Cypriniformes, Bleeker, 1859

Tan, Milton & Armbruster, Jonathan W., 2018, Phylogenetic classification of extant genera of fishes of the order Cypriniformes (Teleostei: Ostariophysi), Zootaxa 4476 (1), pp. 6-39 : 6-39

publication ID

https://doi.org/ 10.11646/zootaxa.4476.1.4

publication LSID

lsid:zoobank.org:pub:C2F41B7E-0682-4139-B226-3BD32BE8949D

persistent identifier

https://treatment.plazi.org/id/03BD87B1-2D11-F814-FF2C-BC49E711AAFC

treatment provided by

Plazi

scientific name

Cypriniformes
status

 

Classification of Genera of Cypriniformes View in CoL View at ENA

Gyrinocheiloidei Gyrinocheilidae Gill 1905 (Type genus: Gyrinocheilus Vaillant 1902 ) Gyrinocheilus Vaillant 1902 (syn. Gyrinocheilops Fowler 1937 ) Catostomoidei Catostomidae Agassiz 1850 (Type genus: Catostomus Lesueur 1817 ) Catostominae Agassiz 1850 (Type genus: Catostomus Lesueur 1817 ) Catostomini Agassiz 1850 (Type genus: Catostomus Lesueur 1817 ) Catostomus Lesueur 1817 (syn. Acomus Girard 1856 , Decactylus Rafinesque 1820 , Minomus Girard 1856 , Notolepidomyzon Fowler 1913 , Pantosteus Cope in Cope & Yarrow 1875, Stomocatus Bonaparte 1839 ) Chasmistes Jordan 1878 (syn. Lipomyzon Cope 1881 , Pithecomyzon Fowler 1913 ) Deltistes Seale 1896 Xyrauchen Eigenmann & Kirsch 1899 Erimyzontini Hubbs 1930 Erimyzon Jordan 1876 Minytrema Jordan 1878 Moxostomatini Bleeker 1863 (Type genus: Moxostoma Rafinesque 1820 ) Moxostoma Rafinesque 1820 (syn. Lagochila Jordan & Brayton 1877 , Megapharynx Legendre 1942 , Placopharynx Cope 1870 , Ptychostomus Agassiz 1855 , Quassilabia Jordan & Brayton in Jordan 1878, Scartomyzon Fowler 1913 , Teretulus Rafinesque 1820 ) Thoburniini Hubbs 1930 (Type genus: Thoburnia Jordan & Snyder 1917 ) Hypentelium Rafinesque 1818 (syn. Hylomyzon Agassiz 1855 ) Thoburnia Jordan & Snyder in Jordan 1917 Cycleptinae Gill 1861 (Type genus: Cycleptus Rafinesque 1819 ) Cycleptus Rafinesque 1819 (syn. Rhytidostomus Heckel 1843 ) Ictiobinae Bleeker 1863 (Type genus: Ictiobus Rafinesque 1820 ) Carpiodes Rafinesque 1820 (syn. Sclerognathus Valenciennes in Cuvier & Valenciennes 1844) Ictiobus Rafinesque 1820 (syn. Amblodon Rafinesque 1819 , Bubalichthys Agassiz 1855 , Megastomatobus Fowler 1913 ) Myxocyprininae Fowler 1958 (Type genus: Myxocyprinus Gill 1877 ). Myxocyprinus Gill 1877 Cobitoidei Balitoridae Swainson 1839 (Type genus: Balitora Gray 1830 ) Balitorinae Swainson 1839 (Type genus: Balitora Gray 1830 )

Balitora Gray 1830 (syn. Sinohomaloptera Fang 1930 )

Balitoropsis Smith 1945

Hemimyzon Regan 1911 (syn. Dienbienia Nguyen & Nguyen 2002 )

Homaloptera van Hasselt 1823 (syn. Helgia Vinciguerra 1890 )

Jinshaia Kottelat & Chu 1988

Lepturichthys Regan 1911

Metahomaloptera Chang 1944

Pseudohomaloptera Silas 1953

Sinogastromyzon Fang 1930

Homalopteroidinae Randall & Page 2015 (Type genus: Homalopteroides Fowler 1905 )

Homalopteroides Fowler 1905 (syn. Chopraia Prashad & Mukerji 1929 )

Homalopterula Fowler 1940

Balitoridae incertae sedis

Bhavania Hora 1920

Cryptotora Kottelat 1998

Ghatsa Randall & Page 2015

Neohomaloptera Herre 1944

Travancoria Hora 1941

Barbuccidae Kottelat 2012 (Type genus: Barbucca Roberts 1989 )

Barbucca Roberts 1989

Botiidae Berg 1940 (Type genus: Botia Gray 1831 )

Botiinae Berg 1940 (Type genus: Botia Gray 1831 )

Ambastaia Kottelat 2012

Botia Gray 1831 (syn. Diacantha Swainson 1839 , Hymenphysa McClelland 1839 , often misspelled Hymenophysa )

Chromobotia Kottelat 2004

Sinibotia Fang 1936

Syncrossus Blyth 1860

Yasuhikotakia Nalbant 2002

Leptobotiinae Nalbant 2002 (Type genus: Leptobotia Bleeker 1870 )

Leptobotia Bleeker 1870

Parabotia Dabry de Thiersant 1872

Cobitidae Swainson 1838 (Type genus: Cobitis Linnaeus 1758 )

Acantopsis van Hasselt 1823 (syn. Prostheacanthus Blyth 1860 )

Aperioptus Richardson 1848 (syn. Acanthopsoides Fowler 1934 , Neacanthopsis Smith 1945 )

Bibarba Chen & Chen 2007

Canthophrys Swainson 1838 (syn. Somileptus Swainson 1839 )

