Orizabus mcclevei, Warner, 2011

Warner, William B., 2011, Review of the genus Orizabus Fairmaire in the United States of America (Coleoptera: Scarabaeidae: Dynastinae), Insecta Mundi 2011 (174), pp. 1-42 : 3-4

Key to Orizabus species of the United States of America

1. Clypeal carina usually entire, at most weakly emarginate in anterior view; protibia tridentate (when not worn) in both sexes with apex truncate to transversely rounded ( Fig. 5-7 , 35 ); male genitalia with paramera relatively thin, laterally dentate ( Fig. 20, 21 , 33, 34 ); body usually smaller (12 to 19 mm long), strongly convex, pyriform ........................................................... 2

— Clypeal carina usually obviously bidentate in anterior view (unless worn; Fig. 1, 2 , 12, 13 , 37 ); female protibia tridentate, with apex oblique ( Fig. 10 , 37, 38 ); male with protibia blade-like, edentate ( Fig. 8, 9 , 12 , 36 ); male genitalia and shape different; body normally longer than 19 mm ............................................................................................................................................ 3

2(1). Clypeal carina apparently apical, in dorsal view often weakly angulate with angle opening anteriorly ( Fig. 6 , 32 ); pronotum with posterior marginal bead more or less effaced and anterior marginal bead not angulately broadened at middle; antennal club as long as or shorter than distance between frontoclypeal tubercle and clypeal carina ( Fig. 6 , 32 ); paramera with lateral tooth just anterior to middle, tooth often appressed ( Fig. 33, 34 ) ................... O. pyriformis (LeConte)

— Clypeal carina obviously subapical, in dorsal view straight ( Fig. 5 ); pronotum with posterior marginal bead complete or only shortly interrupted at middle, anterior marginal bead with posterior edge angularly broadened at middle; antennal club obviously longer than distance between clypeal carina and frontal tubercle ( Fig. 5 ); paramera with lateral tooth at about apical third ( Fig. 20, 21 ) ............................................................................ O. mcclevei new species

3(1). Posterior edge of anterior pronotal bead usually not posteriorly prolonged at midline, in females bead often slightly narrowed at midline ( Fig. 10 ); pronotal disc simply convex, without broad depression in anterior half ( Fig. 8 ); male paramera with apex incised ( Fig. 30 ); length usually less than 22 mm ............................................................................ O. pinalicus new species

— Posterior edge of anterior pronotal bead in male posteriorly prolonged at midline into triangular tubercle ( Fig. 1, 3 , 36 ), in female usually at least obtusely angulate (bead medially broader; Fig. 37, 38 ); pronotal disc often with broad depression behind tubercle (lacking in most females); male paramera without apical incisure ( Fig. 26-29 ); length usually more than 22 mm but smallest individuals of similar size .......................................................................................... 4

4(3). Clypeus parabolic, with lateral margins straight to weakly convex; clypeal carina obviously subapical, with teeth apices separated by about a dorsal eye width or more ( Fig. 1 , 37 ) .......... ............................................................................................................. O. clunalis (LeConte)

— Clypeus strongly narrowed, subtriangular, with lateral margins straight or weakly concave; clypeal carina apparently apical, with teeth narrowly separated (much less than a dorsal eye width) such that clypeal carina appears cleft ( Fig. 3 , 38 ) ......................... O. ligyroides Horn

Figures

Figure 36. Orizabus clunalis: Anterolateral view of male (Texas) head and pronotum.

Figures 5-7. Dorsal views of head and anterior third of pronotum (males): 5) O. mcclevei holotype. 6) O. pyriformis (Texas). 7) O. isodonoides.

Figures 32-35. Orizabus spp. 32-34) O. pyriformis: 32) Female (Colorado) dorsal view of head and anterior pronotum. 33-34) Dorsal and lateral views of paramera. 35) O. mcclevei allotype, dorsal view of right protibia.

Figures 20-25. Dorsal and lateral views of paramera: 20-21) O. mcclevei holotype. 22) O. pinalicus holotype. 23- 24) O. isodonoides. 25) O. rubricollis.

Figures 1-4. Dorsal and right lateral views of head and anterior third of pronotum: 1-2) O. clunalis. 3-4) O. ligyroides.

Figures 12-15. Dorsal habitus: 12) O. pinalicus holotype. 13) O. pinalicus allotype. 14) O. mcclevei holotype. 15) O. mcclevei allotype.

Figures 37-38. Dorsal view of female head and anterior pronotum: 37) O. clunalis (Arizona), 38) O. ligyroides.

Figures 10-11. Dorsoanterior views of female head and pronotum: 10) O. pinalicus allotype. 11) O. rubricollis.

Figures 8-9. Dorsoanterior views of male head and pronotum: 8) O. pinalicus holotype. 9) O. rubricollis.

Figures 26-31. Dorsal and lateral views of paramera: 26, 28) O. clunalis. 27, 29) O. ligyroides. 30) O. pinalicus holotype. 31) O. rubricollis.