Scarodytes ruffoi FRANCISCOLO 1961

Fery, H. & Šťastný, J., 2007, Notes on the Scarodytes savinensis-complex with the description of two new taxa (Coleoptera: Dytiscidae), Linzer biologische Beiträge 39 (2), pp. 877-899 : 883-884

publication ID	https://doi.org/ 10.5281/zenodo.5416778
persistent identifier	https://treatment.plazi.org/id/03C387AA-D31C-D90B-A8B5-25C2FDD7FEF7
treatment provided by	Valdenar
scientific name	Scarodytes ruffoi FRANCISCOLO 1961
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Treatment

Scarodytes ruffoi FRANCISCOLO 1961 View in CoL

Deronectes (Potamonectes) halensis (FABRICIUS) ; MÜLLER 1926: 292 (partim).

Scarodytes savinensis View in CoL ruffoi FRANCISCOLO 1961: 159 View in CoL . - PEDERZANI 1967: 153. - ANGELINI 1975: 60 (partim). - FRANCISCOLO 1978: 176 (partim), 1979: 419 (partim).

Scarodytes ruffoi FRANCISCOLO View in CoL ; WEWALKA 1977: 142 (new status). - ANGELINI 1978: 89, 1984: 74, 1993: 5. - NARDI 1998: 75 (partim). - ROCCHI 2000: 13 (partim). - ROCCHI & TERZANI 2004: 210. - NILSSON 2001: 184, 2003: 69.

T y p e l o c a l i t y: Italy, Campania,MontePicentini,Bagnoli Irpino, river Calore, ca. 40.83N

15.07E {7*}.

T y p e m a t e r i a l: Holotype and Paratypes: stored in the Museo Civico di Storia Naturale di

Verona, Italy; types not studied, however, material from the locus typicus (see below).

A d d i t i o n a l m a t e r i a l s t u d i e d: I t a l y: 3., " Lucania, Latronico ( PZ) [ca. 40.09N 16.02E], m 400, 14/7/[19]73, leg Angelini" {2*} ( MNB). 4 exs., idem, ( CFA). 2 exs., " Campania, monti Picentini, Bagnoli Irpino [ca. 40.83N 15.07E], m. 620 II-XII-[19]71, leg. Angelini F. " {7*} ( MRTO). 3 exs., "Monti Picentini, Bagnoli Irpino (AV), f. Calore m. 500, 6-8- [19]72 leg. Angelini F. " {7*} ( MRTO, CHF). 4 exs., " Lucania, Pescopagano ( PZ) [ca. 40.84N 15.40E], lago Saetta m. 1000, 5-8-[19]72 leg Angelini F" {6*} ( CFA). 3 exs., "Parco Naz. d'Abruzzo, Pescasseroli, Fonte Aquarita, 8.VIII.72, m 1250 leg. Angelini F. " {9*} ( MRTO, CHF). 3 exs., " Abruzzo, Parco nazionale, Pescasseroli, sorg. f. Sangro m. 1300, 8-8-[19]72 leg. Angelini F. " {9*} ( MRTO, CHF). 3 exs., " Abruzzo, Pescasseroli ( AQ) [ca. 41.81N 13.79E], [" F. Sangro " crossed out], 8.VIII.[19]72, m. 1300, l. Angelini F." {9*} ( CFA). 3, " Abruzzo, 1270 m, Pescocostanzo [ca. 41.89N 14.07E], 2.VII.[19]72, leg. Angelini " {9*} ( CHF). 2, 3, "24.4.[19]84 Italia, Abruzzen, Chieti, Manoppello [ca. 42.26N 14.06E], Bach, Fery leg." {11 GoogleMaps *}; found together with S. h. halensis (CHF) . 8, 1, "23.3.[19]85 Italia, Abruzzen, b. Chieti, Manoppello, Bach, Fery leg." ; found together with S. h. halensis (CHF) . 1, "5.5.[19]90 Italia, Abruzzen, Chieti, Manoppello, Bach , Fery leg." ( CHF) .

