Cryptocanthon

Cook, Joyce, 2002, A Revision Of The Neotropical Genus Cryptocanthon Balthasar (Coleoptera: Scarabaeidae: Scarabaeinae), The Coleopterists Bulletin (mo 1) 56, pp. 3-96 : 6-11

Key to Species of Cryptocanthon

1a. Pronotal hypomeron carinate ( Fig. 2)........................... 2

1b. Pronotal hypomeron not carinate.............................. 18

2a. Inflexed portion of elytron with basal fovea ( Fig. 2)................ 3

2b. Inflexed portion of elytron without basal fovea.................... 13

3a. Clypeus strongly produced anteriorly ( Fig. 80)..... 14. C. lobatus Howden

3b. Clypeus not strongly produced anteriorly........................ 4

4a. Pygidium with median swelling or longitudinal carina............... 5

4b. Pygidium without median swelling or longitudinal carina............. 11

5a. Vertical surface of clypeus foveate............................. 6

5b. Vertical surface of clypeus not foveate.......................... 7

6a. Metathoracic wings fully formed; clypeal teeth well developed in male ( Fig. 177 )................................... 33. C. parvus Howden

6b. Metathoracic wings reduced, not functional; clypeal teeth poorly developed in male ( Fig. 90 )........................... 16. C. newtoni Howden

7a. Metathoracic wings fully formed............... 18. C. osaensis sp.nov.

7b. Metathoracic wings reduced, not functional...................... 8

8a. Male protibia not lobed medially on inner margin ( Fig. 57 ); parameres broadly rounded apically ( Fig. 59 )............... 10. C. cristobalensis Howden

8b. Male protibia lobed medially; parameres not as above............... 9

9a. Male protibia with a single median lobe on inner margin ( Fig. 52 ); paramere apices narrow ( Fig. 55 ).................. 9. C. brevisetosus Howden

9b. Male protibia bilobed or trilobed on inner margin; paramere apices broad ..................................................... 10

10a. Male protibia bilobed on inner margin ( Fig. 70 ); paramere apices rounded ( Fig. 73 )................................... 12. C. howdeni sp.nov.

10b. Male protibia trilobed on inner margin ( Fig. 94 ); paramere apices angulatetruncate ( Fig. 97 )......................... 17. C. ocosingo sp.nov.

11a. Paramere apices broadly truncate ( Fig. 87 )...... 15. C. montebello sp.nov.

11b. Paramere apices roundly angulate ( Figs. 48 , 106).................. 12

12a. Elytra rounded dorsally ( Fig. 49 )................ 8. C. bochilae sp.nov.

12b. Elytra strongly arched dorsally ( Fig. 107)....... 19. C. rayonensis sp.nov.

13a. Metathoracic wings fully formed.............................. 14

13b. Metathoracic wings reduced, not functional...................... 16

14a. Paramere apices truncate, with a fringe of setae ( Fig. 143)....................................... 26. C. genieri sp.nov.

14b. Paramere apices rounded ( Figs. 113 , 189 )...................... 15

15a. Clypeal emargination of male deeply rounded between teeth ( Fig. 111 ); male protibia shallowly notched before apex ( Fig. 110 ); Costa Rica.................................................... 20. C. solisi sp.nov.

15b. Clypeal emargination of male shallow between teeth ( Fig. 187 ); male protibia strongly notched before apex ( Fig. 186 ); Ecuador.. 35. C. urguensis sp.nov

16a. Pygidium strongly lobed medially ( Fig. 77 ).................................................... 13. C. lindemanae Howden and Gill

16b. Pygidium evenly convex medially............................. 17

17a. Genae strongly produced anteriorly ( Fig. 64); elytra bilobed dorsally ( Fig. 67); parameres narrow, angulate- truncate apically ( Fig. 66)..................................................... 11. C. denticulum sp.nov.