Cobitis Linnaeus 1758 (syn. Acanestrinia Bǎcescu 1962 , Acanthopsis Agassiz 1832 , Acantophthalmus van Hasselt 1823 , Beyshehiria Erk'akan, Atalay-Ekmekçi & Nalbant 1999, Bicanestrinia Bǎcescu 1962 , Choia Kim, Park & Nalbant 1997 , Cobitinula Hankó 1924 , Iberocobitis Bǎcescu 1962 , Iksookimia Nalbant 1993 , Kichulchoia Kim, Park & Nalbant 1999 , Niwaella Nalbant 1963 )

Koreocobitis Kim, Park & Nalbant 1997

Kottelatlimia Nalbant 1994

Lepidocephalichthys Bleeker 1863 (syn. Cobitichthys Bleeker 1858 , Enobarbichthys Whitley 1931 , Enobarbus Whitley 1928 , Jerdonia Day 1871 , Madrasia Nalbant 1963 , Platacanthus Day 1865 )

Lepidocephalus Bleeker 1858

Microcobitis Bohlen & Harant 2011

Misgurnus Lacepède 1803 (syn. Mesomisgurnus Fang 1935 , Ussuria Nikolskii 1903 )

Neoeucirrhichthys Bănărescu & Nalbant 1968

Pangio Blyth 1860 (syn. Apua Blyth 1860 , Cobitophis Myers 1927 , Eucirrhichthys Perugia 1892 ) Paralepidocephalus Tchang 1935

Paramisgurnus Dabry de Thiersant 1872 (syn. Paramisgurnus Sauvage 1878 )

Protocobitis Yang & Chen 1993

Sabanejewia Vladykov 1929

Theriodes Kottelat 2012

Ellopostomatidae Bohlen & Šlechtová 2009 (Type genus: Ellopostoma Vaillant 1902 )

Ellopostoma Vaillant 1902

Gastromyzontidae Fowler 1905 (Type genus: Gastromyzon Günther 1874 )

Annamia Hora 1932

Beaufortia Hora 1932

Erromyzon Kottelat 2004

Formosania Oshima 1919 (syn. Crossostoma Sauvage 1878 )

Gastromyzon Günther 1874 (syn. Lepidoglanis Vaillant 1889 )

Glaniopsis Boulenger 1899

Hypergastromyzon Roberts 1989

Katibasia Kottelat 2004

Liniparhomaloptera Fang 1935

Neogastromyzon Popta 1905

Paraprotomyzon Pellegrin & Fang 1935

Parhomaloptera Vaillant 1902

Plesiomyzon Zheng & Chen 1980

Protomyzon Hora 1932 (syn. Progastromyzon Hora & Jayaram 1952 )

Pseudogastromyzon Nichols 1925 (syn. Labigastromyzon Tang & Chen 1996 )

Sewellia Hora 1932 (syn. Diardichthys Roberts 1998 , Parasewellia Nguyen & Nguyen in Nguyen 2005) Vanmanenia Hora 1932 (syn. Homalosoma Boulenger 1901 , Praeformosania Fang 1935 )

Yaoshania Yang, Kottelat, Yang & Chen 2012

Nemacheilidae Regan 1911 (Type genus: Nemacheilus Bleeker 1863 )

Aborichthys Chaudhuri 1913

Acanthocobitis Peters 1861

Afronemacheilus Golubtsov & Prokofiev in Prokofiev 2009

Barbatula Linck 1790 (syn. Cobites Swainson 1839 , Orthrias Jordan & Fowler 1903 )

Claea Kottelat 2011 (syn. Oreias Sauvage 1874 )

Draconectes Kottelat 2012

Dzihunia Prokofiev 2001

Eidinemacheilus Segherloo, Ghaedrahmati & Freyhof 2016

Eonemachilus Berg 1938

Hedinichthys Rendahl 1933 (syn. Minihedinichthys Prokofiev 2017)

Heminoemacheilus Zhu & Cao 1987

Homatula Nichols 1925

Indoreonectes Rita & Bănărescu in Rita, Bănărescu & Nalbant 1978

Iskandaria Prokofiev 2009

Lefua Herzenstein 1888 (syn. Elxis Jordan & Fowler 1903 , Octonema Herzenstein in Herzenstein &

Warpachowski 1888)

Malihkaia Kottelat 2017

Mesonoemacheilus Bănărescu & Nalbant 1982

Micronemacheilus Rendahl 1944

Nemacheilus Bleeker 1863 (syn. Modigliania Perugia 1893 , Pogononemacheilus Fowler 1937 )

Nemachilichthys Day 1878

Neonoemacheilus Zhu & Guo 1985 (syn. Infundibulatus Menon 1987 )

Oreonectes Günther 1868 (syn. Octonema Martens 1868 )

Oxynoemacheilus Bănărescu & Nalbant 1966 (syn. Ilamnemacheilus Coad & Nalbant 2005 , Nun

Bănărescu & Nalbant in Bănărescu, Nalbant & Goren 1982)

Paracanthocobitis Grant 2007

Paracobitis Bleeker 1863 (syn. Adiposia Annandale & Hora 1920 , Pseudodon Kessler 1874 )

Paranemachilus Zhu 1983

Paraschistura Prokofiev 2009 (syn. Metaschistura Prokofiev 2009 )

Petruichthys Menon 1987

Physoschistura Bănărescu & Nalbant 1982

Protonemacheilus Yang & Chu 1990

Pteronemacheilus Bohlen & Ŝlechtová 2011

Sasanidus Freyhof, Geiger, Golzarianpour & Patimar 2016

Schistura McClelland 1838 (syn. Acoura Swainson 1839 , Longischistura Bănărescu & Nalbant 1995 )

Sectoria Kottelat 1990

Seminemacheilus Bănărescu & Nalbant 1995

Speonectes Kottelat 2012

Sphaerophysa Cao & Zhu 1988

Sundoreonectes Kottelat 1990

Traccatichthys Freyhof & Serov 2001

Triplophysa Rendahl 1933 (syn. Deuterophysa Rendahl 1933 , Didymophysa Whitley 1950 , Diplophysa

Kessler 1874, Diplophysoides Fowler 1958 , Indotriplophysa Prokofiev 2010 , Labiatophysa Prokofiev

