergite 1 dark brown. Fore tibia entirely dark brown, or yellow with one pair of distal spots. First flagellomere entirely yellow or with pigment on dorsal or anterior half. Hypandrium+pregonite neither dark nor produced laterally. Males only past this point ................................................... 16
16 Fore femur dark brown with base and apex lighter. Fore coxa brown at base ......................................................................................... H. norrisi D.K. McAlpine View in CoL
—— Fore femur only brown apically, sometimes only on inner face. Fore coxa entirely light yellow .................................................................................................... 17
17 Fore tibia yellow with one pair of distal spots (sometimes united as a band). First flagellomere entirely light yellow in male. Hypandrium+pregonite unbroken and apex of phallus with coiled filament ( Fig. 37 View Figures 28–37 ) ...................................................................................................... H. helina n.sp.
—— Fore tibia brown. First flagellomere dark, at least along anterior and inner-distal margins. Hypandrium+pregonite (seen laterally) broken into two or three sclerites divided by thin suture(s) and phallus uncoiled ........................................................................................................................................ 18
18 Basal 2 ⁄ 3 of palpus brown. Frons sometimes entirely smooth and black. Epandrium nearly closed apically and surstyli almost vestigial ( Figs 81, 82 View Figures 78–87 ) .............................................................................................. H. stegna n.sp.
—— Palpus entirely black or light yellow. Frons tomentose on anterior half, and orange to yellow anteriorly and laterally. Sides of epandrium parallel apically and surstyli well developed .................................................................... 19
19 Palpus entirely black. Wing dusky (clearer to base). Sides of surstylus parallel on basal half ( Fig. 50 View Figures 47–57 ) ................................................................... H. macropa n.sp.
—— Palpus entirely light yellow. Wing with anterodistal infuscation. Surstylus gradually tapering to a point ( Fig. 58 View Figures 58–61 ) .................................................. H. montana n.sp.
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