17b. Genae not strongly produced anteriorly ( Fig. 157 ); elytra evenly convex dorsally; parameres broad, truncate at apex ( Fig. 159 )...................................... 29. C. medinae sp.nov.

18a. Inflexed portion of elytra with basal fovea....................... 19

18b. Inflexed portion of elytra lacking basal fovea.................... 21

19a. Vertical surface of clypeus foveate..................................................... 4. C. campbellorum Howden

19b. Vertical surface of clypeus not foveate......................... 20

20a. Eyes partly visible dorsally when head is retracted...... 2. C. gilli sp.nov.

20b. Eyes not visible dorsally when head is retracted..... 7. C. pumilus sp.nov.

21a. Vertical surface of clypeus foveate........................... 22

21b. Vertical surface of clypeus not foveate........................ 27

22a. Elytra fringed laterally with elongate, curved setae ( Fig. 1 ). genae strongly produced anteriorly, margin of clypeus strongly sinuate lateral to apical teeth ( Fig. 167).................................... 31. C. otonga sp.nov.

22b. Elytral setae normal, genae not strongly produced anteriorly, margin of clypeus not strongly sinuate...................................... 23

23a. Paramere apices not truncate ( Figs. 24 , 39 , 118)................... 24

23b. Paramere apices truncate ( Figs. 133, 138)........................ 26

24a. Metathoracic wings fully formed.............. 6. C. peckorum Howden

24b. Metathoracic wings reduced, not functional...................... 25

25a. Paramere apices conjointly concave ( Fig. 118); Colombia 21. C. altus Howden

25b. Paramere apices divergent, narrowly rounded ( Fig. 24 ); Venezuela............................................. 3. C. nebulinus Howden

26a. Parameres narrow apically in lateral view ( Fig. 137), lacking apical fringe of setae ( Fig. 138)............................... 25. C. foveatus sp.nov.

26b. Parameres broad apically in lateral view ( Fig. 132), with apical fringe of setae ( Fig. 133)............................... 24. C. escobari sp.nov.

27a. Posterior pronotal angles distinctly incised; paramere apices with fringe of setae ( Fig. 174)............................ 32. C. paradoxus Balthasar

27b. Posterior pronotal angles not incised; paramere apices without fringe of setae ..................................................... 28

28a. Metathoracic wings fully formed.............................. 29

28b. Metathoracic wings reduced, not functional...................... 31

29a. Paramere apices rounded ( Fig. 34 ); French Guiana.... 5. C. galboa sp.nov.

29b. Paramere apices truncate or angulate........................... 30

30a. Male protibia deeply notched on inner margin (Fig. 145); paramere apices truncate ( Fig. 148 ); Colombia, Panama......... 27. C. humidus Howden

30b. Male protibia not deeply notched on inner margin ( Fig. 125); paramere apices weakly angulate ( Fig. 128); Ecuador........... 23. C. curticrinis sp.nov.

31a. Apical abdominal sternite with median lobe on posterior margin ( Fig. 154 ) 32

31b. Apical abdominal sternite not lobed posteriorly.................... 33

32a. Male protibia strongly notched on inner margin ( Fig. 120 ); paramere apices uniform, not depressed medially ( Fig. 123 )..................................................... 22. C. chiriquinus Howden and Gill

32b. Male protibia weakly notched on inner margin ( Fig. 150 ); paramere apices depressed medially ( Fig. 153 )................ 28. C. lobipygus sp.nov.

33a. Inflexed part of elytron lacking median striae; male protibia strongly notched near apex ( Fig. 161); parameres truncate at apex ( Fig. 164)...................................................... 30. C. napoensis sp.nov.

33b. Inflexed part of elytron with median stria; male protibia weakly or not notched ( Figs. 11, 181); parameres not truncate at apex.................... 34

34a. Punctures of pronotal disc shallow, annulate; male protibia broadly, shallowly notched medially before apex ( Fig. 11); paramere apices narrowly angled ( Fig. 14)................................... 1. C. andersoni sp.nov.