2010, Paradidymophysa Prokofiev 2017, Qinghaichthys Zhu 1981 , Tarimichthys Prokofiev 2010 ,

Tauphysa Rendahl 1933 )

Troglocobitis Parin 1983

Troglonectes Zhang, Zhao & Tang 2016

Tuberoschistura Kottelat 1990

Turcinoemacheilus Bănărescu & Nalbant 1964

Yunnanilus Nichols 1925 Serpenticobitidae Kottelat 2012 (Type genus: Serpenticobitis Roberts 1997 )

Serpenticobitis Roberts 1997 Vaillantellidae Nalbant & Bănărescu 1977 (Type genus: Vaillantella Fowler 1905 )

Vaillantella Fowler 1905 Cyprinoidei Acheilognathidae Bleeker 1863 (Type genus: Acheilognathus Bleeker 1859 )

Acheilognathus Bleeker 1859 (syn. Acanthorhodeus Bleeker 1871 , Paracheilognathus Bleeker 1863 ,

Rhodeops Fowler 1910 )

Paratanakia Chang, Chen & Mayden 2014

Pseudorhodeus Chang, Chen & Mayden 2014

Rhodeus Agassiz 1832 (syn. Pseudoperilampus Bleeker 1863 ) Sinorhodeus Li, Liao & Arai 2017 Tanakia Jordan & Thompson 1914 Cyprinidae Rafinesque 1815 (Type genus: Cyprinus Linnaeus 1758 ) Acrossocheilinae Yang et al. 2015 (Type genus: Acrossocheilus Oshima 1919 ) Acrossocheilus Oshima 1919 (syn. Lissochilichthys Oshima 1920 , Masticbarbus Tang 1942 , Sinibarbus Sauvage 1874 ) Folifer Wu 1977 Onychostoma Günther 1896 (syn. Scaphesthes Oshima 1919 , Scaphiodontella Oshima 1920 ) Barbinae Bleeker 1859 (Type genus: Barbus Daudin 1805 ) Aulopyge Heckel 1841 Barbus Daudin 1805 (syn. Pseudobarbus Bielz 1853 ) Capoeta Valenciennes 1842 (syn. Scaphiodon Heckel 1843 ) Cyprinion Heckel 1843 Luciobarbus Heckel 1843 (syn. Aspiobarbus Berg 1933 , Bertinichthys Whitley 1953 , Bertinius Fang 1943 , Messinobarbus Bianco 1998 ) Scaphiodonichthys Vinciguerra 1890 (syn. Scaphiodontopsis Fowler 1934 ) Semiplotus Bleeker 1860 Cyprininae Rafinesque 1815 (Type genus: Cyprinus Linnaeus 1758 ) Aaptosyax Rainboth 1991 Carassioides Oshima 1926 Carassius Jarocki 1822 (syn. Cyprinopsis Fitzinger 1832 , Neocarassius Castelnau 1872 ) Cyprinus Linnaeus 1758 (syn. Carpio Heckel 1843, Laichowcypris Nguyen & Doan 1969 , Laichowcypris Mai (ex Hoa & Hoa) 1978, Mesocyprinus Fang 1936 , Mesocyprinus Cheng 1950 ) Luciocyprinus Vaillant 1904 (syn. Fustis Lin 1932 ) Procypris Lin 1933 (syn. Paraprocypris Fang 1936 ) Sinocyclocheilus Fang 1936 (syn. Anchicyclocheilus Li & Lan 1992 , Gibbibarbus Dai 1988 ) Labeoninae Bleeker 1859 (Type genus: Labeo Cuvier 1816 ) Garrini Bleeker 1863 (Type genus: Garra Hamilton 1822 ) Discolabeo Fowler 1937 Garra Hamilton 1822 (syn. Discognathichthys Bleeker 1860 , Discognathus Heckel 1843 , Hemigarra Karaman 1971 , Hemigrammocapoeta Pellegrin 1927 , Horalabiosa Silas 1954 , Iranocypris Bruun & Kaiser 1944 , Lissorhynchus Bleeker 1860 , Mayoa Day 1870 , Neotylognathus Kosswig 1950 , Phreatichthys Vinciguerra 1924 , Platycara McClelland 1838 , Tylognathoides Tortonese 1938 , Typhlogarra Trewavas 1955 ) Paracrossochilus Popta 1904 Tariqilabeo Mirza & Saboohi 1990 (syn. Akrokolioplax Zhang & Kottelat 2006 , Gonorhynchus McClelland 1838 ) Labeonini Bleeker 1859 (Type genus: Labeo Cuvier 1816 ) Altigena Burton (ex Lin) 1934 (syn. Vinalabeo Nguyen, Nguyen & Nguyen 2016) Bangana Hamilton 1822 (syn. Mirolabeo Chu & Wang 1963 , Rohitodes Bleeker 1860 , Tylognathus Heckel 1843 ) Cirrhinus Oken 1817 (syn. Isocephalus Heckel 1843 ) Gymnostomus Heckel 1843 (syn. Cirrhinichthys Bleeker 1863 , Mrigala Bleeker 1860 ) Incisilabeo Fowler 1937 Labeo Cuvier 1816 (syn. Abrostomus Smith 1841 , Acra Bleeker 1860 , Catla Valenciennes in Cuvier & Valenciennes 1844, Chrysophekadion Bleeker 1860 , Gibelion Heckel 1843 ,

Morulius Hamilton 1822 , Nandina Gray 1831 , Rohita Valenciennes in Cuvier & Valenciennes 1842, Rohitichthys Bleeker 1860 , Tambra Bleeker 1860 )

Nukta Hora 1942

Speolabeo Kottelat 2017

‘Osteochilini’ (From genus Osteochilus Günther 1868 )

Barbichthys Bleeker 1860

Crossocheilus Kuhl & van Hasselt 1823 (syn. Crossocheilichthys Bleeker 1860 , Holotylognathus Fowler 1934 )