D e s c r i p t i v e n o t e s: We have not been able to find external characters which would easily separate S. ruffoi from S. antoni . Usually, the black spots on the pronotum are smaller ( Fig. 10 View Figs 9-14 ) and more variable in size and shape in S. ruffoi . The females we have studied have the abdomen not mainly black but more or less dark brown or even reddish brown on the entire surface, and the brownish areas on the metacoxae and metasternum are more extended. The sublateral impressions on the pronotum are weaker, and in several specimens the prosternal process shows a weak but recognisable transversal ridge between the anterior margins of the procoxae. However, at present it is not clear whether these characters hold for all populations of S. ruffoi . The protibia is similar to that of S. antoni ( Fig. 19 View Figs 17-22 ) as the outer margin is also curved outwards at the apex in both sexes; this feature, allowing an easy separation of both species from S. savinensis and S. halensis , is reported here for the first time. N o t e s: The broad lateral margin of a female pronotum is shown in Fig. 5 of PEDERZANI (1990).

: Median lobe of aedeagus in ventral view slightly narrowed far before apex and parallel in apical third, apex broadly rounded ( Fig. 2 View Figs 1-4 ); parameres as in Fig. 6. Protarsal claws slightly shorter than in S. antoni .

: Sexual dimorphism as in S. antoni . Gonocoxosternum and gonocoxa more or less like in S. antoni (compare Figs 15 and 16). N o t e s: Single females cannot be separated with certainty from S. antoni without considering the locality.

M e a s u r e m e n t s: Neither FRANCISCOLO (1961, 1979) nor other works give morphometric measurements. We provide the following data: TL: 4.2-4.7 mm, MW: 2.15-2.4 mm, TL/MW: 1.87-1.98.

D i s t r i b u t i o n: Italy; the distribution of S. ruffoi is restricted to the southern and central Apennines from the Potenza province in the south to the Macerata province in the north (Fig. 29).

N o t e s: In addition to the localities listed above, we also include the following literature data in Fig. 29 ( ANGELINI 1975: 60, 1978: 89, 1984: 74; PEDERZANI 1967: 153) (provinces underlined): Potenza: Terranova Pollino (39.98N 16.30E) {1*}, Cersosimo (40.05N 16.35E) {1*}, San Chirico (40.19N 16.08E) {2*}, Laurenzana (40.46N 16.00E) {4*}, Tramutola (40.32N 15.79E) {4*}, Brienza (40.49N 15.63E) {5}, Baragiano (40.68N 15.58E) {6*}, Sarnelli (40.78N 15.76E) {6*}, Muro Lucano (40.75N 15.48E) {6*}, Bella (40.75N 15.54E) {6*}, Possidente (40.74N 15.71E) {6*}; Matera: Caputo (40.32N 16.35E) {3}; Avellino: Ponteromito (40.90N 15.05E) {7*}; Caserta: Gallo (41.47N 14.23E) {8}; Isernia: Pizzone (41.67N 14.03E) {9*}; Aquila: Opi (41.78N 13.83E) {9*}, Villetta Barrea (41.77N 13.95E) {9*}; Chieti: Fara Filiorum Petri (40.25N 14.19E) {11*}; Pescara: Roccaramanico (42.11N 14.01E) {10}, Farindola (42.44N 13.81E) {12*}; Teramo: Isola del Gran Sasso (42.50N 13.66E) {12*}; Perugia: Sellano (42.89N 12.93E) {13*}; Macerata: Polverina (43.09N 13.12E) {13*}.

E c o l o g y: The specimens from Manoppello were collected in a slowly running brook of ca. 1.5 m width and not more than 50 cm depth. They were found together with Agabus biguttatus (OLIVIER 1795) , Agabus brunneus (FABRICIUS 1798) , Agabus didymus (OLIVIER 1795) , Agabus paludosus (FABRICIUS 1801) , D. moestus inconspectus, Hydroglyphus geminus (FABRICIUS 1792) , Hygrotus inaequalis (FABRICIUS 1777) , Laccophilus hyalinus (DE GEER 1774) , Meladema coriacea LAPORTE 1835 , Nebrioporus sansii (AUBÉ 1838) , and S. h. halensis. Surprisingly, females of the latter species had, in contrast to Central European populations, not a reddish but dark brown or black abdo- men. In addition, this record is in contrast to PEDERZANI (1967: 153) who reported never to have found S. h. halensis in the Abruzzi.

Kingdom	Animalia
Phylum	Arthropoda
Class	Insecta
Order	Coleoptera
Family	Dytiscidae
Genus	Scarodytes