Calapnita lawangan, Bernhard A. Huber, 2017

Bernhard A. Huber, 2017, Revision and cladistic analysis of the Southeast Asian leaf-dwelling spider genus Calapnita Simon (Araneae, Pholcidae), Zootaxa 4219 (1), pp. 1-63 : 39-40

Calapnita lawangan sp. nov. Figs 172–179

“ Calapnita vermiformis ” (misidentification; only specimens listed below): Deeleman-Reinhold 1986b: 212. Huber 2011: 48.

Diagnosis. Distinguished from other species of vermiformis group by combination of: procursus with distinctive prolateral undulating ridge and ventral flap with membranous dorso-distal tip ( Figs 172–173 ), procursus relatively short (in contrast to C. bario ), distal cheliceral apophyses bipartite (in contrast to C. bariengi and C. bario ; Fig. 176 ), male sternum pale, wide epigynal plate with short but distinct ‘knob’ ( Fig. 178 ), and round pore plates close together ( Fig. 179 ).

Etymology. Named for the Lawangan people who inhabit the eastern side of Central Kalimantan and West Kutai Regency, East Kalimantan; noun in apposition.

Material examined. Holotype. INDONESIA-BORNEO: ♂, RMNH ( ARA 17815), Central Kalimantan, Tumbang Tahai (2.033°S, 113.583°E), primary marshy forest, from leaves, 3/ 13.ix.1985 (S. Djojosudharmo).

Other material. INDONESIA-BORNEO: 1♀, RMNH ( ARA 17816), Central Kalimantan, 40 km NW Palangkaraya [~ 1.96°S, 113.75°E?], secondary forest, from leaves, ix.1985 (S. Djojosudharmo).

Assigned tentatively. INDONESIA-BORNEO: 1♂, RMNH ( ARA 17418), East Kalimantan, Russian Road km 5 (1.2°S, 116.867°E) (not 1.2°N as in Huber 2011), near sea level, 15.ii.1976 (J.R. Thompson) . 1♀, ZFMK (Ar 16029), East Kalimantan, Balikpapan [~ 1.25°S, 116.833°E], primary forest, 20.vii.1982 (Murphy, 11872 part) . INDONESIA- JAVA: 1♂ 2♀, RMNH ( ARA 17422), Jawa Barat, Ujung Kulon Reserve [~ 6.75°S, 105.33°E], from leaves, 17–19.xi.1986 (S. Djojosudharmo).

Description. Male (holotype)

MEASUREMENTS. Total body length 5.2, carapace width 0.7. Leg 1: 28.5 (7.3 + 0.3 + 6.3 + 12.2 + 2.4), tibia 2: 4.9, tibia 3: 3.1, tibia 4: 5.3; tibia 1 L/d: 105. Distance PME-PME 190 µm, diameter PME 80 µm, distance PME- ALE ~30 µm; no trace of AME.

COLOR. Entire animal pale gray to whitish, legs slightly yellowish with brown patellae and tibia-metatarsus joints.

BODY. Habitus as in close congeners (cf. Figs 120, 122 ); ocular area barely elevated, each triad on very low hump; carapace without median furrow; clypeus unmodified; sternum slightly wider than long (0.48/0.40), unmodified.

CHELICERAE. As in Figs 176–177 , apophyses near lamellae bipartite with small proximal part, with pair of barely visible lateral processes; without modified hairs; without stridulatory ridges.

PALPS. In general similar to C. vermiformis (cf. figs 139 and 140 in Huber 2011); trochanter apophysis very slender, as in Fig. 174 ; femur as in Fig. 175 , distal process at 37% of femur length; tibia length/diameter 0.48/0.27; procursus as in Figs 172–173 , with distinctive prolateral undulating ridge, simple ventro-distal flat sclerite; bulb length 0.36; embolus length 0.48; embolus tip with weak small fringes.

LEGS. Without spines and curved hairs; few vertical hairs; retrolateral trichobothrium on tibia 1 at 1.5%; prolateral trichobothrium absent on tibia 1, present on other tibiae; tarsus 1 pseudosegments not seen in dissecting microscope.

Male (variation). Tibia 1 in other male: 6.6. Specimens from East Kalimantan and Java share the distinctive undulating ridge prolaterally on the procursus but have more strongly bipartite frontal cheliceral apophyses. They are therefore assigned tentatively to this species.

Female. In general similar to male, also eye triads at same distance. Tibia 1 in 3 females: 5.8, 6.1, 6.3. Epigynum as in Fig. 178 , wider than long, with wide but distinctly separated anterior ‘knob’; internal genitalia as in Fig. 179 , slightly longer than wide, with roundish pore plates close together.

Distribution. Known from Central Kalimantan and (specimens assigned tentatively) East Kalimantan and Java ( Fig. 283 ).

Figures

FIGURE 283. Known distribution of the Calapnita vermiformis group, part 1 (cf. Fig. 284). The Java record of C. lawangan is based on specimens assigned tentatively.

FIGURES 172 – 187. Calapnita lawangan sp. nov. (172 – 179; RMNH, ARA 17815 – 16) and C. bugis sp. nov. (180 – 187; RMNH, ARA 17420). 172 – 173, 180 – 181. Left procursi, prolateral and retrolateral views (arrow points at distinctive undulating ridge). 174, 182. Left palpal trochanters, retrolateral views. 175, 183. Left palpal femora, prolateral views. 176 – 177, 184 – 185. Right male chelicerae, frontal and lateral views. 178, 186. Epigyna, ventral views. 179, 187. Cleared female genitalia, dorsal views. Scale lines: 0.3 mm.

FIGURES 120 – 127. Live specimens, Calapnita vermiformis group. 120 – 121. C. bario sp. nov., ♂ and ♀ with egg-sac from Bario. 122 – 123. C. magaseng sp. nov., ♂ and ♀ with egg-sac from Mount Penrissen. 124 – 125. C. saluang Huber, ♂ from Berastagi and ♀ with egg-sac from Phanom Bencha. 126 – 127. C. vermiformis Simon, ♂♂ from Mount Isarog and Mount Banahaw.