Epalzeorhynchos Bleeker 1855

Henicorhynchus Smith 1945

Labiobarbus van Hasselt 1823 (syn. Cyrene Heckel 1843 , Dangila Valenciennes in Cuvier & Valenciennes 1842)

Lobocheilos Bleeker 1853 (syn. Gobionichthys Bleeker 1859 )

Osteochilus Günther 1868 (syn. Diplocheilos Bleeker 1860 , Kantaka Hora 1942 , Neorohita Fowler 1937 )

Thynnichthys Bleeker 1859 (syn. Thynnichthyina Fowler 1937 )

‘Semilabeonini’ (From genus Semilabeo Peters 1881 )

Ageneiogarra Garman 1912

Cophecheilus Zhu, Zhang, Zhang & Han 2011

Discocheilus Zhang 1997 (syn. Discolabeo Chen in Chen & Lan 1992)

Discogobio Lin 1931

Fivepearlus Li, Yang, Li & Chen 2017

Hongshuia Zhang, Qiang & Lan 2008

Linichthys Zhang & Fang 2005

Mekongina Fowler 1937

Paraqianlabeo Zhao, Sullivan, Zhang & Peng 2014

Parasinilabeo Wu 1939 (syn. Pararectoris Su, Yang & Cui 2001 )

Placocheilus Wu in Wu, Lin, Chen, Chen & He 1977

Prolixicheilus Zheng, Chen & Yang 2016

Pseudocrossocheilus Zhang & Chen 1997

Pseudogyrinocheilus Fang 1933

Ptychidio Myers 1930 (syn. Varicogobio Lin 1931 )

Qianlabeo Zhang & Chen 2004

Rectoris Lin 1935

Semilabeo Peters 1881 (syn. Amplolabrius Lin 1933 )

Sinigarra Zhang & Zhou 2012

Sinocrossocheilus Wu 1977

Stenorynchoacrum Huang, Yang & Chen 2014

Vinagarra Nguyen & Bui 2009

Labeoninae incertae sedis

Diplocheilichthys Bleeker 1860

Longanalus Li 2006

Parapsilorhynchus Hora 1921

Protolabeo Zhang, Zhao & An 2010

Schismatorhynchos Bleeker 1855

Sinilabeo Rendahl 1933

‘Poropuntiinae’ (From genus Poropuntius Smith 1931 )

Albulichthys Bleeker 1860

Amblyrhynchichthys Bleeker 1860

Balantiocheilos Bleeker 1860

Barbonymus Kottelat 1999

Cosmochilus Sauvage 1878 (syn. Papillocheilus Smith 1945 )

Cyclocheilichthys Bleeker 1859 (syn. Anematichthys Bleeker 1859 , Oxybarbus Vaillant 1893 , Siaja Bleeker 1859 )

Cyclocheilos Bleeker 1859

Discherodontus Rainboth 1989

Eirmotus Schultz 1959

Hypsibarbus Rainboth 1996

Mystacoleucus Günther 1868 (syn. Acanthonotus Tickell in Day 1888, Matsya Day 1889 )

Poropuntius Smith 1931

Puntioplites Smith 1929 (syn. Adamacypris Fowler 1934 )

Sawbwa Annandale 1918

Scaphognathops Smith 1945 (syn. Scaphognathus Smith 1931 )

Sikukia Smith 1931 (syn. Xenocheilichthys Smith 1934 )

Probarbinae Yang et al. 2015 (Type genus: Probarbus Sauvage 1880 )

Catlocarpio Boulenger 1898

Probarbus Sauvage 1880

Schizothoracinae McClelland 1842 (Type genus: Schizothorax Heckel 1838 )

Aspiorhynchus Kessler 1879

Oreinus McClelland 1838 (syn. Englottogaster Gistel 1848 , Opistocheilos Bleeker 1860 )

Percocypris Chu 1935

Schizopyge Heckel 1848 (syn. Paraschizothorax Tsao 1964 )

Schizothorax Heckel 1838 (syn. Aspiostoma Nikolskii 1897 , Paraschizothorax Bleeker 1863 , Paratylognathus Sauvage 1880 , Racoma McClelland & Griffith in McClelland 1842, Schizothoraichthys Misra 1962 , Tetrostichodon Tchang, Yueh & Hwang 1964 )

Schizopygopsinae Mirza 1991 (Type genus: Schizopygopsis Steindachner 1866 )

Chuanchia Herzenstein 1891

Diptychus Steindachner 1866

Gymnocypris Günther 1868 (syn. Rugogymnocypris Yueh & Hwang 1964 )

Gymnodiptychus Herzenstein 1892

Herzensteinia Chu 1935

Oxygymnocypris Tsao 1964

Platypharodon Herzenstein 1891

Ptychobarbus Steindachner 1866

Schizopygopsis Steindachner 1866

Smiliogastrinae Bleeker 1863 (Type genus: Smiliogaster Bleeker 1860 )

Barbodes Bleeker 1859 (syn. Cephalakompsus Herre 1924 , Mandibularca Herre 1924 , Ospatulus Herre 1924 , Spratellicypris Herre & Myers 1931 )

Barboides Brüning 1929 (syn. Raddabarbus Thys van den Audenaerde 1971 )

Clypeobarbus Fowler 1936

Dawkinsia Pethiyagoda, Meegaskumbura & Maduwage 2012

Desmopuntius Kottelat 2013

Enteromius Cope 1867 (syn. Afropuntio Karaman 1971 , Agrammobarbus Pellegrin 1935 , Beirabarbus

Herre 1936, Estevea Whitley 1953 , Hemigrammocapoeta Estève 1952 , Hemigrammopuntius

Pellegrin 1923, Mannichthys Schultz 1942 , Nicholsopuntius Pellegrin 1933 , Parapuntius

Karaman 1971, Vanderbiltella Fowler 1936 )

Haludaria Pethiyagoda 2013 (syn. Dravidia Pethiyagoda, Meegaskumbura & Maduwage 2012 )

Hampala Kuhl & van Hasselt 1823 (syn. Hampala Bleeker 1860 , Heteroleuciscus Sauvage 1874 )