34b. Punctures of pronotal disc small, deep, not annulate; male protibia not notched medially before apex ( Fig. 181); paramere apices broadly angled ( Fig. 184)..................................... 34. C. punctatus sp.nov.

Figures

Figs. 176–179. Cryptocanthon parvus. 176) left protibia, male; 177) clypeal outline; 178) aedeagus, lateral; 179) parameres, dorsal. Scale line 5 0.5 mm.

Figs. 89–92. Cryptocanthon newtoni. 89) left protibia, male; 90) clypeal outline; 91) aedeagus, lateral; 92) parameres, dorsal. Scale line 5 0.5 mm.

Figs. 57–61. Cryptocanthon cristobalensis. 57) left protibia, male; 58) clypeal outline; 59) parameres, dorsal; 60) aedeagus, lateral; 61) elytra, posterior. Scale line 5 0.5 mm (Figs. 57–60), 1.0 mm (Fig. 61).

Figs. 52–55. Cryptocanthon brevisetosus. 52) left protibia, male; 53) clypeal outline;

Figs. 70–73. Cryptocanthon howdeni. 70) left protibia, male; 71) clypeal outline; 72) aedeagus, lateral; 73) parameres, dorsal. Scale line 5 0.5 mm.

Figs. 94–97. Cryptocanthon ocosingo. 94) left protibia, male; 95) clypeal outline; 96) aedeagus, lateral; 97) parameres, dorsal. Scale line 5 0.5 mm.

Figs. 84–87. Cryptocanthon montebello. 84) left protibia, male; 85) clypeal outline; 86) aedeagus, lateral; 87) parameres, dorsal. Scale line 5 0.5 mm.

Figs. 46–50. Cryptocanthon bochilae. 46) left protibia, male; 47) clypeal outline; 48) parameres, dorsal; 49) elytra, posterior; 50) aedeagus, lateral. Scale line 5 0.5 mm (Figs. 46–48, 50), 1.0 mm (Fig. 49).

Figs. 110–113. Cryptocanthon solisi. 110) left protibia, male; 111) clypeal outline; 112) aedeagus, lateral; 113) parameres, dorsal. Scale line 5 0.5 mm.

Figs. 186–189. Cryptocanthon urguensis. 186) left protibia, male; 187) clypeal outline; 188) aedeagus, lateral; 189) parameres, dorsal. Scale line 5 0.5 mm.

Figs. 75–77. Cryptocanthon lindemanae. 75) left protibia, female; 76) clypeal outline, female;

Figs. 156–159. Cryptocanthon medinae. 156) left protibia, male; 157) clypeal outline; 158) aedeagus, lateral; 159) parameres, dorsal. Scale line 5 0.5 mm.

Fig. 1. Cryptocanthon otonga. Habitus, male.

Figs. 21–24. Cryptocanthon nebulinus. 21) left protibia, male; 22) clypeal outline;

Figs. 36–39. Cryptocanthon peckorum. 36) left protibia, male; 37) clypeal outline; 38) aedeagus, lateral; 39) parameres, dorsal. Scale line 5 0.5 mm.

Figs. 31–34. Cryptocanthon galbao. 31) left protibia, male; 32) clypeal outline; 33) aedeagus, lateral; 34) parameres, dorsal. Scale line 5 0.5 mm.

Figs. 147–148. Cryptocanthon humidus. 145) left protibia, male; 146) clypeal outline; 147) aedeagus, lateral; 148) parameres, dorsal. Scale line 5 0.5 mm.

Figs. 150–154. Cryptocanthon lobipygus. 150) left protibia, male; 151) clypeal outline; 152) aedeagus, lateral; 153) parameres, dorsal; 154) apical sternite and pygidium, ventral. Scale line 5 0.5 mm.

Figs. 120–123. Cryptocanthon chiriquinus. 120) left protibia, male; 121) clypeal outline; 122) aedeagus, lateral; 123) parameres, dorsal. Scale line 5 0.5 mm.