Oliotius Kottelat 2013

Oreichthys Smith 1933

Osteobrama Heckel 1843 (syn. Smiliogaster Bleeker 1860 )

Pethia Pethiyagoda, Meegaskumbura & Maduwage 2012

Pseudobarbus Smith 1841 (syn. Cheilobarbus Smith 1841 , Gnathendalia Castelnau 1861 ,

Oreodaimon Greenwood & Jubb 1967 )

Puntigrus Kottelat 2013

Puntius Hamilton 1822

Rohtee Sykes 1839

Sahyadria Raghavan, Philip, Ali & Dahanukar 2013

Striuntius Kottelat 2013

Systomus McClelland 1838 Spinibarbinae Yang et al. 2015 (Type genus: Spinibarbus Oshima 1919 )

Spinibarbichthys Oshima 1926

Spinibarbus Oshima 1919 Torinae Karaman 1971 (Type genus: Tor Gray 1834 )

Acapoeta Cockerell 1910

Arabibarbus Borkenhagen 2014

Carasobarbus Karaman 1971 (syn. Kosswigobarbus Karaman 1971 , Pseudotor Karaman 1971 )

Hypselobarbus Bleeker 1860 (syn. Gonoproktopterus Bleeker 1860 )

Labeobarbus Rüppell 1835 (syn. Barbellion Whitley 1931 , Barynotus Günther 1868 , Dillonia Heckel

1847, Lanceabarbus Fowler 1936 , Varicorhinus Rüppell 1835 )

Lepidopygopsis Raj 1941

Mesopotamichthys Karaman 1971

Naziritor Mirza & Javed 1985

Neolissochilus Rainboth 1985 (syn. Lissochilus Weber & de Beaufort 1916 )

Osteochilichthys Hora 1942

Pterocapoeta Günther 1902

Sanagia Holly 1926

Tor Gray 1834 Cyprinidae incertae sedis

Barbopsis Di Caporiacco 1926 (syn. Eilichthys Pellegrin 1929 , Zaccarinia Gianferrari 1934 )

Caecobarbus Boulenger 1921

Caecocypris Banister & Bunni 1980

Chagunius Smith 1938

Coptostomabarbus David & Poll 1937

Eechathalakenda Menon 1999

Kalimantania Bănărescu 1980

Laocypris Kottelat 2000

Neobarynotus Bănărescu 1980 Parasikukia Doi 2000

Paraspinibarbus Chu & Kottelat 1989

Parator Wu, Yang, Yue & Huang 1963

Prolabeo Norman 1932

Prolabeops Schultz 1941

Pseudosinocyclocheilus Zhang & Zhao 2016

Rohteichthys Bleeker 1860

Schizocypris Regan 1914

Troglocyclocheilus Kottelat & Bréhier 1999

Typhlobarbus Chu & Chen 1982

Xenobarbus Norman 1923 Danionidae Bleeker 1863 (Type genus: Danio Hamilton 1822 )

Chedrinae Bleeker 1863 (Type genus: Chedrus Swainson 1839 )

Barilius Hamilton 1822 (syn. Pteropsarion Günther 1868 )

Bengala Gray 1834 (syn. Megarasbora Günther 1868 )

Cabdio Hamilton 1822 (syn. Aspidoparia Heckel 1847 , Morara Bleeker 1860 )

Chelaethiops Boulenger 1899 (syn. Anchovicypris Fowler 1936 )

Engraulicypris Günther 1894 (syn. Mesobola Howes 1984 )

Leptocypris Boulenger 1900

Luciosoma Bleeker 1855 (syn. Trinematichthys Bleeker 1860 )

Malayochela Bănărescu 1968

Nematabramis Boulenger 1894 (syn. Mearnsella Seale & Bean 1907 )

Neobola Vinciguerra 1895

Opsaridium Peters 1854 (syn. Pelotrophus Günther 1864 , Sagittabarilius Fowler 1936 )

Opsarius McClelland 1838 (syn. Allodanio Smith 1945 , Bendilisis Bleeker 1860 , Chedrus Swainson

1839, Pachystomus Heckel 1843 , Paradaniops Nguyen & Doan 1969 , Perilampus McClelland

1838, Shacra Bleeker 1860 , Schacra Günther 1868 )

Raiamas Jordan 1919 (syn. Bola Günther 1868)

Rastrineobola Fowler 1936

Salmostoma Swainson 1839 (syn. Salmophasia Swainson 1839 )

Securicula Günther 1868 (syn. Pseudoxygaster Bănărescu 1967 )

Danioninae Bleeker 1863 (Type genus: Danio Hamilton 1822 )

Betadevario Pramod, Fang, Rema Devi, Liao, Indra, Jameela Beevi & Kullander 2010

Chela Hamilton 1822 (syn. Cachius Günther 1868 )

Danio Hamilton 1822 (syn. Brachydanio Weber & de Beaufort 1916 , Celestichthys Roberts 2007 )

Danionella Roberts 1986

Devario Heckel 1843 (syn. Danioides Chu 1935 , Daniops Smith 1945 , Eustira Günther 1868 ,

Parabarilius Pellegrin & Fang 1940 , Paradanio Day 1865 , Rambaibarnia Fowler 1934 )

Inlecypris Howes 1980

Laubuka Bleeker 1859 (syn. Allochela Silas 1958 )

Microdevario Fang, Norén, Liao, Källersjö & Kullander 2009

Microrasbora Annandale 1918

Neochela Silas 1958

Esominae new subfamily (Type genus: Esomus Swainson 1839 )

Esomus Swainson 1839 (syn. Nuria Valenciennes in Cuvier & Valenciennes 1842, Pogonocharax

Regan 1907)

Rasborinae Günther 1868 (Type genus: Rasbora Bleeker 1859 )

Amblypharyngodon Bleeker 1860 (syn. Brachygramma Day 1865 , Mola Heckel 1848 , Mola Blyth 1860 )

Boraras Kottelat & Vidthayanon 1993

Brevibora Liao, Kullander & Fang 2010

Horadandia Deraniyagala 1943

Kottelatia Liao, Kullander & Fang 2010

Pectenocypris Kottelat 1982

Rasbora Bleeker 1859 (syn. Parluciosoma Howes 1980 )

Rasboroides Brittan 1954

Rasbosoma Liao, Kullander & Fang 2010

Trigonopoma Liao, Kullander & Fang 2010

Trigonostigma Kottelat & Witte 1999

Danionidae incertae sedis

Thryssocypris Roberts & Kottelat 1984

Gobionidae Bleeker 1863 (Type genus: Gobio Cuvier 1816 )

Gobioninae Bleeker 1863 (Type genus: Gobio Cuvier 1816 )

Abbottina Jordan & Fowler 1903 (syn. Pseudogobiops Berg 1914 )

Acanthogobio Herzenstein 1892

Biwia Jordan & Fowler 1903

Gobio Cuvier 1816 (syn. Bungia Keyserling 1861 )

Gobiobotia Kreyenberg 1911 (syn. Progobiobotia Chen & Cao 1982 )

Mesogobio Bănărescu & Nalbant 1973

Microphysogobio Mori 1934 (syn. Huigobio Fang 1938 , Rostrogobio Taranetz 1937 )

Platysmacheilus Lu, Luo & Chen 1977

Pseudogobio Bleeker 1860

Romanogobio Bănărescu 1961 (syn. Rheogobio Bănărescu 1961 )

Saurogobio Bleeker 1870 (syn. Armatogobio Taranetz 1937 , Gobiosoma Dybowski 1872 , Longurio Jordan & Starks 1905 )

Xenophysogobio Chen & Cao 1977

Sarcocheilichthyinae Kryzanowsky 1947 (Type genus Sarcocheilichthys Bleeker 1860 )

Coreoleuciscus Mori 1935

Gnathopogon Bleeker 1860 (syn. Leucogobio Günther 1896 , Otakia Jordan & Snyder 1900 )

Gobiocypris Ye & Fu 1983

Ladislavia Dybowski 1869

Paracanthobrama Bleeker 1864 (syn. Glabrobarbus Fowler 1930 )

Pseudopungtungia Mori 1935

Pseudorasbora Bleeker 1860 (syn. Fundulichthys Bleeker 1860 , Micraspius Dybowski 1869 )

Pungtungia Herzenstein 1892 (syn. Zezera Jordan & Fowler (ex Ishikawa) 1903)

Rhinogobio Bleeker 1870 (syn. Megagobio Kessler 1876 , Rhinogobioides Rendahl 1928 )

Sarcocheilichthys Bleeker 1860 (syn. Barbodon Dybowski 1872 , Chilogobio Berg 1914 , Exoglossops Fowler & Bean 1920 , Georgichthys Nichols 1918 )

Hemibarbus Squalidus clade sensu Tang et al. 2011

Belligobio Jordan & Hubbs 1925 (syn. Hemibarboides Wang 1935 )

Hemibarbus Bleeker 1860 (syn. Gobiobarbus Dybowski 1869 )

Squalidus Dybowski 1872 (syn. Parasqualidus Doi 2000 , Sinigobio Chu 1935 )

Gobionidae incertae sedis Coreius Jordan & Starks 1905 (syn. Coripareius Garman 1912 ) Paraleucogobio Berg 1907 Placogobio Nguyen 2001 Leptobarbidae Bleeker 1864 (Type genus: Leptobarbus Bleeker 1859 ) Leptobarbus Bleeker 1859 (syn. Filirasbora Fowler 1937 ) Leuciscidae Bonaparte 1835 (Type genus: Leuciscus Cuvier 1816 ) Laviniinae Bleeker 1863 (Type genus: Lavinia Girard 1854 ) Acrocheilus Agassiz 1855 Chrosomus Rafinesque 1820 (syn. Parchrosomus Gasowska 1979 , Pfrille Jordan 1924 ) Eremichthys Hubbs & Miller 1948 Evarra Woolman 1894 Gila Baird & Girard 1853 (syn. Moapa Hubbs & Miller 1948 , Myloleucus Cope 1872 , Protoporus Cope 1872 , Siboma Girard 1856 , Temeculina Cockerell 1909 , Tigoma Girard 1856 ) Hesperoleucus Snyder 1913 (syn. Endemichthys Hopkirk 1974 ) Klamathella Miller 1945 Lavinia Girard 1854 (syn. Luxilinus Jordan 1885 ) Mylopharodon Ayres 1855 Orthodon Girard 1856 Ptychocheilus Agassiz 1855 Relictus Hubbs & Miller 1972 Siphateles Cope 1883 (syn. Leucidius Snyder 1917 ) Leuciscinae Bonaparte 1835 (Type genus: Leuciscus Cuvier 1816 ) Abramis Cuvier 1816 (syn. Brama Bleeker (ex Klein) 1863, Sapa Kazanskii 1928 , Zopa Fitzinger 1873 ) Acanthobrama Heckel 1843 (syn. Acanthalburnus Berg 1916 , Culticula Abbott 1901 ) Achondrostoma Robalo, Almada, Levy & Doadrio 2007 Alburnoides Jeitteles 1861 (syn. Spirlinus Fatio 1882 , Squalalburnus Berg 1932 ) Alburnus Rafinesque 1820 (syn. Chalcalburnus Berg 1933 ) Anaecypris Collares-Pereira 1983 Aspiolucius Berg 1907 Ballerus Heckel 1843 Blicca Heckel 1843 Capoetobrama Berg 1916 Chondrostoma Agassiz 1832 (syn. Chondrochilus Heckel 1843 , Chondrorhynchus Heckel 1843 , Machaerochilus Fitzinger 1873 , Nasus Basilewsky (ex Cuvier) 1855) Delminichthys Freyhof, Lieckfeldt, Bogutskaya, Pitra & Ludwig 2006 Iberochondrostoma Robalo, Almada, Levy & Doadrio 2007 Ladigesocypris Karaman 1972 Leucalburnus Berg 1916 Leucaspius Heckel & Kner 1857 (syn. Owsianka Dybowski 1862 ) Leuciscus Cuvier 1816 (syn. Aspiopsis Zugmayer 1912 , Aspius Agassiz 1832 , Aturius Dubalen 1878 , Bathystoma Fitzinger 1873 , Dobula Rafinesque 1820 , Genghis Howes 1984 , Idus Heckel 1843 ) Leucos Heckel 1843 Mirogrex Goren, Fishelson & Trewavas 1973 Notemigonus Rafinesque 1819 (syn. Hemiplus Rafinesque 1820 , Leucosomus Heckel 1843 , Stilbe DeKay 1842 , Stilbius Gill 1865 ) Pachychilon Steindachner 1882 Parachondrostoma Robalo, Almada, Levy & Doadrio 2007 Pelasgus Kottelat & Freyhof 2007 Pelecus Agassiz 1835 Petroleuciscus Bogutskaya 2002 Phoxinellus Heckel 1843 (syn. Paraphoxinus Bleeker 1863 ) Protochondrostoma Robalo, Almada, Levy & Doadrio 2007 Pseudochondrostoma Robalo, Almada, Levy & Doadrio 2007 Pseudophoxinus Bleeker 1860 (syn. Pararhodeus Berg 1907 , Spinophoxinellus Karaman 1972 ) Rutilus Rafinesque 1820 (syn. Cenisophius Bonaparte 1846 , Gardonus Bonaparte 1846 , Liparus Schulze 1892 , Metallites Schulze 1890 , Orfus Fitzinger 1873 , Orthroleucos Derjavin 1937 , Pararutilus Berg 1912 , Pigus Bonaparte 1846 , Rubellus Fitzinger 1873 ) Sarmarutilus Bianco & Ketmaier 2014 Scardinius Bonaparte 1837 (syn. Heegerius Bonaparte 1845 ) Squalius Bonaparte 1837 (syn. Cephalopsis Fitzinger 1873 , Cephalus Bonaparte 1846 , Iberocypris Doadrio 1980 , Microlepis Bonaparte 1846) Telestes Bonaparte 1840 (syn. Habrolepis Fitzinger 1873 ) Tropidophoxinellus Stephanidis 1974 Vimba Fitzinger 1873 (syn. Leucabramis Smitt 1895 ) Phoxininae Bleeker 1863 (Type genus: Phoxinus Rafinesque 1820 ) Phoxinus Rafinesque 1820 (syn. Acahara Jordan & Hubbs 1925 , Eulinneela Dybowski 1916 , Phoxinus Agassiz 1835 ) Plagopterinae Cope 1874 (Type genus: Plagopterus Cope 1874 ) Couesius Jordan 1878 Hemitremia Cope 1870 Lepidomeda Cope 1874 (syn. Snyderichthys Miller 1945 ) Margariscus Cockerell 1909 Meda Girard 1856 Plagopterus Cope 1874 Semotilus Rafinesque 1820 (syn. Cheilonemus Storer (ex Baird) 1855, Chilonemus Baird 1851 ) Pogonichthyinae Girard 1858 (Type genus: Pogonichthys Girard 1854 ) Agosia Girard 1856 (syn. Zophendum Jordan 1878 ) Alburnops Girard 1856 Algansea Girard 1856 (syn. Xystrosus Jordan & Snyder 1899 ) Aztecula Jordan & Evermann 1898 Campostoma Agassiz 1855 Clinostomus Girard 1856 (syn. Rhodopleuriscus Fowler 1944 ) Codoma Girard 1856 Cyprinella Girard 1856 (syn. Erogala Jordan in Jordan & Brayton 1878, Moniana Girard 1856 ) Dionda Girard 1856 Ericymba Cope 1865 Erimonax Jordan 1924 Erimystax Jordan 1882 Exoglossum Rafinesque 1818 (syn. Maxillingua Rafinesque 1818 , Parexoglossum Hubbs 1931 ) Graodus Günther 1868

Hudsonius Girard 1856

Hybognathus Agassiz 1855 (syn. Algoma Girard 1856 , Tirodon Hay 1882 )

Hybopsis Agassiz 1854

Iotichthys Jordan & Evermann 1896

Luxilus Rafinesque 1820 (syn. Coccogenia Cockerell & Callaway 1909 , Coccotis Jordan 1882 , Hypsolepis Agassiz (ex Baird) 1854, Plargyrus Rafinesque 1820 )

Lythrurus Jordan 1876

Macrhybopsis Cockerell & Allison 1909 (syn. Extrarius Jordan 1919 )

Miniellus Jordan 1888

Mylocheilus Agassiz 1855 (syn. Clarkina Jordan & Evermann 1927 )

Nocomis Girard 1856 (syn. Ceratichthys Baird in Girard 1856)

Notropis Rafinesque 1818 (syn. Alburnellus Girard 1856 , Azteca Jordan & Evermann 1896 , Chriope Jordan 1878 , Episema Cope & Jordan in Jordan 1877, Erinemus Jordan 1876 , Hydrophlox Jordan in Jordan & Brayton 1878, Minnilus Rafinesque 1820 , Nazatexico Whitley 1931 , Nototropis Jordan 1877 , Opsopoea Jordan & Evermann 1896 , Orcella Jordan & Evermann 1896 , Orcula Jordan & Evermann 1900 , Paranotropis Fowler 1904 , Photogenis Cope 1867 )

Opsopoeodus Hay 1881 (syn. Trycherodon Forbes in Jordan & Gilbert 1883)

Oregonichthys Hubbs 1929

Phenacobius Cope 1867 (syn. Sarcidium Cope 1871 )

Pimephales Rafinesque 1820 (syn. Ceraticthys Baird & Girard 1853 , Cliola Girard 1856 , Cochlognathus Baird & Girard 1854 , Coliscus Cope 1871 , Hyborhynchus Agassiz 1855 , Hypargyrus Forbes in Gilbert 1884, Spinicephalus Lesueur in Vaillant 1896)

Platygobio Gill 1863

Pogonichthys Girard 1854 (syn. Symmetrurus Jordan 1878 )

Pteronotropis Fowler 1935

Rhinichthys Agassiz 1849 (syn. Apocope Cope 1872 , Argyreus Heckel 1843 , Eritrema Cope & Yarrow 1875 )

Richardsonius Girard 1856

Stypodon Garman 1881

Tampichthys Schönhuth, Doadrio, Dominguez-Dominguez, Hillis & Mayden 2008

Tiaroga Girard 1856

Yuriria Jordan & Evermann 1896 (syn. Falcula Jordan & Snyder 1899 , Falcularius Jordan & Snyder in Jordan 1903)

Pseudaspininae Bogutskaya 1990 (Type genus: Pseudaspius Dybowski 1869 )

Oreoleuciscus Warpachowski 1889 (syn. Acanthorutilus Berg 1912 )

Pseudaspius Dybowski 1869

Rhynchocypris Günther 1889 (syn. Czekanowskiella Dybowski 1916 , Eupallasella Dybowski 1916 , Lagowskiella Dybowski 1916 , Moroco Jordan & Hubbs 1925 )

Tribolodon Sauvage 1883

Paedocyprididae Mayden & Chen 2010 (Type genus: Paedocypris Kottelat, Britz, Tan & Witte 2006 )

Paedocypris Kottelat, Britz, Tan & Witte 2006

Psilorhynchidae Hora 1926 (Type genus: Psilorhynchus McClelland 1838 )

Psilorhynchus McClelland 1838 (syn. Psilorhynchoides Yazdani, Singh & Rao 1993 )

Sundadanionidae Mayden & Chen 2010 (Type genus: Sundadanio Kottelat & Witte 1999 )

Fangfangia Britz, Kottelat & Tan 2012

Sundadanio Kottelat & Witte 1999

Tanichthyidae Mayden & Chen 2010 (Type genus: Tanichthys Lin 1932 )

Tanichthys Lin 1932

Tincidae Jordan 1878 (Type genus: Tinca Garsault 1764 )

Tinca Garsault 1764

Xenocyprididae Günther 1868 (Type genus: Xenocypris Günther 1868 )

Opsariichthyinae Rendahl 1928 (Type genus: Opsariichthys Bleeker 1863 )

Candidia Jordan & Richardson 1909

Nipponocypris Chen, Wu & Hsu 2008

Opsariichthys Bleeker 1863

Parazacco Chen 1982 (syn. Carinozacco Zhu, Wang & Ni 1982 )

Zacco Jordan & Evermann 1902

Xenocypridinae Günther 1868 (Type genus: Xenocypris Günther 1868 )

Ancherythroculter Yih & Wu 1964

Aphyocypris Günther 1868 (syn. Aphyocyprioides Tang 1942 , Caraspius Nichols 1925 , Fusania Jordan & Starks 1905 , Nicholsicypris Chu 1935 , Pararasbora Regan 1908 , Phoxiscus Oshima 1919 , Yaoshanicus Lin 1931 )

Chanodichthys Bleeker 1860 (syn. Erythroculter Berg 1909 , Leptocephalus Basilewsky 1855 , Pseudoculter Bleeker 1860 )

Ctenopharyngodon Steindachner 1866 (syn. Pristiodon Dybowski 1877 )

Culter Basilewsky 1855 (syn. Cultrichthys Smith 1938 )

Distoechodon Peters 1881

Elopichthys Bleeker 1860 (syn. Gymnognathus Sauvage 1884 , Scombrocypris Günther 1889 )

Hainania Koller 1927

Hemiculter Bleeker 1860 (syn. Cultriculus Oshima 1919 , Kendallia Evermann & Shaw 1927 , Siniichthys Bănărescu 1970 )

Hemiculterella Warpachowski 1888 (syn. Semiculter Chu 1935 )

Hemigrammocypris Fowler 1910 (syn. Brevigobio Tanaka 1916 )

Hypophthalmichthys Bleeker 1860 (syn. Abramocephalus Steindachner 1869 , Aristichthys Oshima 1919 , Cephalus Basilewsky 1855 , Onychodon Dybowski 1872 )

Ischikauia Jordan & Snyder 1900

Luciobrama Bleeker 1870

Macrochirichthys Bleeker 1859

Megalobrama Dybowski 1872 (syn. Parosteobrama Tchang 1930 )

Metzia Jordan & Thompson 1914 (syn. Rasborinus Oshima 1920 )

Mylopharyngodon Peters 1881 (syn. Leucisculus Oshima 1920 , Myloleuciscus Garman 1912 , Myloleucops Cockerell 1913 , Myloleucus Günther 1873 )

Ochetobius Günther 1868 (syn. Agenigobio Sauvage 1878 )

Oxygaster van Hasselt 1823

Parabramis Bleeker 1864

Parachela Steindachner 1881 (syn. Grandisquamachela Fowler 1934 )

Paralaubuca Bleeker 1864 (syn. Cultrops Smith 1938)

Plagiognathops Berg 1907 (syn. Plagiognathus Dybowski 1872 )

Pseudobrama Bleeker 1870

Pseudohemiculter Nichols & Pope 1927

Pseudolaubuca Bleeker 1864 (syn. Parapelecus Günther 1889 )

Sinibrama Wu 1939

Squaliobarbus Günther 1868

Toxabramis Günther 1873

Xenocyprioides Chen 1982

Xenocypris Günther 1868

Xenocyprididae incertae sedis

Anabarilius Cockerell 1923 (syn. Nicholsiculter Rendahl 1928 , Rohanus Chu 1935 )

Araiocypris Conway & Kottelat 2008

Atrilinea Chu 1935

Gymnodanio Chen & He 1992

Longiculter Fowler 1937

Pogobrama Luo 1995

Rasborichthys Bleeker 1859

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